91-772: Windsor Hill is a 61.8-hectare (153-acre) biological Site of Special Scientific Interest in Princes Risborough in Buckinghamshire . It lies within the Chilterns Area of Outstanding Natural Beauty , and it is featured in the Nature Conservation Review . A small part is managed by the Berkshire, Buckinghamshire and Oxfordshire Wildlife Trust , and access to this area requires a permit. The wood consists of an extensive tract of
182-505: A chrysalis . When metamorphosis is complete, the pupal skin splits, the adult insect climbs out, expands its wings to dry, and flies off. Some butterflies, especially in the tropics, have several generations in a year, while others have a single generation, and a few in cold locations may take several years to pass through their entire life cycle. Butterflies are often polymorphic , and many species make use of camouflage , mimicry , and aposematism to evade their predators. Some, like
273-637: A diapause (resting) stage, and the hatching may take place only in spring. Some temperate region butterflies, such as the Camberwell beauty , lay their eggs in the spring and have them hatch in the summer. Butterfly larvae, or caterpillars, consume plant leaves and spend practically all of their time searching for and eating food. Although most caterpillars are herbivorous, a few species are predators : Spalgis epius eats scale insects , while lycaenids such as Liphyra brassolis are myrmecophilous , eating ant larvae. Some larvae, especially those of
364-493: A butterfly cannot fly until the wings are unfolded. A newly emerged butterfly needs to spend some time inflating its wings with hemolymph and letting them dry, during which time it is extremely vulnerable to predators. The colourful patterns on many butterfly wings tell potential predators that they are toxic. Hence, the genetic basis of wing pattern formation can illuminate both the evolution of butterflies as well as their developmental biology . The colour of butterfly wings
455-570: A butterfly through metamorphosis has held great appeal to mankind. To transform from the miniature wings visible on the outside of the pupa into large structures usable for flight, the pupal wings undergo rapid mitosis and absorb a great deal of nutrients. If one wing is surgically removed early on, the other three will grow to a larger size. In the pupa, the wing forms a structure that becomes compressed from top to bottom and pleated from proximal to distal ends as it grows, so that it can rapidly be unfolded to its full adult size. Several boundaries seen in
546-417: A common family. In some species, such as the great spangled fritillary , the eggs are deposited close to but not on the food plant. This most likely happens when the egg overwinters before hatching and where the host plant loses its leaves in winter, as do violets in this example. The egg stage lasts a few weeks in most butterflies, but eggs laid close to winter, especially in temperate regions, go through
637-591: A continuation of the historical management of the land. Where an owner or occupier is unwilling or unable to carry out management, ultimately the conservation body can require it to be done. Public bodies which own or occupy an SSSI have a duty to manage it properly. Site management statements for SSSI in Scotland are available to download from the NatureScot website using the "Sitelink" facility. The law protecting SSSIs now covers everyone, not just public bodies and
728-576: A few butterflies (e.g., harvesters ) eat harmful insects, and a few are predators of ants , while others live as mutualists in association with ants. Culturally, butterflies are a popular motif in the visual and literary arts. The Smithsonian Institution says "butterflies are certainly one of the most appealing creatures in nature". The Oxford English Dictionary derives the word straightforwardly from Old English butorflēoge , butter-fly; similar names in Old Dutch and Old High German show that
819-444: A few species. Some butterflies have organs of hearing and some species make stridulatory and clicking sounds. Many species of butterfly maintain territories and actively chase other species or individuals that may stray into them. Some species will bask or perch on chosen perches. The flight styles of butterflies are often characteristic and some species have courtship flight displays. Butterflies can only fly when their temperature
910-606: A hard-ridged outer layer of shell, called the chorion . This is lined with a thin coating of wax which prevents the egg from drying out before the larva has had time to fully develop. Each egg contains a number of tiny funnel-shaped openings at one end, called micropyles ; the purpose of these holes is to allow sperm to enter and fertilize the egg. Butterfly eggs vary greatly in size and shape between species, but are usually upright and finely sculptured. Some species lay eggs singly, others in batches. Many females produce between one hundred and two hundred eggs. Butterfly eggs are fixed to
1001-508: A leaf before eggs are laid on it. Many butterflies use chemical signals, pheromones ; some have specialized scent scales ( androconia ) or other structures ( coremata or "hair pencils" in the Danaidae). Vision is well developed in butterflies and most species are sensitive to the ultraviolet spectrum. Many species show sexual dimorphism in the patterns of UV reflective patches. Colour vision may be widespread but has been demonstrated in only
