104-614: Stell is a name for a deep pool in a river where salmon rest and are fished, such as that found at Amble in Northumberland , England. Fishing using stell nets was made illegal by the Tweed Act in Britain in 1857. Salmon all other members of Salmoninae Salmon ( / ˈ s æ m ən / ; pl. : salmon) is the common name for several commercially important species of euryhaline ray-finned fish from
208-506: A common ancestor . The Eocene salmon's fossil from British Columbia provides evidence that the divergence between Pacific and Atlantic salmon had not yet occurred 40 million years ago. Both the fossil record and analysis of mitochondrial DNA suggest the divergence occurred 10 to 20 million years ago during the Miocene . This independent evidence from DNA analysis and the fossil record indicate that salmon divergence occurred long before
312-738: A demonstrative ( þis , þat ), after a possessive pronoun (e.g., hir , our ), or with a name or in a form of address. This derives from the Old English "weak" declension of adjectives. This inflexion continued to be used in writing even after final -e had ceased to be pronounced. In earlier texts, multisyllable adjectives also receive a final -e in these situations, but this occurs less regularly in later Middle English texts. Otherwise, adjectives have no ending and adjectives already ending in -e etymologically receive no ending as well. Earlier texts sometimes inflect adjectives for case as well. Layamon's Brut inflects adjectives for
416-442: A few species of salmon remain in fresh water throughout their life cycle, the majority are anadromous and migrate to the ocean for maturation: in these species, smolts spend a portion of their out-migration time in brackish water, where their body chemistry becomes accustomed to osmoregulation in the ocean. This body chemistry change is hormone-driven, causing physiological adjustments in the function of osmoregulatory organs such as
520-477: A fish returns after just one year's sea feeding, it is called a grilse in Canada, Britain, and Ireland. Grilse may be present at spawning, and go unnoticed by large males, releasing their own sperm on the eggs. Prior to spawning, depending on the species , salmon undergo changes. They may grow a hump, develop canine-like teeth, or develop a kype (a pronounced curvature of the jaws in male salmon). All change from
624-692: A future in aquaculture that eliminates the need for land, freshwater, or fertilizer to raise fish. Salmon population levels are of concern in the Atlantic and in some parts of the Pacific. The population of wild salmon declined markedly in recent decades, especially North Atlantic populations, which spawn in the waters of western Europe and eastern Canada, and wild salmon in the Snake and Columbia River systems in northwestern United States. Middle English language Middle English (abbreviated to ME )
728-569: A high demand for wild forage fish . As a predator, salmon require large nutritional intakes of protein , and farmed salmon consume more fish than they generate as a final product. On a dry weight basis, 2–4 kg of wild-caught fish are needed to produce one kilogram of salmon. As the salmon farming industry expands, it requires more forage fish for feed, at a time when 75% of the world's monitored fisheries are already near to or have exceeded their maximum sustainable yield . The industrial-scale extraction of wild forage fish for salmon farming affects
832-502: A largely Anglo-Saxon vocabulary (with many Norse borrowings in the northern parts of the country) but a greatly simplified inflectional system. The grammatical relations that were expressed in Old English by the dative and instrumental cases were replaced in Early Middle English with prepositional constructions. The Old English genitive - es survives in the -'s of the modern English possessive , but most of
936-494: A lengthened – and later also modified – pronunciation of a preceding vowel. For example, in name , originally pronounced as two syllables, the /a/ in the first syllable (originally an open syllable) lengthened, the final weak vowel was later dropped, and the remaining long vowel was modified in the Great Vowel Shift (for these sound changes, see Phonology , above). The final ⟨e⟩ , now silent, thus became
1040-495: A lesser extent), and, therefore, it cannot be attributed simply to the influence of French-speaking sections of the population: English did, after all, remain the vernacular . It is also argued that Norse immigrants to England had a great impact on the loss of inflectional endings in Middle English. One argument is that, although Norse and English speakers were somewhat comprehensible to each other due to similar morphology,
1144-548: A lot of work on Henneguya salminicola was done by scientists at the Pacific Biological Station in Nanaimo in the mid-1980s, in particular, an overview report which states, "the fish that have the longest fresh water residence time as juveniles have the most noticeable infections. Hence in order of prevalence , coho are most infected followed by sockeye, chinook, chum and pink. As well, the report says, at
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#17327722099511248-593: A process called apophony ), as in Modern English. With the discontinuation of the Late West Saxon standard used for the writing of Old English in the period prior to the Norman Conquest, Middle English came to be written in a wide variety of scribal forms, reflecting different regional dialects and orthographic conventions. Later in the Middle English period, however, and particularly with
1352-412: A technically demanding kind of sport fishing , not necessarily intuitive for beginning fishermen. A conflict exists between commercial fishermen and recreational fishermen for the right to salmon stock resources . Commercial fishing in estuaries and coastal areas is often restricted so enough salmon can return to their natal rivers where they can spawn and be available for sport fishing. On parts of
