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48-547: C. 90 Rhodiola is a genus of perennial plants in the family Crassulaceae that resemble Sedum and other members of the family. Like sedums, Rhodiola species are often called stonecrops. Some authors merge Rhodiola into Sedum . Rhodiola species grow in high-altitude and other cold regions of the Northern Hemisphere . Plants of the World Online gives the number of accepted species as 74,

96-443: A century plant can live for 80 years and grow 30 meters tall before flowering and dying. However, most perennials are polycarpic (or iteroparous ), flowering over many seasons in their lifetime. Perennials invest more resources than annuals into roots, crowns, and other structures that allow them to live from one year to the next. They often have a competitive advantage because they can commence their growth and leaf out earlier in

144-940: A photolyase and the other, a protein involved in nucleotide excision repair ). Perennials that are cultivated include: woody plants like fruit trees grown for their edible fruits; shrubs and trees grown as landscaping ornamentals ; herbaceous food crops like asparagus , rhubarb , strawberries ; and subtropical plants not hardy in colder areas such as tomatoes , eggplant , and coleus (which are treated as annuals in colder areas). Perennials also include plants grown for their flowering and other ornamental value including bulbs (like tulips, narcissus, and gladiolus); lawn grass, and other groundcovers , (such as periwinkle and Dichondra ). Each type of plant must be separated differently; for example, plants with fibrous root systems like daylilies, Siberian iris, or grasses can be pried apart with two garden forks inserted back to back, or cut by knives. However, plants such as bearded irises have

192-428: A climate as "temperate" C , when the mean temperature is above −3 °C (26.6 °F) but below 18 °C (64.4 °F) in the coldest month to account for the persistence of frost. However, some adaptations of Köppen set the minimum at 0 °C (32.0 °F). Continental climates are classified as D and considered to be varieties of temperate climates, having more extreme temperatures, with mean temperatures in

240-424: A distinct four-season pattern, especially in the continental climate sector. Oceanic climates are created by the on-shore flow from the cool high latitude oceans to their west. This causes the climate to have mild summers and cool (but not cold) winters, and relative humidity and precipitation evenly distributed throughout the year. These climates are frequently cloudy and cool, and winters are milder than those in

288-809: A few highland areas of Tasmania , the Australian Alps and Southern Alps of New Zealand . This type of climate is even found in tropical areas such as the Papuan Highlands in Indonesia. Cfc is the categorization for this regime. Even in the middle of summer, temperatures exceeding 20°C (68 °F) are exceptional weather events in the most maritime of those locations impacted by this regime. In some parts of this climate, temperatures as high as 30°C (86°F) have been recorded on rare occasions, while temperatures as low as −15 °C (5 °F) have still been recorded on rare occasions. A cold variant of

336-421: A huge impact on the productivity of agriculture which is less common. Temperate regions have the majority of the world's population, which leads to large cities . There are a couple of factors why the climate of large city landscapes differs from the climate of rural areas. One factor is the strength of the absorption rate of buildings and asphalt, which is higher than that of natural land. The other large factor

384-434: A humid subtropical climate (most notably southeast China and North India ), there is an even sharper wet-dry season, called a monsoon subtropical climate or subtropical monsoon (Cwa). In these regions, winters are quite chilly and dry and summers have very heavy rainfall. Some Cwa areas in southern China report more than 80% of annual precipitation in the five warmest months (southwest monsoon). Mediterranean climates have

432-470: A root system of rhizomes; these root systems should be planted with the top of the rhizome just above ground level, with leaves from the following year showing. The point of dividing perennials is to increase the amount of a single breed of plant in your garden. In the United States more than 900 million dollars worth of potted herbaceous perennial plants were sold in 2019. Although most of humanity

480-469: A summer accent on rainfall. Regions with humid continental climates include southeastern Canada, the upper portions of the eastern United States , portions of eastern Europe , parts of China , Japan and the Korean Peninsula . These are temperate climates that compared to the subtropics are on the poleward edge of the temperate zone. Therefore, they still have four marked seasons including

528-435: A temperate region is found in southern Africa , where some 24,000 taxa (species and infraspecific taxa) have been described. Farming is a large-scale practice in the temperate regions (except for boreal/subarctic regions) due to the plentiful rainfall and warm summers, because most agricultural activity occurs in the spring and summer, cold winters have a small effect on agricultural production. Extreme winters or summers have

