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Strombidae

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In biology , a common name of a taxon or organism (also known as a vernacular name , English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrasted with the scientific name for the same organism, which is often based in Latin . A common name is sometimes frequently used, but that is not always the case.

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59-719: See text Strombidae , commonly known as the true conchs , is a taxonomic family of medium-sized to very large sea snails in the superfamily Stromboidea , and the Epifamily Neostromboidae. The term true conchs , being a common name, does not have an exact meaning. It may generally refer to any of the Strombidae but sometimes is used more specifically to include only Strombus and Lambis . The family currently includes 26 extant , and 10 extinct genera. Strombid gastropods live mainly in tropical and subtropical waters. These animals are widespread in

118-774: A flora of his homeland Sweden, Flora Svecica (1745), and in this, he recorded the Swedish common names, region by region, as well as the scientific names. The Swedish common names were all binomials (e.g. plant no. 84 Råg-losta and plant no. 85 Ren-losta); the vernacular binomial system thus preceded his scientific binomial system. Linnaean authority William T. Stearn said: By the introduction of his binomial system of nomenclature, Linnaeus gave plants and animals an essentially Latin nomenclature like vernacular nomenclature in style but linked to published, and hence relatively stable and verifiable, scientific concepts and thus suitable for international use. The geographic range over which

177-431: A percentage divergence , by dividing the number of substitutions by the number of base pairs analysed: the hope is that this measure will be independent of the location and length of the section of DNA that is sequenced. An older and superseded approach was to determine the divergences between the genotypes of individuals by DNA–DNA hybridization . The advantage claimed for using hybridization rather than gene sequencing

236-530: A common name as one that, although it unambiguously defines a chemical, does not follow the current systematic naming convention, such as acetone , systematically 2-propanone , while a vernacular name describes one used in a lab, trade or industry that does not unambiguously describe a single chemical, such as copper sulfate , which may refer to either copper(I) sulfate or copper(II) sulfate. Sometimes common names are created by authorities on one particular subject, in an attempt to make it possible for members of

295-524: A constant rate of mutation, provide a molecular clock for dating divergence. Molecular phylogeny uses such data to build a "relationship tree" that shows the probable evolution of various organisms. With the invention of Sanger sequencing in 1977, it became possible to isolate and identify these molecular structures. High-throughput sequencing may also be used to obtain the transcriptome of an organism, allowing inference of phylogenetic relationships using transcriptomic data . The most common approach

354-427: A particular species or in a group of related species, it has been found empirically that only a minority of sites show any variation at all, and most of the variations that are found are correlated, so that the number of distinct haplotypes that are found is relatively small. In a molecular systematic analysis, the haplotypes are determined for a defined area of genetic material ; a substantial sample of individuals of

413-485: A particularly common name is used varies; some common names have a very local application, while others are virtually universal within a particular language. Some such names even apply across ranges of languages; the word for cat , for instance, is easily recognizable in most Germanic and many Romance languages . Many vernacular names, however, are restricted to a single country and colloquial names to local districts. Some languages also have more than one common name for

472-547: A phylogenetic tree. The third stage includes different models of DNA and amino acid substitution. Several models of substitution exist. A few examples include Hamming distance , the Jukes and Cantor one-parameter model, and the Kimura two-parameter model (see Models of DNA evolution ). The fourth stage consists of various methods of tree building, including distance-based and character-based methods. The normalized Hamming distance and

531-641: A significant complication to molecular systematics, indicating that different genes within the same organism can have different phylogenies. HGTs can be detected and excluded using a number of phylogenetic methods (see Inferring horizontal gene transfer § Explicit phylogenetic methods ). In addition, molecular phylogenies are sensitive to the assumptions and models that go into making them. Firstly, sequences must be aligned; then, issues such as long-branch attraction , saturation , and taxon sampling problems must be addressed. This means that strikingly different results can be obtained by applying different models to

