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105-445: Ricinus communis , the castor bean or castor oil plant , is a species of perennial flowering plant in the spurge family, Euphorbiaceae . It is the sole species in the monotypic genus , Ricinus , and subtribe , Ricininae . The evolution of castor and its relation to other species are currently being studied using modern genetic tools. It reproduces with a mixed pollination system which favors selfing by geitonogamy but at

210-443: A century plant can live for 80 years and grow 30 meters tall before flowering and dying. However, most perennials are polycarpic (or iteroparous ), flowering over many seasons in their lifetime. Perennials invest more resources than annuals into roots, crowns, and other structures that allow them to live from one year to the next. They often have a competitive advantage because they can commence their growth and leaf out earlier in

315-564: A laxative . The use of castor bean oil ( eranda ) in India has been documented since 2000 BC in lamps and in local medicine as a laxative, purgative , and cathartic in Unani , Ayurvedic , siddha and other ethnomedical systems. Traditional Ayurvedic and siddha medicine considers castor oil the king of medicinals for curing arthritic diseases . It is regularly given to children to treat infections with parasitic worms. The ancient Romans had

420-940: A photolyase and the other, a protein involved in nucleotide excision repair ). Perennials that are cultivated include: woody plants like fruit trees grown for their edible fruits; shrubs and trees grown as landscaping ornamentals ; herbaceous food crops like asparagus , rhubarb , strawberries ; and subtropical plants not hardy in colder areas such as tomatoes , eggplant , and coleus (which are treated as annuals in colder areas). Perennials also include plants grown for their flowering and other ornamental value including bulbs (like tulips, narcissus, and gladiolus); lawn grass, and other groundcovers , (such as periwinkle and Dichondra ). Each type of plant must be separated differently; for example, plants with fibrous root systems like daylilies, Siberian iris, or grasses can be pried apart with two garden forks inserted back to back, or cut by knives. However, plants such as bearded irises have

525-548: A "disagreement" with Rudbeck's wife and had to move out of his mentor's house; his relationship with Rudbeck did not appear to suffer. That Christmas, Linnaeus returned home to Stenbrohult to visit his parents for the first time in about three years. His mother had disapproved of his failing to become a priest, but she was pleased to learn he was teaching at the University. During a visit with his parents, Linnaeus told them about his plan to travel to Lapland ; Rudbeck had made

630-483: A catalogue of the botanical holdings in the herbarium and botanical garden of Hartekamp. He wrote it in nine months (completed in July 1737), but it was not published until 1738. It contains the first use of the name Nepenthes , which Linnaeus used to describe a genus of pitcher plants . Linnaeus stayed with Clifford at Hartekamp until 18 October 1737 (new style), when he left the house to return to Sweden. Illness and

735-475: A child's talents than develop them". Two years after his tutoring had begun, he was sent to the Lower Grammar School at Växjö in 1717. Linnaeus rarely studied, often going to the countryside to look for plants. At some point, his father went to visit him and, after hearing critical assessments by his preceptors, he decided to put the youth as an apprentice to some honest cobbler. He reached

840-659: A collector of natural history, and to see his cabinet , as well as to visit the Chelsea Physic Garden and its keeper, Philip Miller . He taught Miller about his new system of subdividing plants, as described in Systema Naturae . At first, Miller was reluctant to use the new binomial nomenclature , preferring instead the classifications of Joseph Pitton de Tournefort and John Ray . Nevertheless, Linnaeus applauded Miller's Gardeners Dictionary . The conservative Miller actually retained in his dictionary

945-430: A copy of Sir Hans Sloane's Natural History of Jamaica , a rare book, if he let Linnaeus stay with him, and Burman accepted. On 24 September 1735, Linnaeus moved to Hartekamp to become personal physician to Clifford, and curator of Clifford's herbarium. He was paid 1,000 florins a year, with free board and lodging. Though the agreement was only for a winter of that year, Linnaeus practically stayed there until 1738. It

1050-559: A curriculum designed for boys preparing for the priesthood. In the last year at the gymnasium, Linnaeus's father visited to ask the professors how his son's studies were progressing; to his dismay, most said that the boy would never become a scholar. Rothman believed otherwise, suggesting Linnaeus could have a future in medicine. The doctor offered to have Linnaeus live with his family in Växjö and to teach him physiology and botany. Nils accepted this offer. Rothman showed Linnaeus that botany

1155-443: A dark green, sometimes with a reddish tinge, as they mature. The leaves of some other varieties are green practically from the beginning, whereas in yet others a pigment masks the green color of all the chlorophyll -bearing parts, leaves, stems and young fruit, so that they remain a dramatic purple-to-reddish-brown throughout the life of the plant. Plants with the dark leaves can be found growing next to those with green leaves, so there

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1260-575: A director of the Dutch East India Company and the owner of a rich botanical garden at the estate of Hartekamp in Heemstede . Clifford was very impressed with Linnaeus's ability to classify plants, and invited him to become his physician and superintendent of his garden. Linnaeus had already agreed to stay with Burman over the winter, and could thus not accept immediately. However, Clifford offered to compensate Burman by offering him

1365-538: A dissertation, written back in Sweden, entitled Dissertatio medica inauguralis in qua exhibetur hypothesis nova de febrium intermittentium causa , in which he laid out his hypothesis that malaria arose only in areas with clay-rich soils. Although he failed to identify the true source of disease transmission, (i.e., the Anopheles mosquito ), he did correctly predict that Artemisia annua ( wormwood ) would become