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576-479: A very few (e.g. Zostera ) occurring in shallow sea water. Herbaceous perennial plants are particularly dominant in conditions too fire-prone for trees and shrubs, e.g., most plants on prairies and steppes are perennials; they are also dominant on tundra too cold for tree growth. Nearly all forest plants are perennials, including trees and shrubs. Perennial plants are usually better long-term competitors, especially under stable, resource-poor conditions. This

624-646: A warmer one, but are far more influenced by the polar zones than any other but the very polar climates ( tundra and ice cap climate ). Areas with subpolar oceanic climates feature an oceanic climate but are usually located closer to polar regions. As a result of their location, these regions tend to be on the cool end of oceanic climates. Snowfall tends to be more common here than in other oceanic climates. Subpolar oceanic climates are less prone to temperature extremes than subarctic climates or continental climates , featuring milder winters than these climates but still with similar summers. This variant of an oceanic climate

672-467: Is difficult outside their native subarctic and alpine climates. In traditional Chinese medicine , Rhodiola is used for fatigue, hypoxia, radiation, etc. Perennial plant Perennials (especially small flowering plants ) that grow and bloom over the spring and summer, die back every autumn and winter, and then return in the spring from their rootstock or other overwintering structure, are known as herbaceous perennials . However, depending on

720-783: Is due to the development of larger root systems which can access water and soil nutrients deeper in the soil and to earlier emergence in the spring. Annual plants have an advantage in disturbed environments because of their faster growth and reproduction rates. Each section contains a short list of species related to that topic, these are an example as the true lists would fill several books. Perennials grown for their decorative flowers include very many species and types. Some examples include: The majority of fruit bearing plants are perennial even in temperate climates. Examples include: Many herbs are perennial, including these examples: Many vegetable plants can grow as perennials in tropical climates, but die in cold weather. Examples of some of

768-483: Is fed by the re-sowing of the seeds of annual grain crops, (either naturally or by the manual efforts of humans), perennial crops provide numerous benefits. Perennial plants often have deep, extensive root systems which can hold soil to prevent erosion , capture dissolved nitrogen before it can contaminate ground and surface water, and out-compete weeds (reducing the need for herbicides ). These potential benefits of perennials have resulted in new attempts to increase

816-626: Is found in parts of coastal Iceland , the Faroe Islands , parts of Scotland , northwestern coastal areas of Norway such as Lofoten and reaching to 70° north on some islands, uplands near the coast of southwestern Norway, the Aleutian Islands of Alaska and northern parts of the Alaskan Panhandle , some parts of Southern Argentina and Chile (though most regions are still classified as continental subantarctic), and

864-429: Is too cold or dry. In many parts of the world, seasonality is expressed as wet and dry periods rather than warm and cold periods, and deciduous perennials lose their leaves in the dry season. Some perennial plants are protected from wildfires because they have underground roots that produce adventitious shoots, bulbs, crowns , or stems ; other perennials like trees and shrubs may have thick cork layers that protect

912-611: The Angiosperm Phylogeny Website gives it as 90, and the Flora of China gives it as about 90, with 55 in China and 16 endemic there. Flora of North America lists only three species in the United States and Canada. Among the distinguishing characters of the genus are two series of stamens totaling twice the number of petals ; free or nearly free petals (not joined in a tube); a stout rhizome from whose axils

960-451: The Köppen climate classification . In contrast to oceanic climates, they are created by large land masses and seasonal changes in wind direction. This causes humid continental climates to have severe temperatures for the season compared to other temperate climates, meaning a hot summer and cold winter. Precipitation may be evenly distributed throughout the year, while in some locations there is

1008-475: The astronomical symbol for the planet Jupiter . Perennial plants can be short-lived (only a few years) or long-lived. They include a wide assortment of plant groups from non-flowering plants like ferns and liverworts to highly diverse flowering plants like orchids , grasses , and woody plants . Plants that flower and fruit only once and then die are termed monocarpic or semelparous ; these species may live for many years before they flower. For example,

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1056-513: The Cwb classification have significant monsoon influence, usually having dry winters and wet summers. These climates occur in the middle latitudes, between approximately 35° and 66.5° north and south of the equator. There is an equal climatic influence from both the polar and tropical zones in this climate region. Two types of climates are in this zone, a milder oceanic one and more severe seasonal continental one. Most prototypical temperate climates have

1104-446: The advantage of generating larger seedlings that can better compete with other plants. Perennials also produce seeds over many years. An important aspect of cold acclimation is overexpression of DNA repair genes. In Thinopyrum intermedium a perennial relative of common wheat Triticum aestivum , conditions of freezing stress were shown to be associated with large increases in expression of two DNA repair genes (one gene product