1456-524: A variant of the Northumbrian dialect (prevalent in northern England and spoken in southeast Scotland ). During the Middle English period, many Old English grammatical features either became simplified or disappeared altogether. Noun, adjective, and verb inflections were simplified by the reduction (and eventual elimination) of most grammatical case distinctions. Middle English also saw considerable adoption of Anglo-Norman vocabulary, especially in
1560-725: Is a form of the English language that was spoken after the Norman Conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old English period. Scholarly opinion varies, but the University of Valencia states the period when Middle English was spoken as being from 1150 to 1500. This stage of the development of the English language roughly coincided with
1664-858: Is a major contributor to the world production of farmed finfish, representing about US$ 10 billion annually. Other commonly cultured fish species include tilapia , catfish , sea bass , carp and bream . Salmon farming is significant in Chile , Norway , Scotland , Canada and the Faroe Islands ; it is the source for most salmon consumed in the United States and Europe. Atlantic salmon are also, in very small volumes, farmed in Russia and Tasmania , Australia. Salmon are carnivorous , and need to be fed meals produced from catching other wild forage fish and other marine organisms. Salmon farming leads to
1768-680: Is derived from Middle English : samoun , samon and saumon , which in turn are from Anglo-Norman : saumon , from Old French : saumon , and from Latin : salmō (which in turn might have originated from salire , meaning "to leap". ). The unpronounced "l" absent from Middle English was later added as a Latinisation to make the word closer to its Latin root. The term salmon has mostly displaced its now dialectal synonym lax , in turn from Middle English : lax , from Old English : leax , from Proto-Germanic : * lahsaz from Proto-Indo-European : *lakso- . The seven commercially important species of salmon occur in two genera of
1872-433: Is now rare and used only in oxen and as part of a double plural , in children and brethren . Some dialects still have forms such as eyen (for eyes ), shoon (for shoes ), hosen (for hose(s) ), kine (for cows ), and been (for bees ). Grammatical gender survived to a limited extent in early Middle English before being replaced by natural gender in the course of the Middle English period. Grammatical gender
1976-441: Is quite high (perhaps as high as 40 to 50%). To lay her roe , the female salmon uses her tail (caudal fin), to create a low-pressure zone, lifting gravel to be swept downstream, excavating a shallow depression, called a redd. The redd may sometimes contain 5,000 eggs covering 2.8 m (30 sq ft). The eggs usually range from orange to red. One or more males approach the female in her redd, depositing sperm, or milt, over
2080-966: Is strictly a fish parasite that cannot live in or affect warm blooded animals, including man". According to Klaus Schallie, Molluscan Shellfish Program Specialist with the Canadian Food Inspection Agency , " Henneguya salminicola is found in southern B.C. also and in all species of salmon. I have previously examined smoked chum salmon sides that were riddled with cysts and some sockeye runs in Barkley Sound (southern B.C., west coast of Vancouver Island ) are noted for their high incidence of infestation." Sea lice , particularly Lepeophtheirus salmonis and various Caligus species, including C. clemensi and C. rogercresseyi , can cause deadly infestations of both farm-grown and wild salmon. Sea lice are ectoparasites which feed on mucus, blood, and skin, and migrate and latch onto
2184-480: Is under development in Alaska . There, the young salmon are released into the ocean far from any wild salmon streams. When it is time for them to spawn, they return to where they were released, where fishermen can catch them. An alternative method to hatcheries is to use spawning channels. These are artificial streams , usually parallel to an existing stream, with concrete or rip-rap sides and gravel bottoms. Water from
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#17327722099512288-524: The Augustinian canon Orrm wrote the Ormulum , one of the oldest surviving texts in Middle English. The influence of Old Norse aided the development of English from a synthetic language with relatively free word order to a more analytic language with a stricter word order. Both Old English and Old Norse were synthetic languages with complicated inflections. Communication between Vikings in
2392-624: The Cook Inlet Aquaculture Association , as a way to increase salmon populations in situations where they have declined due to overharvesting , construction of dams and habitat destruction or fragmentation . Negative consequences to this sort of population manipulation include genetic "dilution" of the wild stocks. Many jurisdictions are now beginning to discourage supplemental fish planting in favour of harvest controls, and habitat improvement and protection. A variant method of fish stocking , called ocean ranching,
2496-597: The Danelaw and their Anglo-Saxon neighbours resulted in the erosion of inflection in both languages. Old Norse may have had a more profound impact on Middle and Modern English development than any other language. The effect of Old Norse on Old English was substantive, pervasive, and of a democratic character. Like close cousins, Old Norse and Old English resembled each other, and with some words and grammatical structures in common, speakers of each language roughly understood each other, but according to historian Simeon Potler
2600-616: The Early Modern English and Modern English eras. Middle English generally did not have silent letters . For example, knight was pronounced [ˈkniçt] (with both the ⟨k⟩ and the ⟨gh⟩ pronounced, the latter sounding as the ⟨ch⟩ in German Knecht ). The major exception was the silent ⟨e⟩ – originally pronounced but lost in normal speech by Chaucer's time. This letter, however, came to indicate