1470-456: A family, he received permission to marry his fiancée, Sara Elisabeth Moræa. Their wedding was held 26 June 1739. Seventeen months later, Sara gave birth to their first son, Carl . Two years later, a daughter, Elisabeth Christina , was born, and the subsequent year Sara gave birth to Sara Magdalena, who died when 15 days old. Sara and Linnaeus would later have four other children: Lovisa, Sara Christina, Johannes and Sophia. In May 1741, Linnaeus

1575-474: A height of 2–3 metres (6.6–9.8 ft) in a year. In areas prone to frost it is usually shorter, and grown as if it were an annual . However, it can grow well outdoors in cooler climates, at least in southern England, and the leaves do not appear to suffer frost damage in sheltered spots, where it remains evergreen. It was used in Edwardian times in the parks of Toronto, Canada. Although not cultivated there,

1680-480: A naturalist." Linnaeus has been called Princeps botanicorum (Prince of Botanists) and "The Pliny of the North". He is also considered one of the founders of modern ecology . In botany, the abbreviation L. is used to indicate Linnaeus as the authority for a species' name. In zoology, the abbreviation Linnaeus is generally used; the abbreviations L., Linnæus and Linné are also used. In older publications,

1785-475: A new benefactor, Olof Celsius , who was a professor of theology and an amateur botanist. He received Linnaeus into his home and allowed him use of his library, which was one of the richest botanical libraries in Sweden. In 1729, Linnaeus wrote a thesis, Praeludia Sponsaliorum Plantarum on plant sexual reproduction . This attracted the attention of Rudbeck; in May 1730, he selected Linnaeus to give lectures at

1890-569: A number of pre-Linnaean binomial signifiers discarded by Linnaeus but which have been retained by modern botanists. He only fully changed to the Linnaean system in the edition of The Gardeners Dictionary of 1768. Miller ultimately was impressed, and from then on started to arrange the garden according to Linnaeus's system. Linnaeus also travelled to Oxford University to visit the botanist Johann Jacob Dillenius . He failed to make Dillenius publicly fully accept his new classification system, though

1995-630: A physician, and thus to make it possible to support a family. Once again, Linnaeus found a patron; he became acquainted with Count Carl Gustav Tessin , who helped him get work as a physician at the Admiralty. During this time in Stockholm, Linnaeus helped found the Royal Swedish Academy of Science ; he became the first Praeses of the academy by drawing of lots. Because his finances had improved and were now sufficient to support

2100-561: A practical way, making this the first proto-modern Flora . The account covered 534 species, used the Linnaean classification system and included, for the described species, geographical distribution and taxonomic notes. It was Augustin Pyramus de Candolle who attributed Linnaeus with Flora Lapponica as the first example in the botanical genre of Flora writing. Botanical historian E. L. Greene described Flora Lapponica as "the most classic and delightful" of Linnaeus's works. It

2205-472: A range of cultivars for leaf and flower colours, and for oil production. It is a fast-growing, suckering shrub that can reach the size of a small tree, around 12 metres (39 feet), but it is not cold hardy . The glossy leaves are 15–45 centimetres (6–18 inches) long, long-stalked, alternate and palmate with five to twelve deep lobes with coarsely toothed segments. In some varieties they start off dark reddish purple or bronze when young, gradually changing to

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2310-461: A replacement for castoreum , a perfume base made from the dried perineal glands of the beaver ( castor in Latin). It has another common name, palm of Christ , or Palma Christi , that derives from castor oil's reputed ability to heal wounds and cure ailments. Ricinus communis is the host plant of the common castor butterfly ( Ariadne merione ), the eri silkmoth ( Samia cynthia ricini ), and

2415-470: A root system of rhizomes; these root systems should be planted with the top of the rhizome just above ground level, with leaves from the following year showing. The point of dividing perennials is to increase the amount of a single breed of plant in your garden. In the United States more than 900 million dollars worth of potted herbaceous perennial plants were sold in 2019. Although most of humanity

2520-580: A small group of students to Dalarna . Funded by the Governor of Dalarna, the expedition was to catalogue known natural resources and discover new ones, but also to gather intelligence on Norwegian mining activities at Røros . His relations with Nils Rosén having worsened, Linnaeus accepted an invitation from Claes Sohlberg, son of a mining inspector, to spend the Christmas holiday in Falun , where Linnaeus

2625-462: A source of antimalarial medications. Within two weeks he had completed his oral and practical examinations and was awarded a doctoral degree. That summer Linnaeus reunited with Peter Artedi , a friend from Uppsala with whom he had once made a pact that should either of the two predecease the other, the survivor would finish the decedent's work. Ten weeks later, Artedi drowned in the canals of Amsterdam , leaving behind an unfinished manuscript on

2730-414: A suitable climate, castor establishes itself easily where it can become an invasive plant and can often be found on wasteland. It is also used extensively as a decorative plant in parks and other public areas, particularly as a "dot plant" in traditional bedding schemes . If sown early, under glass, and kept at a temperature of around 20 °C (68 °F) until planted out, the castor oil plant can reach