590-672: A strombid has a long and narrow aperture and a siphonal canal . The shell margin has an indentation near the anterior end which accommodates one of the eye stalks. This indentation is called a strombid or stromboid notch . The stromboid notch may be more or less conspicuous, depending on the species. The shells of most species in this family grow a flared lip upon reaching sexual maturity. They lay eggs in long, gelatinous strands. The genera Strombus and Lambis have many similarities between them, both anatomical and reproductive, though their shells show some conspicuous differences. Strombids were widely accepted as carnivores by several authors in

649-1055: A synonym of Lobatus ), in this case, included Eustrombus gigas (now considered a synonym of Lobatus gigas ) and Eustrombus goliath (= Lobatus goliath ); similarly, the genus Aliger included Aliger costatus (= Lobatus costatus ) and Aliger gallus (= Lobatus gallus ). Strombus gallus Strombus gigas Strombus costatus Strombus raninus Strombus peruvianus Strombus galeatus Strombus latus Strombus pugilis Strombus alatus Strombus gracilior Strombus granulatus Strombus bulla Strombus aurisdianae Strombus vomer Strombus gibberulus Strombus luhuanus Lambis chiragra Lambis truncata Lambis lambis Strombus taurus Strombus sinuatus Strombus maculatus Strombus mutabilis Strombus microurceus Strombus labiatus Strombus fragilis Strombus urceus Strombus dentatus Strombus canarium Strombus vittatus Common name In chemistry , IUPAC defines

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708-575: Is a simple method; however, it is less accurate than the neighbor-joining approach. Finally, the last step comprises evaluating the trees. This assessment of accuracy is composed of consistency, efficiency, and robustness. MEGA (molecular evolutionary genetics analysis) is an analysis software that is user-friendly and free to download and use. This software is capable of analyzing both distance-based and character-based tree methodologies. MEGA also contains several options one may choose to utilize, such as heuristic approaches and bootstrapping. Bootstrapping

767-442: Is an approach that is commonly used to measure the robustness of topology in a phylogenetic tree, which demonstrates the percentage each clade is supported after numerous replicates. In general, a value greater than 70% is considered significant. The flow chart displayed on the right visually demonstrates the order of the five stages of Pevsner's molecular phylogenetic analysis technique that have been described. Molecular systematics

826-404: Is an essentially cladistic approach: it assumes that classification must correspond to phylogenetic descent, and that all valid taxa must be monophyletic . This is a limitation when attempting to determine the optimal tree(s), which often involves bisecting and reconnecting portions of the phylogenetic tree(s). The recent discovery of extensive horizontal gene transfer among organisms provides

885-415: Is available at Nature Protocol. Another molecular phylogenetic analysis technique has been described by Pevsner and shall be summarized in the sentences to follow (Pevsner, 2015). A phylogenetic analysis typically consists of five major steps. The first stage comprises sequence acquisition. The following step consists of performing a multiple sequence alignment, which is the fundamental basis of constructing

944-482: Is examined in order to see whether the samples cluster in the way that would be expected from current ideas about the taxonomy of the group. Any group of haplotypes that are all more similar to one another than any of them is to any other haplotype may be said to constitute a clade , which may be visually represented as the figure displayed on the right demonstrates. Statistical techniques such as bootstrapping and jackknifing help in providing reliability estimates for

1003-461: Is in these remarks from a book on marine fish: In scientific binomial nomenclature, names commonly are derived from classical or modern Latin or Greek or Latinised forms of vernacular words or coinages; such names generally are difficult for laymen to learn, remember, and pronounce and so, in such books as field guides, biologists commonly publish lists of coined common names. Many examples of such common names simply are attempts to translate

1062-425: Is possible to determine the processes by which diversity among species has been achieved. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree . Molecular phylogenetics is one aspect of molecular systematics , a broader term that also includes the use of molecular data in taxonomy and biogeography . Molecular phylogenetics and molecular evolution correlate. Molecular evolution