2704-476: The High and Late Middle Ages . Middle English saw significant changes to its vocabulary, grammar, pronunciation, and orthography . Writing conventions during the Middle English period varied widely. Examples of writing from this period that have survived show extensive regional variation. The more standardized Old English literary variety broke down and writing in English became fragmented and localized and was, for
2808-489: The Japanese word for trout. On the other hand, the steelhead and sea trout , the anadromous forms of rainbow trout and brown trout respectively, are from the same genera as salmon and live identical migratory lives, but neither is termed "salmon" . The extinct Eosalmo driftwoodensis , the oldest known Salmoninae fish in the fossil record , helps scientists figure how the different species of salmon diverged from
2912-585: The Norman Conquest , had normally been written in French. Like Chaucer's work, this new standard was based on the East Midlands-influenced speech of London. Clerks using this standard were usually familiar with French and Latin , influencing the forms they chose. The Chancery Standard, which was adopted slowly, was used in England by bureaucrats for most official purposes, excluding those of
3016-526: The Pacific Northwest and Alaska , salmon are keystone species . The migration of salmon represent a massive retrograde nutrient transfer, rich in nitrogen , sulfur , carbon and phosphorus , from the ocean to the inland freshwater ecosystems . Predation by piscivorous land animals (such as ospreys , bears and otters ) along the journey serve to transfer the nutrients from the water to land, and decomposition of salmon carcass benefits
3120-461: The Pacific coast of Canada , the louse-induced mortality of pink salmon in some regions is commonly over 80%. The risk of injury caused by underwater pile driving has been studied by Dr. Halvorsen and her co-workers. The study concluded that the fish are at risk of injury if the cumulative sound exposure level exceeds 210 dB relative to 1 μPa s. As can be seen from the production chart at
3224-514: The Quaternary glaciation began the cycle of glacial advance and retreat. There are several other species of fish which are colloquially called "salmon" but are not true salmon. Of those listed below, the Danube salmon or huchen is a large freshwater salmonid closely related (from the same subfamily) to the seven species of salmon above, but others are fishes of unrelated orders , given
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3328-618: The Salmo and Oncorhynchus genera also contain a number of trout species informally referred to as salmon. Within Salmo , the Adriatic salmon ( Salmo obtusirostris ) and Black Sea salmon ( Salmo labrax ) have both been named as salmon in English , although they fall outside the generally recognized seven salmon species. The masu salmon ( Oncorhynchus masou ) is actually considered a trout ("cherry trout") in Japan , with masu actually being
3432-497: The forest ecosystem . In the case of Pacific salmon , most (if not all) of the salmon that survive to reach the headwater spawning grounds will die after laying eggs and their dead bodies sink to cover the gravel beds, with the nutrients released from the biodegradation of their corpses providing a significant boost to these otherwise biomass -poor shallow streams. Grizzly bears function as ecosystem engineers , capturing salmon and carrying them into adjacent dry land to eat
3536-601: The genera Salmo and Oncorhynchus of the family Salmonidae , native to tributaries of the North Atlantic ( Salmo ) and North Pacific ( Oncorhynchus ) basins. Other closely related fish in the same family include trout , char , grayling , whitefish , lenok and taimen , all coldwater fish of the subarctic and cooler temperate regions with some sporadic endorheic populations in Central Asia . Salmon are typically anadromous : they hatch in
3640-524: The myrtle zone . These dams can be overtopped at high tide and hold water at low tide. This provides refuges for juvenile salmon so they do not have to swim into large channels where they are subject to predation by larger fish. It has been discovered that rivers which have seen a decline or disappearance of anadromous lampreys , loss of the lampreys also affects the salmon in a negative way. Like salmon, anadromous lampreys stop feeding and die after spawning, and their decomposing bodies release nutrients into
3744-590: The otolith (annuli), analogous to the growth rings visible in a tree trunk. Freshwater growth shows as densely crowded rings, sea growth as widely spaced rings; spawning is marked by significant erosion as body mass is converted into eggs and milt. Freshwater streams and estuaries provide important habitat for many salmon species. They feed on terrestrial and aquatic insects , amphipods , and other crustaceans while young, and primarily on other fish when older. Eggs are laid in deeper water with larger gravel and need cool water and good water flow (to supply oxygen) to
3848-849: The subfamily Salmoninae . The genus Salmo contains the Atlantic salmon , found in both sides of the North Atlantic , as well as more than 40 other species commonly named as trout . The genus Oncorhynchus contains 12 recognised species which occur naturally only in the North Pacific, six of which are known as Pacific salmon while the remainder are considered trout. Outside their native habitats, Chinook salmon have been successfully introduced in New Zealand and Patagonia , while coho , sockeye and Atlantic salmon have been established in Patagonia, as well. Both
3952-645: The 12th century, incorporating a unique phonetic spelling system; and the Ancrene Wisse and the Katherine Group , religious texts written for anchoresses , apparently in the West Midlands in the early 13th century. The language found in the last two works is sometimes called the AB language . Additional literary sources of the 12th and 13th centuries include Layamon's Brut and The Owl and
4056-422: The 13th century and was replaced by thorn. Thorn mostly fell out of use during the 14th century and was replaced by ⟨th⟩ . Anachronistic usage of the scribal abbreviation [REDACTED] ( þe , "the") has led to the modern mispronunciation of thorn as ⟨ y ⟩ in this context; see ye olde . Wynn, which represented the phoneme /w/ , was replaced by ⟨ w ⟩ during