2835-452: A thesis along with Frederick Lindberg, a physician student, based on their experiences. In the tradition of the period, this dissertation was essentially an idea of the presiding reviewer ( prases ) expounded upon by the student. Linnaeus's dissertation was translated into French by J. E. Gilibert in 1770 as La Nourrice marâtre, ou Dissertation sur les suites funestes du nourrisage mercénaire . Linnaeus suggested that children might absorb

2940-552: A variety of medicinal/cosmetic uses for both the seeds and the leaves of Ricinus communis . The naturalist Pliny the Elder cited the poisonous qualities of the seeds, but mentioned that they could be used to form wicks for oil lamps (possibly if crushed together), and the oil for use as a laxative and lamp oil. He also recommends the use of the leaves as follows: "The leaves are applied topically with vinegar for erysipelas , and fresh-gathered, they are used by themselves for diseases of

3045-479: A very few (e.g. Zostera ) occurring in shallow sea water. Herbaceous perennial plants are particularly dominant in conditions too fire-prone for trees and shrubs, e.g., most plants on prairies and steppes are perennials; they are also dominant on tundra too cold for tree growth. Nearly all forest plants are perennials, including trees and shrubs. Perennial plants are usually better long-term competitors, especially under stable, resource-poor conditions. This

3150-443: A yellow nodule full of fats one end of the seed that are nutritious for young ants. After hauling their harvest into their nests and pulling off the delicious part, ants discard the rest of the seed into their trash pile, where the future plant starts to grow. Although Ricinus communis is indigenous to the southeastern Mediterranean Basin , Eastern Africa , and India, today it is widespread throughout tropical regions. In areas with

3255-494: Is considered to be four to eight seeds, reports of actual poisoning are relatively rare. According to the Guinness World Records , this is the world's most poisonous common plant. If ricin is ingested, symptoms commonly begin within two to four hours, but may be delayed by up to 36 hours. These include a burning sensation in mouth and throat, abdominal pain, purging and bloody diarrhea. Within several days there

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3360-783: Is due to the development of larger root systems which can access water and soil nutrients deeper in the soil and to earlier emergence in the spring. Annual plants have an advantage in disturbed environments because of their faster growth and reproduction rates. Each section contains a short list of species related to that topic, these are an example as the true lists would fill several books. Perennials grown for their decorative flowers include very many species and types. Some examples include: The majority of fruit bearing plants are perennial even in temperate climates. Examples include: Many herbs are perennial, including these examples: Many vegetable plants can grow as perennials in tropical climates, but die in cold weather. Examples of some of

3465-483: Is fed by the re-sowing of the seeds of annual grain crops, (either naturally or by the manual efforts of humans), perennial crops provide numerous benefits. Perennial plants often have deep, extensive root systems which can hold soil to prevent erosion , capture dissolved nitrogen before it can contaminate ground and surface water, and out-compete weeds (reducing the need for herbicides ). These potential benefits of perennials have resulted in new attempts to increase

3570-615: Is known as the "father of modern taxonomy ". Many of his writings were in Latin; his name is rendered in Latin as Carolus Linnæus and, after his 1761 ennoblement , as Carolus a Linné . Linnaeus was the son of a curate and was born in Råshult , in the countryside of Småland , southern Sweden . He received most of his higher education at Uppsala University and began giving lectures in botany there in 1730. He lived abroad between 1735 and 1738, where he studied and also published

3675-533: Is most likely only a single gene controlling the production of the pigment in some varieties. The stems and the spherical, spiny seed capsules also vary in pigmentation. The fruit capsules of some varieties are more showy than the flowers. The flowers lack petals and are unisexual (male and female) where both types are borne on the same plant ( monoecious ) in terminal panicle -like inflorescences of green or, in some varieties, shades of red. The male flowers are numerous, yellowish-green with prominent creamy stamens ;

3780-717: Is not toxic to humans in normal doses, whether internal or external. Castor oil and the plant's roots and leaves are used in the ancient Indian medicinal system of Ayurveda . Global castor seed production is around two million tons per year. Leading producing areas are India (with over three-quarters of the global yield), China and Mozambique, and it is widely grown as a crop in Ethiopia. There are several active breeding programmes. Other modern uses of natural, blended, or chemically altered castor products include: Castor seeds have been found in Egyptian tombs dating back to 4000 BC;

3885-612: Is severe dehydration, a drop in blood pressure and a decrease in urine. Unless treated, death can be expected to occur within 3–5 days; however, in most cases a full recovery can be made. Poisoning occurs when animals, including humans, ingest broken castor beans or break the seed by chewing: intact seeds may pass through the digestive tract without releasing the toxin. The toxin provides the castor oil plant with some degree of natural protection from insect pests such as aphids . Ricin has been investigated for its potential use as an insecticide . Commercially available cold-pressed castor oil

3990-535: Is the source of castor oil , which has a wide variety of uses. The seeds contain between 40% and 60% oil that is rich in triglycerides , mainly ricinolein . The seed also contains ricin , a highly potent water-soluble toxin , which is also present in lower concentrations throughout the plant . The plant known as "false castor oil plant", Fatsia japonica , is not closely related. Ricinus communis can vary greatly in its growth habit and appearance. The variability has been increased by breeders who have selected