1152-521: The basis of well developed rhizomes and annual flowering stems , arising from axils of the scaly radical leaves. This separation was subsequently confirmed by molecular phylogenetic studies. Rhodiola is placed within family Crassulaceae , in subfamily Sempervivoideae , tribe Umbiliceae. There it is a sister group to Pseudosedum , though some authors have suggested that the latter genus be submersed within Rhodiola . Traditionally Rhodiola

1200-614: The coldest month usually being below −3 °C (26.6 °F). The north temperate zone extends from the Tropic of Cancer (approximately 23.5° north latitude) to the Arctic Circle (approximately 66.5° north latitude). The south temperate zone extends from the Tropic of Capricorn (approximately 23.5° south latitude) to the Antarctic Circle (at approximately 66.5° south latitude). In some climate classifications,

1248-410: The continental climate. Regions with oceanic climates include northwestern Europe , northwestern North America , southeastern and southwestern South America , southeastern Australia and most of New Zealand . Humid continental climates are considered as a variety of temperate climates due to lying in the temperate zones, although they are classified separately from other temperate climates in

1296-510: The dormancy period is over and new growth begins. In climates that are warm all year long, perennials may grow continuously. Annuals which complete their life cycle in one growing season, in contrast with perennials, produce seeds as the next generation and die; the seeds may survive cold or dry periods or germinate soon after dispersal depending on the climate. Some perennials retain their foliage year-round; these are evergreen perennials. Deciduous perennials shed all their leaves part of

1344-409: The flowering stems rise; and a basal rosette of leaves. This genus contains the only species of Crassulaceae that have unisexual flowers. Rhodionin is a herbacetin rhamnoside found in Rhodiola species. Although Linnaeus distinguished Rhodiola from Sedum on the basis of being dioecious , it was later submerged in the latter genus until the twentieth century, when it was restored, on

1392-450: The genus, and reversals to a hermaphrodite condition have also occurred, which is a rare occurrence in flowering plants. It has been suggested that dioecy in the genus may correlate with abiotic pollination in the cold environment. Rhodiola is sold to promote a variety of uses, specifically in physical health, with the plant being used historically in medicine throughout Europe. A number of species are grown as ornamentals, but growing them

1440-541: The growing season, and can grow taller than annuals. In doing so they can better compete for space and collect more light. Perennials typically grow structures that allow them to adapt to living from one year to the next through a form of vegetative reproduction rather than seeding. These structures include bulbs , tubers , woody crowns, rhizomes , turions , woody stems, or crowns which allows them to survive periods of dormancy over cold or dry seasons; these structures typically store carbohydrates which are used once

1488-620: The hormones produced due to environmental situations (i.e., seasons), reproduction, and stage of development to begin and halt the ability to grow or flower. There is also a distinction between the ability to grow and the actual task of growth. For example, most trees regain the ability to grow during winter but do not initiate physical growth until the spring and summer months. The start of dormancy can be seen in perennial plants through withering flowers, loss of leaves on trees, and halting of reproduction in both flowering and budding plants. Perennial species may produce relatively large seeds that have

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1536-476: The humid subtropics. Warm ocean currents are usually found in coastal areas with humid subtropical climates. This type of climate is normally located along leeward lower east coasts of continents such as in southeast and central Argentina , Uruguay and south of Brazil , the southeast portions of East Asia , southern and portions of the northeast and midwestern United States and portions of, South Africa , Ethiopia , and eastern Australia . In some areas with

1584-547: The monsoon-influenced subtropical highland climate similar to subpolar oceanic climates occurs in small areas in Yunnan, Sichuan and parts of the Altiplano between Bolivia , Peru and Chile , where summers are sufficiently short to be Cwc with fewer than four months over 10 °C (50 °F) due to the high altitudes at these locations. El Alto, Bolivia , is one of the few confirmed towns that features this variation of

1632-430: The more completely perennial vegetables are: Many aquatic plants are perennial even though many do not have woody tissue. Examples include: Temperate In geography , the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth . These zones generally have wider temperature ranges throughout

1680-517: The opposite rainfall pattern to dry-winter climates, with a dry summer and wet winter. This climate occurs mostly at the western edges and coasts of the continents and are bounded by arid deserts on their equatorward sides that brings dry winds causing the dry season of summer, and oceanic climates to the poleward sides that are influenced by cool ocean currents and air masses that bring the rainfall of winter. The five main Mediterranean regions of