1121-634: Is significant value to understanding evolutionary processes within Stromboidea to recognise the universal similarity in the position of the eye on the end of peduncles and a diminished cephalic tentacle that arises from the middle to the end on that peduncle. This is in contrast to other members of the Stromboidea where the eye is located at the base of the cephalic tentacle. (from Maxwell et al. 2019) ( https://www.researchgate.net/publication/334746905_Recognising_and_defining_a_new_crown_clade_within_Stromboidea_Rafinesque_1815_Mollusca_Gastropoda ) Historically

1180-659: Is the Cape dikkop (or "gewone dikkop", not to mention the presumably much older Zulu name "umBangaqhwa"); Burhinus vermiculatus is the "water dikkop". The thick joints in question are not even, in fact, the birds' knees, but the intertarsal joints —in lay terms the ankles. Furthermore, not all species in the genus have "thick knees", so the thickness of the "knees" of some species is not of clearly descriptive significance. The family Burhinidae has members that have various common names even in English, including " stone curlews ", so

1239-482: Is the comparison of homologous sequences for genes using sequence alignment techniques to identify similarity. Another application of molecular phylogeny is in DNA barcoding , wherein the species of an individual organism is identified using small sections of mitochondrial DNA or chloroplast DNA . Another application of the techniques that make this possible can be seen in the very limited field of human genetics, such as

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1298-406: Is the process of selective changes (mutations) at a molecular level (genes, proteins, etc.) throughout various branches in the tree of life (evolution). Molecular phylogenetics makes inferences of the evolutionary relationships that arise due to molecular evolution and results in the construction of a phylogenetic tree. The theoretical frameworks for molecular systematics were laid in the 1960s in

1357-552: Is very characteristic of each species. Usually, large strombid gastropods, such as the queen conch Eustrombus gigas and the spider conch Lambis lambis , do not bury themselves, except during their juvenile stages. However, smaller species such as Strombus canarium and Strombus epidromis may bury themselves even after adulthood. For a long time, all conchs and their allies (the strombids) were classified in only two genera, namely Strombus and Lambis . This classification can still be found in many textbooks and on websites on

1416-519: The ICZN has formal rules for biological nomenclature and convenes periodic international meetings to further that purpose. The form of scientific names for organisms, called binomial nomenclature , is superficially similar to the noun-adjective form of vernacular names or common names which were used by non-modern cultures. A collective name such as owl was made more precise by the addition of an adjective such as screech . Linnaeus himself published

1475-619: The Indo-West Pacific , where most species and genera occur. Nearly 40 of the living species that used to belong to the genus Strombus can be found in the Indo-Pacific region. They also occur in the eastern Pacific and Western Atlantic , and a single species can be found on the African Atlantic coast. Six species of strombids are found in the wider Caribbean region, including the queen conch Aliger gigas ,

1534-642: The 19th century, an erroneous concept that persisted for several decades into the first half of the 20th century. This ideology was probably born in the writings of Lamarck , who classified strombids alongside other supposedly carnivorous snails, and was copied in this by subsequent authors. However, the many claims of those authors were never supported by the observation of animals feeding in their natural habitat . Nowadays, strombids are known to be specialized herbivores and occasional detritivores . They are usually associated with shallow-water reefs and seagrass meadows . Unlike most snails, which glide slowly across

1593-817: The Hebrew Language publish from time to time short dictionaries of common name in Hebrew for species that occur in Israel or surrounding countries e.g. for Reptilia in 1938, Osteichthyes in 2012, and Odonata in 2015. Molecular phylogeny Molecular phylogenetics ( / m ə ˈ l ɛ k j ʊ l ər ˌ f aɪ l oʊ dʒ ə ˈ n ɛ t ɪ k s , m ɒ -, m oʊ -/ ) is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it

1652-529: The Jukes-Cantor correction formulas provide the degree of divergence and the probability that a nucleotide changes to another, respectively. Common tree-building methods include unweighted pair group method using arithmetic mean ( UPGMA ) and Neighbor joining , which are distance-based methods, Maximum parsimony , which is a character-based method, and Maximum likelihood estimation and Bayesian inference , which are character-based/model-based methods. UPGMA