4095-429: Is too cold or dry. In many parts of the world, seasonality is expressed as wet and dry periods rather than warm and cold periods, and deciduous perennials lose their leaves in the dry season. Some perennial plants are protected from wildfires because they have underground roots that produce adventitious shoots, bulbs, crowns , or stems ; other perennials like trees and shrubs may have thick cork layers that protect

4200-450: The Beast of Revelation . Even at the risk of incurring the mayor's wrath, Linnaeus made his observations public, dashing the mayor's dreams of selling the hydra for an enormous sum. Linnaeus and Sohlberg were forced to flee from Hamburg. Linnaeus began working towards his degree as soon as he reached Harderwijk , a university known for awarding degrees in as little as a week. He submitted

4305-475: The astronomical symbol for the planet Jupiter . Perennial plants can be short-lived (only a few years) or long-lived. They include a wide assortment of plant groups from non-flowering plants like ferns and liverworts to highly diverse flowering plants like orchids , grasses , and woody plants . Plants that flower and fruit only once and then die are termed monocarpic or semelparous ; these species may live for many years before they flower. For example,

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4410-495: The curate of the small village of Stenbrohult in Småland. Christina was the daughter of the rector of Stenbrohult, Samuel Brodersonius. A year after Linnaeus's birth, his grandfather Samuel Brodersonius died, and his father Nils became the rector of Stenbrohult. The family moved into the rectory from the curate's house. Even in his early years, Linnaeus seemed to have a liking for plants, flowers in particular. Whenever he

4515-469: The larvae of muga and endi silkworms . Castor oil is an effective motor lubricant and has been used in internal combustion engines , including those of World War I airplanes, some racing cars and some model airplanes . It has historically been popular for lubricating two-stroke engines due to high resistance to heat compared to petroleum-based oils. It does not mix well with petroleum products, particularly at low temperatures, but mixes better with

4620-624: The Italian poet and Fascist supporter, during the First World War . Extract of Ricinus communis exhibited acaricidal and insecticidal activities against the adult of Haemaphysalis bispinosa ( Acarina : Ixodidae ) and hematophagous fly Hippobosca maculata ( Diptera : Hippoboscidae ). Members of the Bodo tribe of Bodoland in Assam , India, use the leaves of the plant to feed

4725-578: The Scottish doctor Isaac Lawson , the manuscript was published as Systema Naturae (1735). Linnaeus became acquainted with one of the most respected physicians and botanists in the Netherlands, Herman Boerhaave , who tried to convince Linnaeus to make a career there. Boerhaave offered him a journey to South Africa and America, but Linnaeus declined, stating he would not stand the heat. Instead, Boerhaave convinced Linnaeus that he should visit

4830-558: The Swedish province of Västergötland . He set out from Uppsala on 12 June and returned on 11 August. On the expedition his primary companion was Erik Gustaf Lidbeck, a student who had accompanied him on his previous journey. Linnaeus described his findings from the expedition in the book Wästgöta-Resa , published the next year. After he returned from the journey, the Government decided Linnaeus should take on another expedition to

4935-490: The University although the young man was only a second-year student. His lectures were popular, and Linnaeus often addressed an audience of 300 people. In June, Linnaeus moved from Celsius's house to Rudbeck's to become the tutor of the three youngest of his 24 children. His friendship with Celsius did not wane and they continued their botanical expeditions. Over that winter, Linnaeus began to doubt Tournefort's system of classification and decided to create one of his own. His plan

5040-534: The abbreviation "Linn." is found. Linnaeus's remains constitute the type specimen for the species Homo sapiens following the International Code of Zoological Nomenclature , since the sole specimen that he is known to have examined was himself. Linnaeus was born in the village of Råshult in Småland , Sweden, on 23 May 1707. He was the first child of Nicolaus (Nils) Ingemarsson (who later adopted

5145-446: The advantage of generating larger seedlings that can better compete with other plants. Perennials also produce seeds over many years. An important aspect of cold acclimation is overexpression of DNA repair genes. In Thinopyrum intermedium a perennial relative of common wheat Triticum aestivum , conditions of freezing stress were shown to be associated with large increases in expression of two DNA repair genes (one gene product

5250-543: The age of 17, Linnaeus had become well acquainted with the existing botanical literature. He remarks in his journal that he "read day and night, knowing like the back of my hand, Arvidh Månsson's Rydaholm Book of Herbs, Tillandz's Flora Åboensis, Palmberg's Serta Florea Suecana, Bromelii's Chloros Gothica and Rudbeckii's Hortus Upsaliensis". Linnaeus entered the Växjö Katedralskola in 1724, where he studied mainly Greek , Hebrew , theology and mathematics ,

5355-442: The botanist Johannes Burman . After his visit, Burman, impressed with his guest's knowledge, decided Linnaeus should stay with him during the winter. During his stay, Linnaeus helped Burman with his Thesaurus Zeylanicus . Burman also helped Linnaeus with the books on which he was working: Fundamenta Botanica and Bibliotheca Botanica . In August 1735, during Linnaeus's stay with Burman, he met George Clifford III ,

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5460-413: The castor semi-looper moth ( Achaea janata ). It is also used as a food plant by the larvae of some other species of Lepidoptera , including Hypercompe hambletoni and the nutmeg ( Discestra trifolii ). A jumping spider Evarcha culicivora has an association with R. communis . They consume the nectar for food and preferentially use these plants as a location for courtship. Each castor seed has