4160-409: The 13th century. Due to its similarity to the letter ⟨p⟩ , it is mostly represented by ⟨w⟩ in modern editions of Old and Middle English texts even when the manuscript has wynn. Under Norman influence, the continental Carolingian minuscule replaced the insular script that had been used for Old English. However, because of the significant difference in appearance between
4264-473: The 14th century, even after the loss of the majority of the continental possessions of the English monarchy . In the aftermath of the Black Death of the 14th century, there was significant migration into London , of people to the counties of the southeast of England and from the east and central Midlands of England, and a new prestige London dialect began to develop as a result of this clash of
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4368-604: The 1540s after the printing and wide distribution of the English Bible and Prayer Book , which made the new standard of English publicly recognizable and lasted until about 1650. The main changes between the Old English sound system and that of Middle English include: The combination of the last three processes listed above led to the spelling conventions associated with silent ⟨e⟩ and doubled consonants (see under Orthography , below). Middle English retains only two distinct noun-ending patterns from
4472-572: The Church and legalities, which used Latin and Law French respectively. The Chancery Standard's influence on later forms of written English is disputed, but it did undoubtedly provide the core around which Early Modern English formed. Early Modern English emerged with the help of William Caxton 's printing press, developed during the 1470s. The press stabilized English through a push towards standardization, led by Chancery Standard enthusiast and writer Richard Pynson . Early Modern English began in
4576-557: The Nightingale . Some scholars have defined "Early Middle English" as encompassing English texts up to 1350. This longer time frame would extend the corpus to include many Middle English Romances (especially those of the Auchinleck manuscript c. 1330 ). Gradually, the wealthy and the government Anglicised again, although Norman (and subsequently French ) remained the dominant language of literature and law until
4680-591: The Norse speakers' inability to reproduce the ending sounds of English words influenced Middle English's loss of inflectional endings. Important texts for the reconstruction of the evolution of Middle English out of Old English are the Peterborough Chronicle , which continued to be compiled up to 1154; the Ormulum , a biblical commentary probably composed in Lincolnshire in the second half of
4784-479: The North American West Coast salmon sport fishing has completely replaced inshore commercial salmon fishing. In most cases, the commercial value of a salmon sold as seafood can be several times less than the value attributed to the same fish caught by a sport fisherman. This is "a powerful economic argument for allocating stock resources preferentially to sport fishing". Salmon aquaculture
4888-544: The Old English -eþ , Midland dialects showing -en from about 1200, and Northern forms using -es in the third person singular as well as the plural. The past tense of weak verbs was formed by adding an -ed(e) , -d(e) , or -t(e) ending. The past-tense forms, without their personal endings, also served as past participles with past-participle prefixes derived from Old English: i- , y- , and sometimes bi- . Strong verbs , by contrast, formed their past tense by changing their stem vowel (e.g., binden became bound ,
4992-713: The Old Norse influence was strongest in the dialects under Danish control that composed the southern part of the Northern England (corresponding to the Scandinavian Kingdom of Jórvík ), the East Midlands and the East of England , words in the spoken language emerged in the 10th and 11th centuries near the transition from Old to Middle English. Influence on the written languages only appeared from
5096-570: The Pacific Ocean, the second host releases a stage infective to salmon. The parasite is then carried in the salmon until the next spawning cycle. The myxosporean parasite that causes whirling disease in trout has a similar life cycle. However, as opposed to whirling disease, the Henneguya infestation does not appear to cause disease in the host salmon—even heavily infected fish tend to return to spawn successfully. According to Dr. Kieser,
5200-710: The abundance of Modern English words for the mechanisms of government that are derived from Anglo-Norman, such as court , judge , jury , appeal , and parliament . There are also many Norman-derived terms relating to the chivalric cultures that arose in the 12th century, an era of feudalism , seigneurialism , and crusading . Words were often taken from Latin, usually through French transmission. This gave rise to various synonyms, including kingly (inherited from Old English), royal (from French, inherited from Vulgar Latin), and regal (from French, which borrowed it from Classical Latin). Later French appropriations were derived from standard, rather than Norman, French. Examples of
5304-432: The adjacent stream is piped into the top of the channel, sometimes via a header pond, to settle out sediment. Spawning success is often much better in channels than in adjacent streams due to the control of floods, which in some years can wash out the natural redds. Because of the lack of floods, spawning channels must sometimes be cleaned out to remove accumulated sediment. The same floods that destroy natural redds also clean
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#17327722099515408-494: The areas of politics, law, the arts, and religion, as well as poetic and emotive diction. Conventional English vocabulary remained primarily Germanic in its sources, with Old Norse influences becoming more apparent. Significant changes in pronunciation took place, particularly involving long vowels and diphthongs, which in the later Middle English period began to undergo the Great Vowel Shift . Little survives of early Middle English literature , due in part to Norman domination and