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#17327833260161092-416: A leaf with a special glue which hardens rapidly. As it hardens it contracts, deforming the shape of the egg. This glue is easily seen surrounding the base of every egg forming a meniscus. The nature of the glue has been little researched but in the case of Pieris brassicae , it begins as a pale yellow granular secretion containing acidophilic proteins. This is viscous and darkens when exposed to air, becoming
1183-478: A long way upstream of a wetland SSSI might require consultation. Some developments might be neutral or beneficial, even if they are within the SSSI itself – the critical point is whether they harm the interest features. The owners and occupiers of SSSIs are required (Scotland, England, Wales) to obtain consent from the relevant nature conservation body if they want to carry out, cause or permit to be carried out within
1274-547: A longer lifespan of several months as adults. The thorax of the butterfly is devoted to locomotion. Each of the three thoracic segments has two legs (among nymphalids , the first pair is reduced and the insects walk on four legs). The second and third segments of the thorax bear the wings. The leading edges of the forewings have thick veins to strengthen them, and the hindwings are smaller and more rounded and have fewer stiffening veins. The forewings and hindwings are not hooked together ( as they are in moths ) but are coordinated by
1365-452: A necessity for their successful establishment. Many butterflies, such as the painted lady , monarch, and several danaine migrate for long distances. These migrations take place over a number of generations and no single individual completes the whole trip. The eastern North American population of monarchs can travel thousands of miles south-west to overwintering sites in Mexico . There is
1456-407: A proposed activity would not affect the interest or is beneficial to it, then the conservation body will issue a "consent" allowing it to be carried out without further consultation. If it would be harmful, the conservation body may issue consent subject to conditions or refuse the application. If consent in writing is not given the operation must not proceed. Conditions may cover any relevant aspect of
1547-658: A reverse migration in the spring. It has recently been shown that the British painted lady undertakes a 9,000-mile round trip in a series of steps by up to six successive generations, from tropical Africa to the Arctic Circle — almost double the length of the famous migrations undertaken by monarch. Spectacular large-scale migrations associated with the monsoon are seen in peninsular India. Migrations have been studied in more recent times using wing tags and also using stable hydrogen isotopes . Butterflies navigate using
1638-412: A segmented palp. Adjoining these is the labium-hypopharynx which houses a tubular spinneret which is able to extrude silk. Caterpillars such as those in the genus Calpodes (family Hesperiidae) have a specialized tracheal system on the 8th segment that function as a primitive lung. Butterfly caterpillars have three pairs of true legs on the thoracic segments and up to six pairs of prolegs arising from
1729-435: A silken girdle may be spun to keep the pupa in a head-up position. Most of the tissues and cells of the larva are broken down inside the pupa, as the constituent material is rebuilt into the imago. The structure of the transforming insect is visible from the exterior, with the wings folded flat on the ventral surface and the two halves of the proboscis, with the antennae and the legs between them. The pupal transformation into
1820-428: A site may contain strata containing vertebrate fossils, insect fossils and plant fossils and it may also be of importance for stratigraphy . Geological sites fall into two types, having different conservation priorities: exposure sites, and deposit sites. Exposure sites are where quarries , disused railway cuttings, cliffs or outcrops give access to extensive geological features, such as particular rock layers. If
1911-425: A standard list for that country. The ORCs/OLDs are not "banned" activities – the list includes activities which would damage the interest, but also many which might be beneficial. For example, " grazing " (a standard item on the list) would require consent, even on a chalk grassland or heathland where grazing is an essential part of management. In England and Wales the list of OLDs is almost the same for each SSSI – and
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#17327833260162002-410: A time-compensated sun compass. They can see polarized light and therefore orient even in cloudy conditions. The polarized light near the ultraviolet spectrum appears to be particularly important. Many migratory butterflies live in semi-arid areas where breeding seasons are short. The life histories of their host plants also influence butterfly behaviour. Butterflies in their adult stage can live from
2093-541: A total of about 20,000 species. Traditionally, butterflies have been divided into the superfamilies Papilionoidea and the moth-like Hedyloidea . Recent work has discovered that Hedylidae, the only family within Hedyloidea, is nested within the Papilionoidea, meaning that Papilionoidea would be synonymous with Rhopalocera. The relationships between the rest of the 6 families are extremely well resolved, which