5565-424: The class of organisms. Linnaeus published Species Plantarum , the work which is now internationally accepted as the starting point of modern botanical nomenclature , in 1753. The first volume was issued on 24 May, the second volume followed on 16 August of the same year. The book contained 1,200 pages and was published in two volumes; it described over 7,300 species. The same year the king dubbed him knight of

5670-409: The classification of fish. One of the first scientists Linnaeus met in the Netherlands was Johan Frederik Gronovius , to whom Linnaeus showed one of the several manuscripts he had brought with him from Sweden. The manuscript described a new system for classifying plants. When Gronovius saw it, he was very impressed, and offered to help pay for the printing. With an additional monetary contribution by

5775-468: The dispersal of the seed by ants (myrmecochory). Three terpenoids and a tocopherol -related compound have been found in the aerial parts of Ricinus . Compounds named (3 E ,7 Z ,11 E )-19-hydroxycasba-3,7,11-trien-5-one, 6α-hydroxy-10β-methoxy-7α,8α-epoxy-5-oxocasbane-20,10-olide, 15α-hydroxylup-20(29)-en-3-one, and (2 R ,4a R ,8a R )-3,4,4a,8a-tetrahydro-4a-hydroxy-2,6,7,8a-tetramethyl-2-(4,8, 12-trimethyltridecyl)-2 H -chromene-5,8-dione were isolated from

5880-510: The dormancy period is over and new growth begins. In climates that are warm all year long, perennials may grow continuously. Annuals which complete their life cycle in one growing season, in contrast with perennials, produce seeds as the next generation and die; the seeds may survive cold or dry periods or germinate soon after dispersal depending on the climate. Some perennials retain their foliage year-round; these are evergreen perennials. Deciduous perennials shed all their leaves part of

5985-440: The expedition were later published in Öländska och Gothländska Resa , written in Swedish. Like Flora Lapponica , it contained both zoological and botanical observations, as well as observations concerning the culture in Öland and Gotland. During the summer of 1745, Linnaeus published two more books: Flora Suecica and Fauna Suecica . Flora Suecica was a strictly botanical book, while Fauna Suecica

6090-411: The family name Linnaeus) and Christina Brodersonia. His siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus (who would eventually succeed their father as rector of Stenbrohult and write a manual on beekeeping ), and Emerentia Linnæa. His father taught him Latin as a small child. One of a long line of peasants and priests, Nils was an amateur botanist , a Lutheran minister , and

6195-429: The female flowers, borne at the tips of the spikes, lie within the immature spiny capsules, are relatively few in number and have prominent red stigmas . The fruit is a spiny, greenish (to reddish-purple) capsule containing large, oval, shiny, bean-like, highly poisonous seeds with variable brownish mottling. Castor seeds have a warty appendage called the caruncle , which is a type of elaiosome . The caruncle promotes

6300-422: The first edition of his Systema Naturae in the Netherlands. He then returned to Sweden where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on several journeys through Sweden to find and classify plants and animals. In the 1750s and 1760s, he continued to collect and classify animals, plants, and minerals, while publishing several volumes. By the time of his death in 1778, he

6405-544: The flora and fauna in the vicinity of Uppsala. Linnaeus published Philosophia Botanica in 1751. The book contained a complete survey of the taxonomy system he had been using in his earlier works. It also contained information of how to keep a journal on travels and how to maintain a botanical garden. During Linnaeus's time it was normal for upper class women to have wet nurses for their babies. Linnaeus joined an ongoing campaign to end this practice in Sweden and promote breast-feeding by mothers. In 1752 Linnaeus published

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6510-541: The growing season, and can grow taller than annuals. In doing so they can better compete for space and collect more light. Perennials typically grow structures that allow them to adapt to living from one year to the next through a form of vegetative reproduction rather than seeding. These structures include bulbs , tubers , woody crowns, rhizomes , turions , woody stems, or crowns which allows them to survive periods of dormancy over cold or dry seasons; these structures typically store carbohydrates which are used once

6615-620: The hormones produced due to environmental situations (i.e., seasons), reproduction, and stage of development to begin and halt the ability to grow or flower. There is also a distinction between the ability to grow and the actual task of growth. For example, most trees regain the ability to grow during winter but do not initiate physical growth until the spring and summer months. The start of dormancy can be seen in perennial plants through withering flowers, loss of leaves on trees, and halting of reproduction in both flowering and budding plants. Perennial species may produce relatively large seeds that have

6720-889: The journey in 1695, but the detailed results of his exploration were lost in a fire seven years afterwards. Linnaeus's hope was to find new plants, animals and possibly valuable minerals. He was also curious about the customs of the native Sami people , reindeer-herding nomads who wandered Scandinavia's vast tundras. In April 1732, Linnaeus was awarded a grant from the Royal Society of Sciences in Uppsala for his journey. Linnaeus began his expedition from Uppsala on 12 May 1732, just before he turned 25. He travelled on foot and horse, bringing with him his journal, botanical and ornithological manuscripts and sheets of paper for pressing plants. Near Gävle he found great quantities of Campanula serpyllifolia , later known as Linnaea borealis ,