5512-583: The beginning of the 13th century, this delay in Scandinavian lexical influence in English has been attributed to the lack of written evidence from the areas of Danish control, as the majority of written sources from Old English were produced in the West Saxon dialect spoken in Wessex , the heart of Anglo-Saxon political power at the time. The Norman Conquest of England in 1066 saw the replacement of
5616-414: The clergy for written communication and record-keeping. A significant number of Norman words were borrowed into English and used alongside native Germanic words with similar meanings. Examples of Norman/Germanic pairs in Modern English include pig and pork , calf and veal , wood and forest , and freedom and liberty . The role of Anglo-Norman as the language of government and law can be seen in
5720-608: The common name "salmon" simply due to similar shapes, behaviors and niches occupied: Salmon eggs are laid in freshwater streams typically at high latitudes. The eggs hatch into alevin or sac fry. The fry quickly develop into parr with camouflaging vertical stripes. The parr stay for six months to three years in their natal stream before becoming smolts, which are distinguished by their bright, silvery colour with scales that are easily rubbed off. Only 10% of all salmon eggs are estimated to survive to this stage. The smolt body chemistry changes, allowing them to live in salt water. While
5824-507: The comparative and superlative (e.g., greet , great; gretter , greater). Adjectives ending in -ly or -lich formed comparatives either with -lier , -liest or -loker , -lokest . A few adjectives also displayed Germanic umlaut in their comparatives and superlatives, such as long , lenger . Other irregular forms were mostly the same as in modern English. Middle English personal pronouns were mostly developed from those of Old English , with
5928-430: The developing embryos. Mortality of salmon in the early life stages is usually high due to natural predation and human-induced changes in habitat, such as siltation, high water temperatures, low oxygen concentration, loss of stream cover, and reductions in river flow. Estuaries and their associated wetlands provide vital nursery areas for the salmon prior to their departure to the open ocean. Wetlands not only help buffer
6032-486: The development of the Chancery Standard in the 15th century, orthography became relatively standardised in a form based on the East Midlands-influenced speech of London. Spelling at the time was mostly quite regular . (There was a fairly consistent correspondence between letters and sounds.) The irregularity of present-day English orthography is largely due to pronunciation changes that have taken place over
6136-495: The different dialects, that was based chiefly on the speech of the East Midlands but also influenced by that of other regions. The writing of this period, however, continues to reflect a variety of regional forms of English. The Ayenbite of Inwyt , a translation of a French confessional prose work, completed in 1340, is written in a Kentish dialect . The best known writer of Middle English, Geoffrey Chaucer , wrote in
6240-531: The double consonant represented a sound that was (or had previously been) geminated (i.e., had genuinely been "doubled" and would thus have regularly blocked the lengthening of the preceding vowel). In other cases, by analogy, the consonant was written double merely to indicate the lack of lengthening. The basic Old English Latin alphabet consisted of 20 standard letters plus four additional letters: ash ⟨æ⟩ , eth ⟨ð⟩ , thorn ⟨þ⟩ , and wynn ⟨ƿ⟩ . There
6344-472: The elimination of beavers from large parts of the river system, salmon runs plummeted, even in the absence of many of the factors usually associated with the demise of salmon runs. Salmon recruitment can be affected by beavers' dams because dams can: Beaver dams are able to nurture salmon juveniles in estuarine tidal marshes where the salinity is less than 10 ppm. Beavers build small dams of generally less than 60 cm (2 ft) high in channels in
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#17327722099516448-468: The end of the Middle English period only the strong -'s ending (variously spelled) was in use. Some formerly feminine nouns, as well as some weak nouns, continued to make their genitive forms with -e or no ending (e.g., fole hoves , horses' hooves), and nouns of relationship ending in -er frequently have no genitive ending (e.g., fader bone , "father's bane"). The strong -(e)s plural form has survived into Modern English. The weak -(e)n form
6552-821: The estuary from silt and pollutants, but also provide important feeding and hiding areas. Salmon not killed by other means show greatly accelerated deterioration ( phenoptosis , or "programmed aging") at the end of their lives. Their bodies rapidly deteriorate right after they spawn as a result of the release of massive amounts of corticosteroids . Salmon are mid- level carnivores whose diet change according to their life stage. Salmon fry predominantly feed upon zooplanktons until they reach fingerling sizes, when they start to consume more aquatic invertebrates such as insect larvae , micro crustaceans and worms . As juveniles (parrs), they become more predatory and actively prey upon aquatic insects , small crustaceans, tadpoles and small bait fishes . They are also known to breach
6656-413: The exact stream where they themselves hatched to spawn , and tracking studies have shown this to be mostly true. A portion of a returning salmon run may stray and spawn in different freshwater systems; the percent of straying depends on the species of salmon. Homing behavior has been shown to depend on olfactory memory . Salmon are important food fish and are intensively farmed in many parts of