1711-710: The SSAR switched to an online version with a searchable database. Standardized names for the amphibians and reptiles of Mexico in Spanish and English were first published in 1994, with a revised and updated list published in 2008. A set of guidelines for the creation of English names for birds was published in The Auk in 1978. It gave rise to Birds of the World: Recommended English Names and its Spanish and French companions. The Academy of

1770-534: The Secretariat for the AFNC. SSA is an accredited Standards Australia (Australia's peak non-government standards development organisation) Standards Development The Entomological Society of America maintains a database of official common names of insects, and proposals for new entries must be submitted and reviewed by a formal committee before being added to the listing. Efforts to standardize English names for

1829-415: The Strombidae, Rostellariidae, and Seraphsidae from their sister families Struthiolariidae and Aporrhaidae. There is significant value to understanding evolutionary processes within Stromboidea to recognise the universal similarity in the position of the eye on the end of peduncles and a diminished cephalic tentacle that arises from the middle to the end on that peduncle. This is in contrast to other members of

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1888-481: The Stromboidea where the eye is located at the base of the cephalic tentacle. These physiological differences represent two set of organisms with divergent and independent evolutionary life histories and therefore these differences need to be identifiable within the nomenclature to bring meaning to the way we name things.new crown clade Neostromboidae to separate the Strombidae, Rostellariidae, and Seraphsidae from their sister families Struthiolariidae and Aporrhaidae. There

1947-505: The amphibians and reptiles of North America (north of Mexico) began in the mid-1950s. The dynamic nature of taxonomy necessitates periodical updates and changes in the nomenclature of both scientific and common names. The Society for the Study of Amphibians and Reptiles (SSAR) published an updated list in 1978, largely following the previous established examples, and subsequently published eight revised editions ending in 2017. More recently

2006-515: The author introduced into it so many new English names, that are to be found in no dictionary, and that do not preclude the necessity of learning with what Latin names they are synonymous. A tolerable idea may be given of the danger of too great a multiplicity of vulgar names, by imagining what geography would be, or, for instance, the Post-office administration, supposing every town had a totally different name in every language. Various bodies and

2065-493: The authors of many technical and semi-technical books do not simply adapt existing common names for various organisms; they try to coin (and put into common use) comprehensive, useful, authoritative, and standardised lists of new names. The purpose typically is: Other attempts to reconcile differences between widely separated regions, traditions, and languages, by arbitrarily imposing nomenclature, often reflect narrow perspectives and have unfortunate outcomes. For example, members of

2124-408: The choice of the name "thick-knees" is not easy to defend but is a clear illustration of the hazards of the facile coinage of terminology. For collective nouns for various subjects, see a list of collective nouns (e.g. a flock of sheep, pack of wolves). Some organizations have created official lists of common names, or guidelines for creating common names, hoping to standardize

2183-606: The ever-more-popular use of genetic testing to determine a child's paternity , as well as the emergence of a new branch of criminal forensics focused on evidence known as genetic fingerprinting . There are several methods available for performing a molecular phylogenetic analysis. One method, including a comprehensive step-by-step protocol on constructing a phylogenetic tree, including DNA/Amino Acid contiguous sequence assembly, multiple sequence alignment , model-test (testing best-fitting substitution models), and phylogeny reconstruction using Maximum Likelihood and Bayesian Inference,

2242-419: The exact sequences of nucleotides or bases in either DNA or RNA segments extracted using different techniques. In general, these are considered superior for evolutionary studies, since the actions of evolution are ultimately reflected in the genetic sequences. At present, it is still a long and expensive process to sequence the entire DNA of an organism (its genome ). However, it is quite feasible to determine

2301-464: The general public (including such interested parties as fishermen, farmers, etc.) to be able to refer to one particular species of organism without needing to be able to memorise or pronounce the scientific name. Creating an "official" list of common names can also be an attempt to standardize the use of common names, which can sometimes vary a great deal between one part of a country and another, as well as between one country and another country, even where