2184-470: A trend towards multivoltinism . Courtship is often aerial and often involves pheromones . Butterflies then land on the ground or on a perch to mate. Copulation takes place tail-to-tail and may last from minutes to hours. Simple photoreceptor cells located at the genitals are important for this and other adult behaviours. The male passes a spermatophore to the female; to reduce sperm competition, he may cover her with his scent, or in some species such as
2275-486: A tubular proboscis which is curled up at rest and expanded when needed to feed. The first and second maxillae bear palps which function as sensory organs. Some species have a reduced proboscis or maxillary palps and do not feed as adults. Many Heliconius butterflies also use their proboscis to feed on pollen; in these species only 20% of the amino acids used in reproduction come from larval feeding, which allow them to develop more quickly as caterpillars, and gives them
2366-437: A water-insoluble, rubbery material which soon sets solid. Butterflies in the genus Agathymus do not fix their eggs to a leaf; instead, the newly laid eggs fall to the base of the plant. Eggs are almost invariably laid on plants. Each species of butterfly has its own host plant range and while some species of butterfly are restricted to just one species of plant, others use a range of plant species, often including members of
2457-422: A week to nearly a year depending on the species. Many species have long larval life stages while others can remain dormant in their pupal or egg stages and thereby survive winters. The Melissa Arctic ( Oeneis melissa ) overwinters twice as a caterpillar. Butterflies may have one or more broods per year. The number of generations per year varies from temperate to tropical regions with tropical regions showing
2548-450: Is above 27 °C (81 °F); when it is cool, they can position themselves to expose the underside of the wings to the sunlight to heat themselves up. If their body temperature reaches 40 °C (104 °F), they can orientate themselves with the folded wings edgewise to the sun. Basking is an activity which is more common in the cooler hours of the morning. Some species have evolved dark wingbases to help in gathering more heat and this
2639-795: Is associated with various other typical chalk shrubs and chalk grassland with an abundance of the moss Pseudoscleropodium purum and common valerian ( Valeriana officinalis ). There are patches of bare chalk with forget-me-nots ( Myosotis species) and common mullein ( Verbascum thapsus ). On Kop Hill, hawthorn dominates the scrub, and the grassland is less mossy. Several localised plant species such as squinancywort ( Asperula cynanchica ) chalk eyebright ( Euphrasia pseudokerneri ), horseshoe vetch ( Hippocrepis comosa ), autumn gentian ( Gentianella amarella ) and clustered bellflower ( Campanula glomerata ) occur. A further small area of chalk grassland dominated by wood false-brome ( Brachypodium sylvaticum ) and glaucous sedge ( Carex flacca ) occurs at
2730-458: Is derived from tiny structures called scales, each of which have their own pigments . In Heliconius butterflies, there are three types of scales: yellow/white, black, and red/orange/brown scales. Some mechanism of wing pattern formation are now being solved using genetic techniques. For instance, a gene called cortex determines the colour of scales: deleting cortex turned black and red scales yellow. Mutations, e.g. transposon insertions of
2821-403: Is especially evident in alpine forms. As in many other insects, the lift generated by butterflies is more than can be accounted for by steady-state, non-transitory aerodynamics . Studies using Vanessa atalanta in a wind tunnel show that they use a wide variety of aerodynamic mechanisms to generate force. These include wake capture , vortices at the wing edge, rotational mechanisms and
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2912-410: Is extruded and inserted into the female's vagina. A spermatophore is deposited in the female, following which the sperm make their way to a seminal receptacle where they are stored for later use. In both sexes, the genitalia are adorned with various spines, teeth, scales and bristles, which act to prevent the butterfly from mating with an insect of another species. After it emerges from its pupal stage,
3003-451: Is good, especially in some species in the blue/violet range. The antennae are composed of many segments and have clubbed tips (unlike moths that have tapering or feathery antennae). The sensory receptors are concentrated in the tips and can detect odours. Taste receptors are located on the palps and on the feet. The mouthparts are adapted to sucking and the mandibles are usually reduced in size or absent. The first maxillae are elongated into
3094-516: Is governed by published SSSI Selection Guidelines. Within each area, a representative series of the best examples of each significant natural habitat may be notified, and for rarer habitats all examples may be included. Sites of particular significance for various taxonomic groups may be selected (for example birds, dragonflies , butterflies , reptiles, amphibians , etc.)—each of these groups has its own set of selection guidelines. Conservation of biological SSSI/ASSIs usually involves continuation of