6760-418: The exception of the third person plural, a borrowing from Old Norse (the original Old English form clashed with the third person singular and was eventually dropped). Also, the nominative form of the feminine third person singular was replaced by a form of the demonstrative that developed into sche (modern she ), but the alternative heyr remained in some areas for a long time. As with nouns, there
6864-443: The fish. There they deposit nutrient-rich urine and feces and partially eaten carcasses. Bears preparing for hibernation tend to preferentially consume the more nutrient- and energy -rich salmon roes and brain over the actual flesh , and are estimated to discard up to half the salmon they've harvested uneaten on the forest floor, in densities that can reach 4,000 kg (8,800 lb) per hectare, providing as much as 24% of
6968-566: The gills, which leads to large increases in their ability to secrete salt. Hormones involved in increasing salinity tolerance include insulin-like growth factor I , cortisol , and thyroid hormones , which permits the fish to endure the transition from a freshwater environment to the ocean. The salmon spend about one to five years (depending on the species) in the open ocean, where they gradually become sexually mature. The adult salmon then return primarily to their natal streams to spawn. Atlantic salmon spend between one and four years at sea. When
7072-412: The indicator of the longer and changed pronunciation of ⟨a⟩ . In fact, vowels could have this lengthened and modified pronunciation in various positions, particularly before a single consonant letter and another vowel or before certain pairs of consonants. A related convention involved the doubling of consonant letters to show that the preceding vowel was not to be lengthened. In some cases,
7176-684: The left, the global capture reported by different countries to the FAO of commercial wild salmon has remained fairly steady since 1990 at about one million tonnes per year. This is in contrast to farmed salmon (below) which has increased in the same period from about 0.6 million tonnes to well over two million tonnes. Nearly all captured wild salmon are Pacific salmon . The capture of wild Atlantic salmon has always been relatively small, and has declined steadily since 1990. In 2011 only 2,500 tonnes were reported. In contrast, about half of all farmed salmon are Atlantic salmon. Recreational salmon fishing can be
7280-458: The longer the fish remain in fresh water, and they then deteriorate further after they spawn, when they are known as kelts. In all species of Pacific salmon, the mature individuals die within a few days or weeks of spawning, a trait known as semelparity . Between 2 and 4% of Atlantic salmon kelts survive to spawn again, all females. However, even in those species of salmon that may survive to spawn more than once ( iteroparity ), postspawning mortality
7384-561: The main difference lied on their inflectional endings, which led to much confusion within the mixed population that existed in the Danelaw, this endings tended gradually to become obscured and finally lost, "simplifying English grammar" in the process. In time, the inflections melted away and the analytic pattern emerged. Viking influence on Old English is most apparent in pronouns , modals, comparatives, pronominal adverbs (like hence and together ), conjunctions, and prepositions show
7488-417: The masculine accusative, genitive, and dative, the feminine dative, and the plural genitive. The Owl and the Nightingale adds a final -e to all adjectives not in the nominative, here only inflecting adjectives in the weak declension (as described above). Comparatives and superlatives were usually formed by adding -er and -est . Adjectives with long vowels sometimes shortened these vowels in
7592-427: The more complex system of inflection in Old English : Nouns of the weak declension are primarily inherited from Old English n -stem nouns but also from ō -stem, wō -stem, and u -stem nouns, which did not inflect in the same way as n -stem nouns in Old English, but joined the weak declension in Middle English. Nouns of the strong declension are inherited from the other Old English noun stem classes. Some nouns of
7696-490: The most marked Danish influence. The best evidence of Scandinavian influence appears in extensive word borrowings; however, texts from the period in Scandinavia and Northern England do not provide certain evidence of an influence on syntax. However, at least one scholarly study of this influence shows that Old English may have been replaced entirely by Norse, by virtue of the change from Old English to Norse syntax. While
7800-551: The most part, being improvised. By the end of the period (about 1470), and aided by the invention of the printing press by Johannes Gutenberg in 1439, a standard based on the London dialects (Chancery Standard) had become established. This largely formed the basis for Modern English spelling, although pronunciation has changed considerably since that time. Middle English was succeeded in England by Early Modern English , which lasted until about 1650. Scots developed concurrently from
7904-693: The old insular g and the Carolingian g (modern g ), the former continued in use as a separate letter, known as yogh , written ⟨ȝ⟩ . This was adopted for use to represent a variety of sounds: [ɣ], [j], [dʒ], [x], [ç] , while the Carolingian g was normally used for [g]. Instances of yogh were eventually replaced by ⟨j⟩ or ⟨y⟩ and by ⟨gh⟩ in words like night and laugh . In Middle Scots , yogh became indistinguishable from cursive z , and printers tended to use ⟨z⟩ when yogh
8008-423: The other case endings disappeared in the Early Middle English period, including most of the roughly one dozen forms of the definite article ("the"). The dual personal pronouns (denoting exactly two) also disappeared from English during this period. The loss of case endings was part of a general trend from inflections to fixed word order that also occurred in other Germanic languages (though more slowly and to