2360-522: The genus Burhinus occur in Australia, Southern Africa, Eurasia, and South America. A recent trend in field manuals and bird lists is to use the name " thick-knee " for members of the genus. This, in spite of the fact that the majority of the species occur in non-English-speaking regions and have various common names, not always English. For example, "Dikkop" is the centuries-old South African vernacular name for their two local species: Burhinus capensis

2419-482: The genus Strombus . However, according to Simone, only Strombus gracilior , Strombus alatus , and Strombus pugilis , the type species , remained within Strombus , as they constituted a distinct group based on at least five synapomorphies. The remaining taxa were previously considered as subgenera , and were elevated to genus level by Simone in the end of his analysis. The genus Eustrombus (now considered

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2478-426: The genus Terebellum as the most basal taxon , distinguished from the remaining strombids by 13 synapomorphies, including a rounded foot. Though the genus Tibia was left out of the analysis, Simone regarded it as probably closely related to Terebellum , apparently due to some well known morphological similarities between them. With the exception of Lambis , the remaining taxa were previously allocated within

2537-566: The goliath conch Titanostrombus goliath , the hawk-wing conch Lobatus raninus , the rooster tail conch Aliger gallus , the milk conch Macrostrombus costatus , the West Indian fighting conch Strombus pugilis , and the Florida fighting conch Strombus alatus . Until recently, all of these species were placed in the genus Strombus , but now many species are being moved into new genera. Strombids have long eye stalks. The shell of

2596-555: The internet. Based on molecular phylogeny in addition to an extensively documented fossil record, both genera have been subdivided into several new genera by different authors. The family Strombidae actually comprises 26 extant genera and 10 extinct genera (marked with a dagger †). Terebellum terebellum Canarium urceus Conomurex luhuanus Tricornis raninus Lambis lambis Strombus Eustrombus Aliger The most recent revision of Maxwell 2019 (ZooKeys 867:1-7) new crown clade Neostromboidae to separate

2655-711: The modern (now binding) International Code of Nomenclature for algae, fungi, and plants contains the following: Art. 68. Every friend of science ought to be opposed to the introduction into a modern language of names of plants that are not already there unless they are derived from a Latin botanical name that has undergone but a slight alteration. ... ought the fabrication of names termed vulgar names, totally different from Latin ones, to be proscribed. The public to whom they are addressed derives no advantage from them because they are novelties. Lindley's work, The Vegetable Kingdom, would have been better relished in England had not

2714-645: The phylogenetic relationships among the Strombidae have been mainly accessed on two occasions, using two methods. In a 2005 monograph , Simone proposed a cladogram (a tree of descent) based on an extensive morpho anatomical analysis of representatives of the Aporrhaidae , Strombidae, Xenophoridae , and Struthiolariidae . In his analysis, Simone recognized the Strombidae as a monophyletic taxon supported by 13 synapomorphies (traits that are shared by two or more taxa and their most recent common ancestor ), comprising at least eight distinct genera. He considered

2773-486: The positions of haplotypes within the evolutionary trees. Every living organism contains deoxyribonucleic acid ( DNA ), ribonucleic acid ( RNA ), and proteins . In general, closely related organisms have a high degree of similarity in the molecular structure of these substances, while the molecules of organisms distantly related often show a pattern of dissimilarity. Conserved sequences, such as mitochondrial DNA, are expected to accumulate mutations over time, and assuming

2832-659: The results were not quantitative and did not initially improve on morphological classification, they provided tantalizing hints that long-held notions of the classifications of birds , for example, needed substantial revision. In the period of 1974–1986, DNA–DNA hybridization was the dominant technique used to measure genetic difference. Early attempts at molecular systematics were also termed chemotaxonomy and made use of proteins, enzymes , carbohydrates , and other molecules that were separated and characterized using techniques such as chromatography . These have been replaced in recent times largely by DNA sequencing , which produces