3185-581: Is native to the Americas, but in the nineteenth century or before, spread across the world, and is now found in Australia, New Zealand, other parts of Oceania, and the Iberian Peninsula . It is not clear how it dispersed; adults may have been blown by the wind or larvae or pupae may have been accidentally transported by humans, but the presence of suitable host plants in their new environment was
3276-420: Is not necessarily absolute—generally it requires the SSSI interest to be considered properly against other factors. Local planning authorities are required to have policies in their development plans which protect SSSIs. They are then required to consult the appropriate conservation body over planning applications which might affect the interest of an SSSI (such a development might not be within or even close to
3367-441: Is often pronounced "triple-S I". Sites notified for their biological interest are known as Biological SSSIs (or ASSIs), and those notified for geological or physiographic interest are Geological SSSIs (or ASSIs). Sites may be divided into management units, with some areas including units that are noted for both biological and geological interest. Biological SSSI/ASSIs may be selected for various reasons, which for Great Britain
3458-465: Is restricted to the males, and studies have suggested that the nutrients collected may be provided as a nuptial gift , along with the spermatophore, during mating. In hilltopping , males of some species seek hilltops and ridge tops, which they patrol in search for females. Since it usually occurs in species with low population density, it is assumed these landscape points are used as meeting places to find mates. Butterflies use their antennae to sense
3549-698: Is summarized in the below cladogram. Papilionidae [REDACTED] Hedylidae [REDACTED] Hesperiidae [REDACTED] Pieridae [REDACTED] Nymphalidae [REDACTED] Lycaenidae [REDACTED] Riodinidae [REDACTED] Butterfly adults are characterized by their four scale-covered wings, which give the Lepidoptera their name ( Ancient Greek λεπίς lepís, scale + πτερόν pterón, wing). These scales give butterfly wings their colour: they are pigmented with melanins that give them blacks and browns, as well as uric acid derivatives and flavones that give them yellows, but many of
3640-405: Is taken up by the gut, but there may also be large silk glands, and special glands which secrete distasteful or toxic substances. The developing wings are present in later stage instars and the gonads start development in the egg stage. When the larva is fully grown, hormones such as prothoracicotropic hormone (PTTH) are produced. At this point the larva stops feeding, and begins "wandering" in
3731-658: The Isle of Man and Northern Ireland , is a conservation designation denoting a protected area in the United Kingdom and Isle of Man . SSSI/ASSIs are the basic building block of site-based nature conservation legislation and most other legal nature/geological conservation designations in the United Kingdom are based upon them, including national nature reserves , Ramsar sites , Special Protection Areas , and Special Areas of Conservation . The acronym "SSSI"
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3822-479: The Lycaenidae , form mutual associations with ants. They communicate with the ants using vibrations that are transmitted through the substrate as well as using chemical signals. The ants provide some degree of protection to these larvae and they in turn gather honeydew secretions . Large blue ( Phengaris arion ) caterpillars trick Myrmica ants into taking them back to the ant colony where they feed on
3913-453: The Marine and Coastal Access Act 2010 ). Access to SSSIs is the same as for the rest of the countryside of the relevant country. Most SSSIs/ASSIs are in private ownership and form parts of working farms, forests and estates. In Scotland, people may use their rights of responsible access to visit SSSIs. When designating an SSSI/ASSI, the relevant nature conservation body must formally notify
4004-983: The National Parks and Access to the Countryside Act 1949 , but the current legal framework for SSSIs is provided in England and Wales by the Wildlife and Countryside Act 1981 , amended in 1985 and further substantially amended in 2000 (by the Countryside and Rights of Way Act 2000 ), in Scotland by the Nature Conservation (Scotland) Act 2004 and in Northern Ireland by the Nature Conservation and Amenity Lands (Northern Ireland) Order 1985 . SSSIs are also covered under
4095-528: The Paleocene , about 56 million years ago, though molecular likely originated in the Cretaceous . Butterflies have a four-stage life cycle , and like other holometabolous insects they undergo complete metamorphosis . Winged adults lay eggs on the food plant on which their larvae , known as caterpillars , will feed. The caterpillars grow, sometimes very rapidly, and when fully developed, pupate in
4186-570: The Water Resources Act 1991 and related legislation. An SSSI may be made on any area of land which is considered to be of special interest by virtue of its fauna , flora , geological or physiographical / geomorphological features. SSSI notification can cover any "land" within the area of the relevant nature conservation body, including dry land, land covered by fresh water . The extent to which an SSSI/ASSI may extend seawards differs between countries. In Scotland an SSSI may include