6825-586: The kindness of Dutch friends obliged him to stay some months longer in Holland. In May 1738, he set out for Sweden again. On the way home, he stayed in Paris for about a month, visiting botanists such as Antoine de Jussieu . After his return, Linnaeus never again left Sweden. When Linnaeus returned to Sweden on 28 June 1738, he went to Falun , where he entered into an engagement to Sara Elisabeth Moræa . Three months later, he moved to Stockholm to find employment as

6930-530: The last year of the Lower School when he was fifteen, which was taught by the headmaster, Daniel Lannerus, who was interested in botany. Lannerus noticed Linnaeus's interest in botany and gave him the run of his garden. He also introduced him to Johan Rothman, the state doctor of Småland and a teacher at Katedralskolan (a gymnasium ) in Växjö. Also a botanist, Rothman broadened Linnaeus's interest in botany and helped him develop an interest in medicine. By

7035-399: The lungs, triggering allergic reactions. The sap of the plant causes skin rashes. People who are allergic to the plant can also develop rashes from touching the leaves, flowers, or seeds. They can also have cross-allergic reactions to latex sap from the related Hevea brasiliensis plant. The toxicity of raw castor beans is due to the presence of ricin . Although the lethal dose in adults

7140-626: The mamillæ [breasts] and de- fluxions; a decoction of them in wine, with polenta and saffron , is good for inflammations of various kinds. Boiled by themselves, and applied to the face for three successive days, they improve the complexion." In Haiti it is called maskreti , where the plant is turned into a red oil that is then given to newborns as a purgative to cleanse the insides of their first stools. Castor seed and its oil have also been used in China for centuries, mainly prescribed in local medicine for internal use or use in dressings. Castor oil

7245-451: The methanol extracts of Ricinus communis by chromatographic methods. Partitioned h-hexane fraction of Ricinus root methanol extract resulted in enrichment of two triterpenes: lupeol and urs-6-ene-3,16-dione (erandone). Crude methanolic extract, enriched n-hexane fraction and isolates at doses 100 mg/kg p.o. exhibited significant (P < 0.001) anti-inflammatory activity in carrageenan-induced hind paw oedema model. Carl Linnaeus used

7350-412: The methanol-based fuels used in glow model engines . In total-loss-lubrication applications, it tends to leave carbon deposits and varnish within the engine. It has been largely replaced by synthetic oils that are more stable and less toxic. Jewellery can be made of castor beans, particularly necklaces and bracelets. Holes must not be drilled into the castor beans as the shell protects the wearer from

7455-407: The more completely perennial vegetables are: Many aquatic plants are perennial even though many do not have woody tissue. Examples include: Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné , was a Swedish biologist and physician who formalised binomial nomenclature , the modern system of naming organisms. He

7560-433: The name Ricinus because it is a Latin word for tick ; the seed is so named because it has markings and a bump at the end that resemble certain ticks. The genus Ricinus also exists in zoology, and designates insects (not ticks) which are parasites of birds; this is possible because the names of animals and plants are governed by different nomenclature codes . The common name "castor oil" probably comes from its use as

7665-459: The personality of their wet nurse through the milk. He admired the child care practices of the Lapps and pointed out how healthy their babies were compared to those of Europeans who employed wet nurses. He compared the behaviour of wild animals and pointed out how none of them denied their newborns their breastmilk. It is thought that his activism played a role in his choice of the term Mammalia for

7770-669: The plant grows wild in the US, notably Griffith Park in Los Angeles. Cultivars have been developed by breeders for use as ornamental plants (heights refer to plants grown as annuals) and for commercial production of castor oil. Ricinus is extremely allergenic, and has an OPALS allergy scale rating of 10 out of 10. The plant is also a very strong trigger for asthma, and allergies to Ricinus are commonplace and severe. The castor oil plant produces abundant amounts of very light pollen, which easily become airborne and can be inhaled into

7875-461: The quality of its ferruginous water. The journey was successful, and Linnaeus's observations were published the next year in Skånska Resa . In 1750, Linnaeus became rector of Uppsala University, starting a period where natural sciences were esteemed. Perhaps the most important contribution he made during his time at Uppsala was to teach; many of his students travelled to various places in

7980-459: The ricin. Any chips in the shell can cause poisoning of the wearer. Pets who chew the jewellery can become ill. Ricinus communis leaves are used in botanical printing (also known as ecoprinting) in Asia. When bundled with cotton or silk fabric and steamed, the leaves can produce a green-colored imprint. Perennial Perennials (especially small flowering plants ) that grow and bloom over

8085-448: The same time can be an out-crosser by anemophily (wind pollination) or entomophily (insect pollination). Its seed is the castor bean, which despite the term is not a bean (as it is not the seed of a member of the family Fabaceae ). Castor is indigenous to the southeastern Mediterranean Basin , East Africa , and India , but is widespread throughout tropical regions (and widely grown elsewhere as an ornamental plant). Castor seed

8190-465: The seed yield of perennial species, which could result in the creation of new perennial grain crops. Some examples of new perennial crops being developed are perennial rice and intermediate wheatgrass . A perennial rice developed in 2018, was reported in 2023, to have provided a similar yield to replanted annual rice when evaluated over eight consecutive harvests. Perennial plants dominate many natural ecosystems on land and in fresh water, with only