8112-446: The parasitic infection into a number of cysts that contain milky fluid. This fluid is an accumulation of a large number of parasites. Henneguya and other parasites in the myxosporean group have complex life cycles, where the salmon is one of two hosts. The fish releases the spores after spawning. In the Henneguya case, the spores enter a second host, most likely an invertebrate, in the spawning stream. When juvenile salmon migrate to
8216-526: The preferred prey of seals and sea lions, which can eat 30 lampreys to every salmon, allowing more adult salmon to enter the rivers to spawn without being eaten by the marine mammals. According to Canadian biologist Dorothy Kieser, the myxozoan parasite Henneguya salminicola is commonly found in the flesh of salmonids. It has been recorded in the field samples of salmon returning to the Haida Gwaii Islands . The fish responds by walling off
8320-403: The prestige that came with writing in French rather than English. During the 14th century, a new style of literature emerged with the works of writers including John Wycliffe and Geoffrey Chaucer , whose Canterbury Tales remains the most studied and read work of the period. The transition from Late Old English to Early Middle English had taken place by the 1150s to 1180s, the period when
8424-408: The problems of rising costs for buying hatchery fish feed . Yet another attractive alternative is the increased use of seaweed . Seaweed provides essential minerals and vitamins for growing organisms. It offers the advantage of providing natural amounts of dietary fiber and having a lower glycemic load than grain-based fish meal . In the best-case scenario, widespread use of seaweed could yield
8528-466: The regular streams. Spawning channels preserve the natural selection of natural streams, as there is no benefit, as in hatcheries, to use prophylactic chemicals to control diseases. Farm-raised salmon are fed the carotenoids astaxanthin and canthaxanthin to match their flesh colour to wild salmon to improve their marketability. Wild salmon get these carotenoids , primarily astaxanthin, from eating shellfish and krill . One proposed alternative to
8632-569: The resulting doublet pairs include warden (from Norman) and guardian (from later French; both share a common ancestor loaned from Germanic). The end of Anglo-Saxon rule did not result in immediate changes to the language. The general population would have spoken the same dialects as they had before the Conquest. Once the writing of Old English came to an end, Middle English had no standard language, only dialects that evolved individually from Old English. Early Middle English (1150–1350) has
8736-407: The roe. The female then covers the eggs by disturbing the gravel at the upstream edge of the depression before moving on to make another redd. The female may make as many as seven redds before her supply of eggs is exhausted. Each year, the fish experiences a period of rapid growth, often in summer, and one of slower growth, normally in winter. This results in ring formation around an earbone called
8840-509: The salmon population. This system is referred to as ranching . It was very common in countries such as Sweden, before the Norwegians developed salmon farming, but is seldom done by private companies. As anyone may catch the salmon when they return to spawn, a company is limited in benefiting financially from their investment. Because of this, the ranching method has mainly been used by various public authorities and non-profit groups, such as
8944-807: The second half of the 14th century in the emerging London dialect, although he also portrays some of his characters as speaking in northern dialects, as in " The Reeve's Tale ". In the English-speaking areas of lowland Scotland , an independent standard was developing, based on the Northumbrian dialect . This would develop into what came to be known as the Scots language . A large number of terms for abstract concepts were adopted directly from scholastic philosophical Latin (rather than via French). Examples are "absolute", "act", "demonstration", and "probable". The Chancery Standard of written English emerged c. 1430 in official documents that, since
9048-458: The second person singular in -(e)st (e.g., þou spekest , "thou speakest"), and the third person singular in -eþ (e.g., he comeþ , "he cometh/he comes"). ( þ (the letter "thorn") is pronounced like the unvoiced th in "think", but under certain circumstances, it may be like the voiced th in "that"). The following table illustrates a typical conjugation pattern: Plural forms vary strongly by dialect, with Southern dialects preserving
9152-415: The shallow gravel beds of freshwater headstreams and spend their juvenile years in rivers , lakes and freshwater wetlands , migrate to the ocean as adults and live like sea fish , then return to their freshwater birthplace to reproduce . However, populations of several species are restricted to fresh waters (i.e. landlocked) throughout their lives. Folklore has it that the fish return to
9256-421: The silvery blue of a fresh-run fish from the sea to a darker colour. Salmon can make amazing journeys, sometimes moving hundreds of miles upstream against strong currents and rapids to reproduce. Chinook and sockeye salmon from central Idaho, for example, travel over 1,400 km (900 mi) and climb nearly 2,100 m (7,000 ft) from the Pacific Ocean as they return to spawn. Condition tends to deteriorate
9360-546: The skin of wild salmon during free-swimming, planktonic nauplii and copepodid larval stages, which can persist for several days. Large numbers of highly populated, open-net salmon farms can create exceptionally large concentrations of sea lice; when exposed in river estuaries containing large numbers of open-net farms, many young wild salmon are infected, and do not survive as a result. Adult salmon may survive otherwise critical numbers of sea lice, but small, thin-skinned juvenile salmon migrating to sea are highly vulnerable. On