4277-474: The Weis-Fogh ' clap-and-fling ' mechanism. Butterflies are able to change from one mode to another rapidly. Butterflies are threatened in their early stages by parasitoids and in all stages by predators, diseases and environmental factors. Braconid and other parasitic wasps lay their eggs in lepidopteran eggs or larvae and the wasps' parasitoid larvae devour their hosts, usually pupating inside or outside
4368-455: The ZW sex-determination system where females are the heterogametic sex (ZW) and males homogametic (ZZ). Butterflies are distributed worldwide except Antarctica, totalling some 18,500 species. Of these, 775 are Nearctic ; 7,700 Neotropical ; 1,575 Palearctic ; 3,650 Afrotropical ; and 4,800 are distributed across the combined Oriental and Australian / Oceania regions. The monarch butterfly
4459-402: The monarch and the painted lady , migrate over long distances. Many butterflies are attacked by parasites or parasitoids , including wasps , protozoans , flies , and other invertebrates, or are preyed upon by other organisms. Some species are pests because in their larval stages they can damage domestic crops or trees; other species are agents of pollination of some plants. Larvae of
4550-600: The non-coding DNA around the cortex gene can turn a black-winged butterfly into a butterfly with a yellow wing band. When the butterfly Bicyclus anynana is subjected to repeated inbreeding in the laboratory, there is a dramatic decrease in egg hatching. This severe inbreeding depression is considered to be likely due to a relatively high mutation rate to recessive alleles with substantial damaging effects and infrequent episodes of inbreeding in nature that might otherwise purge such mutations. Although B. anynana experiences inbreeding depression when forcibly inbred in
4641-540: The Apollos ( Parnassius ) plugs her genital opening to prevent her from mating again. The vast majority of butterflies have a four-stage life cycle: egg , larva (caterpillar), pupa (chrysalis) and imago (adult). In the genera Colias , Erebia , Euchloe , and Parnassius , a small number of species are known that reproduce semi-parthenogenetically ; when the female dies, a partially developed larva emerges from her abdomen. Butterfly eggs are protected by
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#17327833260164732-536: The Chiltern escarpment. It contains beech woodlands, scrub and chalk grassland . It is one of three extant British locations for the red helleborine orchid. The beech woodlands incorporate communities characteristic of soils on the Clay-with-flints of the Chiltern plateau and the chalky deposits of the scarp slopes. Associated with the beech on the plateau are oak and occasional birch whilst on
4823-563: The ORC lists for each SSSI and removed those activities that were unlikely to happen and if they were to would be unlikely to damage the protected natural features, and other activities adequately regulated by other statutory regimes. The intention of this was to remove the need for owners and occupiers to obtain SSSI consent as well as licences/ permits from other authorities (who must consult NatureScot prior to determining such applications). Purely geological SSSIs often have much shorter OLD lists. If
4914-607: The SSSI Register, hosted by The Registers of Scotland . Further information about SSSIs in Scotland is available on the NatureScot website. The decision to notify an SSSI is made by the relevant nature conservation body (the appropriate conservation body ) for that part of the United Kingdom: Northern Ireland Environment Agency , Natural England , NatureScot or Natural Resources Wales . SSSIs were originally set up by
5005-414: The SSSI any of the activities listed in the notification. Formerly these activities were called 'potentially damaging operations' or PDOs. Under the current legal arrangements they are called 'operations requiring consent' or ORCs (Scotland), or 'operations likely to damage the SSSI interest' or OLDs (England & Wales). The list of ORCs/OLDs for each SSSI is unique to that site – though all are derived from
5096-402: The SSSI itself). The effect of this is to prevent development which harms the interest – except where the value of that interest is over-ridden by some more important factor, for example a requirement for a major road or port or oil pipe. The requirement for consultation covers any development which might affect the interest, not just developments within the SSSI itself – for example, a development
5187-430: The abdomen, generally with short prolegs on segments 3–6 and 10; the three pairs of true legs on the thorax have five segments each. Many are well camouflaged; others are aposematic with bright colours and bristly projections containing toxic chemicals obtained from their food plants. The pupa or chrysalis, unlike that of moths, is not wrapped in a cocoon. Many butterflies are sexually dimorphic . Most butterflies have
5278-405: The abdominal segments. These prolegs have rings of tiny hooks called crochets that are engaged hydrostatically and help the caterpillar grip the substrate. The epidermis bears tufts of setae , the position and number of which help in identifying the species. There is also decoration in the form of hairs, wart-like protuberances, horn-like protuberances and spines. Internally, most of the body cavity