8295-407: The slow-burning oil was mostly used to fuel lamps. Herodotus and other Greek travellers noted the use of castor seed oil for lighting, body ointments, and improving hair growth and texture. Cleopatra is reputed to have used it to brighten the whites of her eyes. The Ebers Papyrus is an ancient Egyptian medical treatise believed to date from 1552 BC. Translated in 1872, it describes castor oil as

8400-570: The southernmost province Scania . This journey was postponed, as Linnaeus felt too busy. In 1747, Linnaeus was given the title archiater , or chief physician, by the Swedish king Adolf Frederick —a mark of great respect. The same year he was elected member of the Academy of Sciences in Berlin . In the spring of 1749, Linnaeus could finally journey to Scania , again commissioned by the government. With him he brought his student Olof Söderberg. On

8505-668: The spring and summer, die back every autumn and winter, and then return in the spring from their rootstock or other overwintering structure, are known as herbaceous perennials . However, depending on the rigours of the local climate (temperature, moisture, organic content in the soil, microorganisms), a plant that is a perennial in its native habitat, may be treated by a gardener as an annual and planted out every year, from seed, from cuttings, or from divisions. Tomato vines, for example, live several years in their natural tropical/ subtropical habitat but are grown as annuals in temperate regions because their above-ground biomass does not survive

8610-431: The stems. Herbaceous perennials from temperate and alpine regions of the world can tolerate the cold during winter. Perennial plants may remain dormant for long periods and then recommence growth and reproduction when the environment is more suitable, while most annual plants complete their life cycle during one growing period, and biennials have two growing periods. The meristem of perennial plants communicates with

8715-758: The twinflower that would become his favourite. He sometimes dismounted on the way to examine a flower or rock and was particularly interested in mosses and lichens , the latter a main part of the diet of the reindeer , a common and economically important animal in Lapland. Linnaeus travelled clockwise around the coast of the Gulf of Bothnia , making major inland incursions from Umeå , Luleå and Tornio . He returned from his six-month-long, over 2,000 kilometres (1,200 mi) expedition in October, having gathered and observed many plants, birds and rocks. Although Lapland

8820-636: The two men remained in correspondence for many years afterwards. Linnaeus dedicated his Critica Botanica to him, as " opus botanicum quo absolutius mundus non-vidit ". Linnaeus would later name a genus of tropical tree Dillenia in his honour. He then returned to Hartekamp, bringing with him many specimens of rare plants. The next year, 1737, he published Genera Plantarum , in which he described 935 genera of plants, and shortly thereafter he supplemented it with Corollarium Generum Plantarum , with another sixty ( sexaginta ) genera. His work at Hartekamp led to another book, Hortus Cliffortianus ,

8925-474: The two professors who taught at the medical faculty at Uppsala: Olof Rudbeck the Younger and Lars Roberg . Although Rudbeck and Roberg had undoubtedly been good professors, by then they were older and not so interested in teaching. Rudbeck no longer gave public lectures, and had others stand in for him. The botany, zoology, pharmacology and anatomy lectures were not in their best state. In Uppsala, Linnaeus met

9030-455: The use of his library, which included many books about botany. He also gave the student free admission to his lectures. In his spare time, Linnaeus explored the flora of Skåne, together with students sharing the same interests. In August 1728, Linnaeus decided to attend Uppsala University on the advice of Rothman, who believed it would be a better choice if Linnaeus wanted to study both medicine and botany. Rothman based this recommendation on

9135-550: The way to Scania, he made his last visit to his brothers and sisters in Stenbrohult since his father had died the previous year. The expedition was similar to the previous journeys in most aspects, but this time he was also ordered to find the best place to grow walnut and Swedish whitebeam trees; these trees were used by the military to make rifles. While there, they also visited the Ramlösa mineral spa , where he remarked on

9240-476: The way, the pair stopped in Hamburg , where they met the mayor, who proudly showed them a supposed wonder of nature in his possession: the taxidermied remains of a seven-headed hydra . Linnaeus quickly discovered the specimen was a fake , cobbled together from the jaws and paws of weasels and the skins of snakes. The provenance of the hydra suggested to Linnaeus that it had been manufactured by monks to represent

9345-405: The winter. There is also a class of evergreen perennials which lack woody stems, such as Bergenia which retain a mantle of leaves throughout the year. An intermediate class of plants is known as subshrubs , which retain a vestigial woody structure in winter, e.g. Penstemon . The symbol for a perennial plant, based on Species Plantarum by Linnaeus , is [REDACTED] , which is also

9450-485: The world to collect botanical samples. Linnaeus called the best of these students his "apostles". His lectures were normally very popular and were often held in the Botanical Garden. He tried to teach the students to think for themselves and not trust anybody, not even him. Even more popular than the lectures were the botanical excursions made every Saturday during summer, where Linnaeus and his students explored

9555-439: The year. Deciduous perennials include herbaceous and woody plants; herbaceous plants have stems that lack hard, fibrous growth, while woody plants have stems with buds that survive above ground during dormancy. Some perennials are semi-deciduous, meaning they lose some of their leaves in either winter or summer. Deciduous perennials shed their leaves when growing conditions are no longer suitable for photosynthesis, such as when it

9660-537: The æ ligature, both in handwritten documents and in publications. Carl's patronymic would have been Nilsson, as in Carl Nilsson Linnæus. Linnaeus's father began teaching him basic Latin, religion, and geography at an early age. When Linnaeus was seven, Nils decided to hire a tutor for him. The parents picked Johan Telander, a son of a local yeoman . Linnaeus did not like him, writing in his autobiography that Telander "was better calculated to extinguish