9464-458: The stream. Also, along with species like rainbow trout and Sacramento sucker , lampreys clean the gravel in the rivers during spawning. Their larvae, called ammocoetes, are filter feeders which contribute to the health of the waters. They are also a food source for the young salmon, and being fattier and oilier, it is assumed predators prefer them over salmon offspring, taking off some of the predation pressure on smolts. Adult lampreys are also
9568-409: The strong type have an -e in the nominative/accusative singular, like the weak declension, but otherwise strong endings. Often, these are the same nouns that had an -e in the nominative/accusative singular of Old English (they, in turn, were inherited from Proto-Germanic ja -stem and i -stem nouns). The distinct dative case was lost in early Middle English, and although the genitive survived, by
9672-534: The survivability of other wild predatory fish which rely on them for food. Research is ongoing into sustainable and plant-based salmon feeds. Intensive salmon farming uses open-net cages, which have low production costs. It has the drawback of allowing disease and sea lice to spread to local wild salmon stocks. Another form of salmon production, which is safer but less controllable, is to raise salmon in hatcheries until they are old enough to become independent. They are released into rivers in an attempt to increase
9776-496: The time the studies were conducted, stocks from the middle and upper reaches of large river systems in British Columbia such as Fraser , Skeena , Nass and from mainland coastal streams in the southern half of B.C., "are more likely to have a low prevalence of infection." The report also states, "It should be stressed that Henneguya , economically deleterious though it is, is harmless from the view of public health . It
9880-440: The top levels of the English-speaking political and ecclesiastical hierarchies by Norman rulers who spoke a dialect of Old French , now known as Old Norman , which developed in England into Anglo-Norman . The use of Norman as the preferred language of literature and polite discourse fundamentally altered the role of Old English in education and administration, even though many Normans of this period were illiterate and depended on
9984-467: The total nitrogen available to the riparian woodlands . The foliage of spruce trees up to 500 m (1,600 ft) from a stream where grizzlies fish salmon have been found to contain nitrogen originating from the fished salmon. Beavers also function as ecosystem engineers; in the process of tree-cutting and damming , beavers alter the local ecosystems extensively. Beaver ponds can provide critical habitat for juvenile salmon . An example of this
10088-597: The use of wild-caught fish as feed for the salmon, is the use of soy -based products. This should be better for the local environment of the fish farm, but producing soy beans has a high environmental cost for the producing region. The fish omega-3 fatty acid content would be reduced compared to fish-fed salmon. Another possible alternative is a yeast-based coproduct of bioethanol production, proteinaceous fermentation biomass. Substituting such products for engineered feed can result in equal (sometimes enhanced) growth in fish. With its increasing availability, this would address
10192-417: The water to attack terrestrial insects such as grasshoppers and dragonflies , as well as consuming fish eggs (even those of other salmon). As adults, salmon behave like other mid-sized pelagic fish , eating a variety of sea creatures including smaller forage fish such as lanternfish , herrings , sand lances , mackerels and barracudina . They also eat krill , squid and polychaete worms . In
10296-581: The world, with Norway being the world's largest producer of farmed salmon, followed by Chile . They are also highly prized game fish for recreational fishing , by both freshwater and saltwater anglers . Many species of salmon have since been introduced and naturalized into non-native environments such as the Great Lakes of North America , Patagonia in South America and South Island of New Zealand . The Modern English term salmon
10400-425: Was indicated by agreement of articles and pronouns (e.g., þo ule "the feminine owl") or using the pronoun he to refer to masculine nouns such as helm ("helmet"), or phrases such as scaft stærcne (strong shaft), with the masculine accusative adjective ending -ne . Single-syllable adjectives added -e when modifying a noun in the plural and when used after the definite article ( þe ), after
10504-520: Was not yet a distinct j , v , or w , and Old English scribes did not generally use k , q , or z . Ash was no longer required in Middle English, as the Old English vowel /æ/ that it represented had merged into /a/ . The symbol nonetheless came to be used as a ligature for the digraph ⟨ae⟩ in many words of Greek or Latin origin, as did ⟨œ⟩ for ⟨oe⟩ . Eth and thorn both represented /θ/ or its allophone / ð / in Old English. Eth fell out of use during
10608-417: Was ousted by it in most dialects by the 15th. The following table shows some of the various Middle English pronouns. Many other variations are noted in Middle English sources because of differences in spellings and pronunciations at different times and in different dialects. As a general rule, the indicative first person singular of verbs in the present tense ended in -e (e.g., ich here , "I hear"),
10712-644: Was seen in the years following 1818 in the Columbia River Basin. In 1818, the British government made an agreement with the U.S. government to allow U.S. citizens access to the Columbia catchment (see Treaty of 1818 ). At the time, the Hudson's Bay Company sent word to trappers to extirpate all furbearers from the area in an effort to make the area less attractive to U.S. fur traders. In response to
10816-475: Was some inflectional simplification (the distinct Old English dual forms were lost), but pronouns, unlike nouns, retained distinct nominative and accusative forms. Third person pronouns also retained a distinction between accusative and dative forms, but that was gradually lost: The masculine hine was replaced by him south of the River Thames by the early 14th century, and the neuter dative him
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