5369-502: The adult colour pattern are marked by changes in the expression of particular transcription factors in the early pupa. The reproductive stage of the insect is the winged adult or imago . The surface of both butterflies and moths is covered by scales, each of which is an outgrowth from a single epidermal cell. The head is small and dominated by the two large compound eyes . These are capable of distinguishing flower shapes or motion but cannot view distant objects clearly. Colour perception
5460-467: The air for wind and scents. The antennae come in various shapes and colours; the hesperiids have a pointed angle or hook to the antennae, while most other families show knobbed antennae. The antennae are richly covered with sensory organs known as sensillae . A butterfly's sense of taste is coordinated by chemoreceptors on the tarsi , or feet, which work only on contact, and are used to determine whether an egg-laying insect's offspring will be able to feed on
5551-434: The ant eggs and larvae in a parasitic relationship. Caterpillars mature through a series of developmental stages known as instars . Near the end of each stage, the larva undergoes a process called apolysis , mediated by the release of a series of neurohormones . During this phase, the cuticle , a tough outer layer made of a mixture of chitin and specialized proteins , is released from the softer epidermis beneath, and
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#17327833260165642-658: The blues, greens, reds and iridescent colours are created by structural coloration produced by the micro-structures of the scales and hairs. As in all insects, the body is divided into three sections: the head, thorax , and abdomen . The thorax is composed of three segments, each with a pair of legs. In most families of butterfly the antennae are clubbed, unlike those of moths which may be threadlike or feathery. The long proboscis can be coiled when not in use for sipping nectar from flowers. Nearly all butterflies are diurnal , have relatively bright colours, and hold their wings vertically above their bodies when at rest, unlike
5733-441: The desiccated husk. Most wasps are very specific about their host species and some have been used as biological controls of pest butterflies like the large white butterfly . When the small cabbage white was accidentally introduced to New Zealand, it had no natural enemies. In order to control it, some pupae that had been parasitised by a chalcid wasp were imported, and natural control was thus regained. Some flies lay their eggs on
5824-867: The designating authority is NatureScot ; the role in Wales is performed by Natural Resources Wales (formerly the Countryside Council for Wales ). In the Isle of Man the role is performed by the Department of Environment, Food and Agriculture . Geological SSSI/ASSIs are selected by a different mechanism to biological ones, with a minimalistic system selecting one site for each geological feature in Great Britain. Academic geological specialists have reviewed geological literature, selecting sites within Great Britain of at least national importance for each of
5915-419: The epidermis begins to form a new cuticle. At the end of each instar, the larva moults , the old cuticle splits and the new cuticle expands, rapidly hardening and developing pigment. Development of butterfly wing patterns begins by the last larval instar. Caterpillars have short antennae and several simple eyes . The mouthparts are adapted for chewing with powerful mandibles and a pair of maxillae, each with
6006-461: The exposure becomes obscured, the feature could in principle be re-exposed elsewhere. Conservation of these sites usually concentrates on maintenance of access for future study. Deposit sites are features which are limited in extent or physically delicate—for example, they include small lenses of sediment , mine tailings , caves and other landforms . If such features become damaged they cannot be recreated, and conservation usually involves protecting
6097-741: The family Hesperiidae (skippers). Molecular clock estimates suggest that butterflies originated sometime in the Late Cretaceous , but only significantly diversified during the Cenozoic, with one study suggesting a North American origin for the group. The oldest American butterfly is the Late Eocene Prodryas persephone from the Florissant Fossil Beds , approximately 34 million years old. Butterflies are divided into seven families that contain
6188-507: The feature from erosion or other damage. Following devolution, legal arrangements for SSSIs (Scotland, England, Wales) and ASSIs (Northern Ireland) differ between the countries of the UK. The Isle of Man ASSI system is a separate entity. NatureScot publishes a summary of the SSSI arrangements for SSSI owners and occupiers (other than public bodies) which can be downloaded from its website. Legal documents for all SSSIs in Scotland are available on
6279-758: The foot of Pink Hill. 23 species of butterflies have been recorded, including brown hairstreak ( Thecla betulae ), which requires scrub thickets and woodland edge habitats. The juniper colony supports several species of insects specific to this host plant. The snail fauna includes Abida secale , Helicella itala and Pomatias elegans . There is access from Kop Hill Road and Peters Lane. 51°37′01″N 0°49′14″W / 51.616849°N 0.820652°W / 51.616849; -0.820652 Site of Special Scientific Interest A site of special scientific interest ( SSSI ) in Great Britain , or an area of special scientific interest ( ASSI ) in