2891-488: The same animal. For example, in Irish, there are many terms that are considered outdated but still well-known for their somewhat humorous and poetic descriptions of animals. w/ literal translations of the poetic terms Common names are used in the writings of both professionals and laymen . Lay people sometimes object to the use of scientific names over common names, but the use of scientific names can be defended, as it

2950-576: The same language is spoken in both places. A common name intrinsically plays a part in a classification of objects, typically an incomplete and informal classification, in which some names are degenerate examples in that they are unique and lack reference to any other name, as is the case with say, ginkgo , okapi , and ratel . Folk taxonomy , which is a classification of objects using common names, has no formal rules and need not be consistent or logical in its assignment of names, so that say, not all flies are called flies (for example Braulidae ,

3009-402: The scientific name into English or some other vernacular. Such translation may be confusing in itself, or confusingly inaccurate, for example, gratiosus does not mean "gracile" and gracilis does not mean "graceful". The practice of coining common names has long been discouraged; de Candolle's Laws of Botanical Nomenclature , 1868, the non-binding recommendations that form the basis of

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3068-476: The sequence of a defined area of a particular chromosome . Typical molecular systematic analyses require the sequencing of around 1000 base pairs . At any location within such a sequence, the bases found in a given position may vary between organisms. The particular sequence found in a given organism is referred to as its haplotype . In principle, since there are four base types, with 1000 base pairs, we could have 4 distinct haplotypes. However, for organisms within

3127-423: The simplest case, the difference between two haplotypes is assessed by counting the number of locations where they have different bases: this is referred to as the number of substitutions (other kinds of differences between haplotypes can also occur, for example, the insertion of a section of nucleic acid in one haplotype that is not present in another). The difference between organisms is usually re-expressed as

3186-407: The so-called "bee lice") and not every animal called a fly is indeed a fly (such as dragonflies and mayflies ). In contrast, scientific or biological nomenclature is a global system that attempts to denote particular organisms or taxa uniquely and definitively , on the assumption that such organisms or taxa are well-defined and generally also have well-defined interrelationships; accordingly

3245-457: The substrate on their feet, strombid gastropods have a characteristic means of locomotion, using their pointed, sickle-shaped, horny operculum to propel themselves forward in a so-called leaping motion. Burrowing behavior, in which an individual sinks itself entirely or partially into the substrate, is also frequent among strombid gastropods. The burrowing process itself, which involves distinct sequential movements and sometimes complex behaviors,

3304-403: The target species or other taxon is used; however, many current studies are based on single individuals. Haplotypes of individuals of closely related, yet different, taxa are also determined. Finally, haplotypes from a smaller number of individuals from a definitely different taxon are determined: these are referred to as an outgroup . The base sequences for the haplotypes are then compared. In

3363-832: The use of common names. For example, the Australian Fish Names List or AFNS was compiled through a process involving work by taxonomic and seafood industry experts, drafted using the CAAB (Codes for Australian Aquatic Biota) taxon management system of the CSIRO , and including input through public and industry consultations by the Australian Fish Names Committee (AFNC). The AFNS has been an official Australian Standard since July 2007 and has existed in draft form (The Australian Fish Names List) since 2001. Seafood Services Australia (SSA) serve as

3422-424: The works of Emile Zuckerkandl , Emanuel Margoliash , Linus Pauling , and Walter M. Fitch . Applications of molecular systematics were pioneered by Charles G. Sibley ( birds ), Herbert C. Dessauer ( herpetology ), and Morris Goodman ( primates ), followed by Allan C. Wilson , Robert K. Selander , and John C. Avise (who studied various groups). Work with protein electrophoresis began around 1956. Although

3481-398: Was that it was based on the entire genotype, rather than on particular sections of DNA. Modern sequence comparison techniques overcome this objection by the use of multiple sequences. Once the divergences between all pairs of samples have been determined, the resulting triangular matrix of differences is submitted to some form of statistical cluster analysis , and the resulting dendrogram

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