1728-580: The opposite rainfall pattern. This climate is rare and is predominantly found in climate fringes and isolated areas of the Cascades and Andes Mountains , as the dry-summer climate extends further poleward in the Americas than elsewhere. The vast majority of the world's human population resides in temperate zones, especially in the Northern Hemisphere , due to its greater mass of land and lack of extreme temperatures. The biggest described number of taxa in

1776-461: The rigours of the local climate (temperature, moisture, organic content in the soil, microorganisms), a plant that is a perennial in its native habitat, may be treated by a gardener as an annual and planted out every year, from seed, from cuttings, or from divisions. Tomato vines, for example, live several years in their natural tropical/ subtropical habitat but are grown as annuals in temperate regions because their above-ground biomass does not survive

1824-465: The seed yield of perennial species, which could result in the creation of new perennial grain crops. Some examples of new perennial crops being developed are perennial rice and intermediate wheatgrass . A perennial rice developed in 2018, was reported in 2023, to have provided a similar yield to replanted annual rice when evaluated over eight consecutive harvests. Perennial plants dominate many natural ecosystems on land and in fresh water, with only

1872-431: The stems. Herbaceous perennials from temperate and alpine regions of the world can tolerate the cold during winter. Perennial plants may remain dormant for long periods and then recommence growth and reproduction when the environment is more suitable, while most annual plants complete their life cycle during one growing period, and biennials have two growing periods. The meristem of perennial plants communicates with

1920-509: The subtropical highland climate. Cold summer mediterranean climates ( Csc ) are present in high-elevation areas around coastal Csb climate areas, where the strong maritime influence prevents the average winter monthly temperature from dropping below 0 °C. Despite the maritime influence, they are classified alongside other mediterranean climates in the Köppen classification rather than oceanic climates like subtropical highland climates due to

1968-449: The temperate zone may be divided into several smaller climate zones, based on monthly temperatures, the coldest month, and rainfall. These can include the subtropical zone ( humid subtropical and Mediterranean climate ), and the cool temperate zone ( oceanic and continental climates ). These climates are typically found in the more equatorial regions of the temperate zone, between 23.5° and 35° north or south. They are influenced more by

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2016-472: The tropics or subtropics. They have characteristically mild temperatures year-round, featuring the four seasons in the subtropics and no marked seasons in the tropics, the latter usually remaining mild to cool through most of the year. Subtropical highland climates under the Cfb classification usually have rainfall spread relatively evenly in all months of the year similar to most oceanic climates while climates under

2064-458: The tropics than by other temperate climate types, usually experiencing warmer temperatures throughout the year, with longer, hotter summers and shorter, milder winters. Freezing precipitation is uncommon in this part of the temperate zone. Humid subtropical climates generally have long, hot and humid summers with frequent convective showers in summer, and a peak seasonal rainfall in the hottest months. Winters are normally mild and above freezing in

2112-405: The winter. There is also a class of evergreen perennials which lack woody stems, such as Bergenia which retain a mantle of leaves throughout the year. An intermediate class of plants is known as subshrubs , which retain a vestigial woody structure in winter, e.g. Penstemon . The symbol for a perennial plant, based on Species Plantarum by Linnaeus , is [REDACTED] , which is also

2160-717: The world are the Mediterranean Basin in North Africa , Southern Europe , and West Asia , coastal California in the United States , the South and West states of Australia , the Western Cape of South Africa and the south and southwestern coast of Chile . Subtropical highland climates are climate variants often grouped together with oceanic climates found in some mountainous areas of either

2208-454: The year and more distinct seasonal changes compared to tropical climates , where such variations are often small; they usually differ only in the amount of precipitation . In temperate climates, not only do latitudinal positions influence temperature changes, but various sea currents, prevailing wind direction, continentality (how large a landmass is) and altitude also shape temperate climates. The Köppen climate classification defines

2256-439: The year. Deciduous perennials include herbaceous and woody plants; herbaceous plants have stems that lack hard, fibrous growth, while woody plants have stems with buds that survive above ground during dormancy. Some perennials are semi-deciduous, meaning they lose some of their leaves in either winter or summer. Deciduous perennials shed their leaves when growing conditions are no longer suitable for photosynthesis, such as when it

2304-620: Was divided into subgenera, sections and series, based on plant characteristics. Four subgenera were recognised; Rhodiola , Primuloides , Crassipedes and Clementsia . However molecular studies have failed to demonstrate monophyly of these subtaxa. Species include: The name combines the Greek rhodon , meaning rose and referring to the rose-like smell of the roots, with the Latin diminutive suffix -iola . Dioecy , having separate male and female flowers, has evolved at least three times in

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