6370-409: The friction of their overlapping parts. The front two segments have a pair of spiracles which are used in respiration. The abdomen consists of ten segments and contains the gut and genital organs. The front eight segments have spiracles and the terminal segment is modified for reproduction. The male has a pair of clasping organs attached to a ring structure, and during copulation, a tubular structure
6461-555: The grass was growing. The earliest Lepidoptera fossils date to the Triassic - Jurassic boundary, around 200 million years ago. Butterflies evolved from moths, so while the butterflies are monophyletic (forming a single clade ), the moths are not. The oldest known butterfly is Protocoeliades kristenseni from the Palaeocene aged Fur Formation of Denmark, approximately 55 million years old, which belongs to
6552-405: The interest), but not illegal trail biking. This loophole was closed by the Countryside and Rights of Way Act 2000 and section 19 of the Nature Conservation (Scotland) Act 2004. Funding for the monitoring of SSSIs in England has been cut from £1.58 million in 2010 to £700,000 in 2018, causing concern that many have not been inspected over the last six years, as required by guidelines. Since
6643-473: The interested parties and allow a period for them to make representations before confirming the notification. When creating a new SSSI/ASSI the designation has legal effect from the date of notification. The interested parties include central government, local planning authorities , national park authorities, all the owners and occupiers of the land, relevant public bodies such as the utility providers e.g., water companies . In Scotland, NatureScot must also notify
6734-469: The intertidal land down to mean low water spring or to the extent of the local planning authority area, thus only limited areas of estuaries and coastal waters beyond MLWS may be included. In England, Natural England may notify an SSSI over estuarial waters and further adjacent waters in certain circumstances (section 28(1A & 1B) of the Wildlife and Countryside Act 1981 as amended by Part 2 of Annex 13 of
6825-1030: The laboratory it recovers within a few generation when allowed to breed freely. During mate selection, adult females do not innately avoid or learn to avoid siblings, implying that such detection may not be critical to reproductive fitness. Inbreeding may persist in B anynana because the probability of encountering close relatives is rare in nature; that is, movement ecology may mask the deleterious effect of inbreeding resulting in relaxation of selection for active inbreeding avoidance behaviors. Butterflies feed primarily on nectar from flowers. Some also derive nourishment from pollen , tree sap, rotting fruit, dung, decaying flesh, and dissolved minerals in wet sand or dirt. Butterflies are important as pollinators for some species of plants. In general, they do not carry as much pollen load as bees , but they are capable of moving pollen over greater distances. Flower constancy has been observed for at least one species of butterfly. Adult butterflies consume only liquids, ingested through
6916-402: The list for an SSSI will only omit activities impossible on the particular SSSI (such as fishing where there is no water), and things requiring planning permission (which are covered by the local planning authority consultation process). In Scotland, and following the implementation of the Nature Conservation (Scotland) Act 2004, Scottish Natural Heritage (the former name for NatureScot) reviewed
7007-506: The majority of moths which fly by night, are often cryptically coloured (well camouflaged), and either hold their wings flat (touching the surface on which the moth is standing) or fold them closely over their bodies. Some day-flying moths, such as the hummingbird hawk-moth , are exceptions to these rules. Butterfly larvae , caterpillars , have a hard ( sclerotised ) head with strong mandibles used for cutting their food, most often leaves. They have cylindrical bodies, with ten segments to
7098-531: The most important features within each geological topic (or block ). Each of these sites is described, with most published in the Geological Conservation Review series, and so becomes a GCR site . Almost all GCR sites (but no other sites) are subsequently notified as geological SSSIs, except some that coincide with designated biological SSSI management units. A GCR site may contain features from several different topic blocks, for example
7189-401: The name is ancient, but modern Dutch and German use different words ( vlinder and Schmetterling ) and the common name often varies substantially between otherwise closely related languages. A possible source of the name is the bright yellow male of the brimstone ( Gonepteryx rhamni ); another is that butterflies were on the wing in meadows during the spring and summer butter season while
7280-400: The natural and artificial processes which resulted in their development and survival, for example the continued traditional grazing of heathland or chalk grassland . In England, the designating body for SSSIs, Natural England , selects biological SSSIs from within natural areas which are areas with particular landscape and ecological characteristics, or on a county basis. In Scotland,