9765-433: Was zoological . Anders Celsius had created the temperature scale named after him in 1742. Celsius's scale was originally inverted compared to the way it is used today, with water boiling at 0 °C and freezing at 100 °C. Linnaeus was the one who inverted the scale to its present usage, in 1745. In the summer of 1746, Linnaeus was once again commissioned by the Government to carry out an expedition, this time to

9870-413: Was a region with limited biodiversity , Linnaeus described about 100 previously unidentified plants. These became the basis of his book Flora Lapponica . However, on the expedition to Lapland, Linnaeus used Latin names to describe organisms because he had not yet developed the binomial system. In Flora Lapponica Linnaeus's ideas about nomenclature and classification were first used in

9975-640: Was a serious subject. He taught Linnaeus to classify plants according to Tournefort's system. Linnaeus was also taught about the sexual reproduction of plants, according to Sébastien Vaillant . In 1727, Linnaeus, age 21, enrolled in Lund University in Skåne . He was registered as Carolus Linnæus , the Latin form of his full name, which he also used later for his Latin publications. Professor Kilian Stobæus , natural scientist, physician and historian, offered Linnaeus tutoring and lodging, as well as

10080-441: Was appointed Professor of Medicine at Uppsala University, first with responsibility for medicine-related matters. Soon, he changed place with the other Professor of Medicine, Nils Rosén, and thus was responsible for the Botanical Garden (which he would thoroughly reconstruct and expand), botany and natural history , instead. In October that same year, his wife and nine-month-old son followed him to live in Uppsala. Ten days after he

10185-487: Was appointed professor, he undertook an expedition to the island provinces of Öland and Gotland with six students from the university to look for plants useful in medicine. They stayed on Öland until 21 June, then sailed to Visby in Gotland. Linnaeus and the students stayed on Gotland for about a month, and then returned to Uppsala. During this expedition, they found 100 previously unrecorded plants. The observations from

10290-448: Was during this expedition that Linnaeus had a flash of insight regarding the classification of mammals. Upon observing the lower jawbone of a horse at the side of a road he was travelling, Linnaeus remarked: "If I only knew how many teeth and of what kind every animal had, how many teats and where they were placed, I should perhaps be able to work out a perfectly natural system for the arrangement of all quadrupeds." In 1734, Linnaeus led

10395-625: Was here that he wrote a book Hortus Cliffortianus , in the preface of which he described his experience as "the happiest time of my life". (A portion of Hartekamp was declared as public garden in April 1956 by the Heemstede local authority, and was named "Linnaeushof". It eventually became, as it is claimed, the biggest playground in Europe. ) In July 1736, Linnaeus travelled to England, at Clifford's expense. He went to London to visit Sir Hans Sloane,

10500-558: Was named Ingemarsson after his father Ingemar Bengtsson. When Nils was admitted to the University of Lund , he had to take on a family name. He adopted the Latinate name Linnæus after a giant linden tree (or lime tree), lind in Swedish, that grew on the family homestead. This name was spelled with the æ ligature . When Carl was born, he was named Carl Linnæus, with his father's family name. The son also always spelled it with

10605-420: Was one of the most acclaimed scientists in Europe. Philosopher Jean-Jacques Rousseau sent him the message: "Tell him I know no greater man on Earth." Johann Wolfgang von Goethe wrote: "With the exception of Shakespeare and Spinoza , I know no one among the no longer living who has influenced me more strongly." Swedish author August Strindberg wrote: "Linnaeus was in reality a poet who happened to become

10710-551: Was permitted to visit the mines. In April 1735, at the suggestion of Sohlberg's father, Linnaeus and Sohlberg set out for the Dutch Republic , where Linnaeus intended to study medicine at the University of Harderwijk while tutoring Sohlberg in exchange for an annual salary. At the time, it was common for Swedes to pursue doctoral degrees in the Netherlands , then a highly revered place to study natural history. On

10815-752: Was to divide the plants by the number of stamens and pistils . He began writing several books, which would later result in, for example, Genera Plantarum and Critica Botanica . He also produced a book on the plants grown in the Uppsala Botanical Garden , Adonis Uplandicus . Rudbeck's former assistant, Nils Rosén , returned to the University in March 1731 with a degree in medicine. Rosén started giving anatomy lectures and tried to take over Linnaeus's botany lectures, but Rudbeck prevented that. Until December, Rosén gave Linnaeus private tutoring in medicine. In December, Linnaeus had

10920-414: Was upset, he was given a flower, which immediately calmed him. Nils spent much time in his garden and often showed flowers to Linnaeus and told him their names. Soon Linnaeus was given his own patch of earth where he could grow plants. Carl's father was the first in his ancestry to adopt a permanent surname. Before that, ancestors had used the patronymic naming system of Scandinavian countries: his father

11025-556: Was used as an instrument of coercion by the paramilitary Blackshirts under the regime of Italian dictator Benito Mussolini and by the Spanish Civil Guard in Francoist Spain. Dissidents and regime opponents were forced to ingest the oil in large amounts, triggering severe diarrhea and dehydration , which could ultimately cause death. This punishment method was originally thought of by Gabriele D'Annunzio ,

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