7371-1003: The new Act, often with boundary changes. This complex process took some ten years to complete for the several thousand SSSIs. For the purposes of selecting the original tranche of SSSIs, Natural England's predecessors (the Nature Conservancy, the Nature Conservancy Council and English Nature ) used a system termed "areas of search" (AOSs). In England these were largely based on the 1974–1996 administrative counties (with larger counties divided into two or more areas), whereas in Scotland and Wales they are based around districts. The individual AOSs are between 400 km (150 sq mi) and 4,000 km (1,500 sq mi) in size. There were 59 AOSs in England, 12 in Wales, and 44 in Scotland. Watsonian vice-counties were formerly used for selection over
7462-399: The notification is then confirmed or withdrawn (in whole or part). At the time of the passing of the Wildlife and Countryside Act 1981 , many SSSIs were already in existence, having been notified over the previous decades under the National Parks and Access to the Countryside Act 1949 . Each of these was considered in turn, and either denotified, or renotified —brought under the provisions of
7553-447: The owners and occupiers of SSSIs. Previously, activities by "third parties" were not illegal under the SSSI legislation. This meant that damaging activities such as fly-tipping , intensive bait-digging or trail biking on an SSSI were only prevented if done (or permitted) by the owner or occupier – not if done by trespassers or under public rights. The effect was, for example, to allow control of legal trail biking on SSSIs (where damaging to
7644-529: The proboscis. They sip water from damp patches for hydration and feed on nectar from flowers, from which they obtain sugars for energy, and sodium and other minerals vital for reproduction. Several species of butterflies need more sodium than that provided by nectar and are attracted by sodium in salt; they sometimes land on people, attracted by the salt in human sweat. Some butterflies also visit dung and scavenge rotting fruit or carcasses to obtain minerals and nutrients. In many species, this mud-puddling behaviour
7735-517: The proposed operation and may, for example, limit its timing, location or intensity. The process is slightly different where the owner or occupier is a public body, but the effect is broadly similar. The relevant nature conservation body sends all SSSI owners and occupiers a site-specific 'site management statement' describing the ideal management (there may be grants available to help fund management). Owners and occupiers are encouraged to carry out this management, which in many (but not all) cases will be
7826-431: The quest for a suitable pupation site, often the underside of a leaf or other concealed location. There it spins a button of silk which it uses to fasten its body to the surface and moults for a final time. While some caterpillars spin a cocoon to protect the pupa, most species do not. The naked pupa, often known as a chrysalis, usually hangs head down from the cremaster, a spiny pad at the posterior end, but in some species
7917-572: The referendum to leave the EU in 2016, more than 450 staff have been transferred to the Department for Environment, Food and Rural Affairs (Defra). Neglected areas include Exmoor , the Lake District , the Pennines , and The Wash . The process of designating a site as of Special Scientific Interest is called notification; this is followed by consultation with the site's owners and occupiers, and
8008-565: The relevant community councils and community group having registered an interest in the land. The notification includes a description of the land and the natural features for which it is notified ("the citation"), a boundary map, and a list of the acts or omissions (activities) that the nature conservation body regulates through the issue of consents. The various laws protect the interest features of SSSIs from development, from other damage, and (since 2000 in England) also from neglect. Protection
8099-555: The rushes Juncus effusus and J. conglomeratus are present in damper areas. On the slopes dog's-mercury ( Mercurialis perennis ) and woodruff ( Galium odoratum ) are abundant, while more local species include nettle-leaved bellflower ( Campanula trachelium ), white ( Cephalanthera damasonium ), broad-leaved ( Epipactis helleborine ) and narrow-lipped ( E. leptochila )helleborines, yellow bird's-nest ( Monotropa hypopitys ) and scaly male-fern ( Dryopteris affinis ). Juniper scrub occurs on Windsor Hill itself, where it
8190-753: The slopes, the associated species are ash and whitebeam . Both even-aged and more mixed stands are represented, the latter typically with a shrub layer including gorse and honeysuckle on the plateau, and elder and hazel on the slopes. On the plateau, brambles , bracken and rosebay willowherb occupy extensive patches, but a more mixed ground flora occurs between these, including creeping soft-grass ( Holcus mollis ), hairy brome ( Bromus ramosus ), tufted hair-grass ( Deschampsia cespitosa ), wood millet ( Milium effusum ) and wood melick ( Melica uniflora ). Pill sedge ( Carex pilulifera ), hairy wood-rush ( Luzula pilosa ) and slender St. John's-wort ( Hypericum pulchrum ) are found in more heathy areas, and
8281-492: The whole of Great Britain. Butterflies Butterflies are winged insects from the lepidopteran suborder Rhopalocera , characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The group comprises the superfamilies Hedyloidea (moth-butterflies in the Americas) and Papilionoidea (all others). The oldest butterfly fossils have been dated to
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