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132-502: Termites are a group of detritophagous eusocial insects which consume a variety of decaying plant material , generally in the form of wood , leaf litter , and soil humus . They are distinguished by their moniliform antennae and the soft-bodied and often unpigmented worker caste for which they have been commonly termed " white ants "; however, they are not ants , to which they are only distantly related. About 2,972 extant species are currently described, 2,105 of which are members of

264-479: A Mediterranean climate . The south-east ranges from oceanic (Tasmania and coastal Victoria) to humid subtropical (upper half of New South Wales), with the highlands featuring alpine and subpolar oceanic climates . The interior is arid to semi-arid . Driven by climate change , average temperatures have risen more than 1°C since 1960 . Associated changes in rainfall patterns and climate extremes exacerbate existing issues such as drought and bushfires . 2019

396-525: A belt of Brigalow grasslands lie between the coast and the mountains, while inland of the dividing range are large areas of grassland and shrubland. These include the western plains of New South Wales, and the Mitchell Grass Downs and Mulga Lands of inland Queensland. The northernmost point of the mainland is the tropical Cape York Peninsula . The landscapes of the Top End and

528-824: A large wave of immigration from across Europe , with such immigrants referred to as " New Australians ". This required a relaxation of the white Australia policy , which was justified to Australians using the slogan "populate or perish". A member of the Western Bloc during the Cold War , Australia participated in the Korean War and the Malayan Emergency during the 1950s and the Vietnam War from 1962 to 1972. During this time, tensions over communist influence in society led to unsuccessful attempts by

660-417: A brief amount of time, so they also rely on their legs. The appearance of the legs is similar in each caste, but the soldiers have larger and heavier legs. The structure of the legs is consistent with other insects: the parts of a leg include a coxa , trochanter , femur , tibia and the tarsus . The number of tibial spurs on an individual's leg varies. Some species of termite have an arolium, located between

792-446: A certain developmental path. Workers may also be fertile or sterile, however the term "worker" is normally reserved for the latter, having evolved in taxa that exhibit a bifurcated developmental pathway. As a result, sterile workers like in the family Termitidae are termed true workers and are the most derived, while those that are undifferentiated and fertile as in the wood-nesting Archotermopsidae are termed pseudergates , which are

924-405: A crucial role in benthic ecosystems, forming essential food chains and participating in the nitrogen cycle . Detritivores and decomposers that reside in the desert live in burrows underground to avoid the hot surface since underground conditions provide favorable living conditions for them. Detritivores are the main organisms in clearing plant litter and recycling nutrients in the desert. Due to

1056-461: A date which later became Australia's national day . Most early settlers were convicts , transported for petty crimes and assigned as labourers or servants to "free settlers" (willing immigrants). Once emancipated , convicts tended to integrate into colonial society. Convict rebellions and uprisings were suppressed under martial law, which lasted for two years following the 1808 Rum Rebellion , Australia's only successful coup d'état . During

1188-425: A few hundred individuals to enormous societies with several million individuals. Most species are rarely seen, having a cryptic life-history where they remain hidden within the galleries and tunnels of their nests for most of their lives. Termites' success as a group has led to them colonizing almost every global landmass, with the highest diversity occurring in the tropics where they are estimated to constitute 10% of

1320-511: A fontanelle, which appears as a circular pore or series of pores in a depressed region within the middle of the head. The fontanelle connects to the frontal gland, a novel organ unique to Neoisopteran termites which evolved to excrete an array of defensive chemicals and secretions, and so is typically most developed in the soldier caste. Cellulose digestion in the family Termitidae has co-evolved with bacterial gut microbiota and many taxa have evolved additional symbiotic relationships such as with

1452-403: A much lesser extent in workers. The main distinction being that while larvae are wholly dependent on other nestmates to survive, workers are independent and are able to feed themselves and contribute to the colony. Workers remain wingless and across many taxa become developmentally arrested, appearing to not change into any other caste until death. In some basal taxa, there is no distinction, with

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1584-452: A pedicel (typically shorter than the scape), and the flagellum (all segments beyond the scape and pedicel). The mouth parts contain a maxillae , a labium, and a set of mandibles . The maxillae and labium have palps that help termites sense food and handling. The cuticle of most castes is soft and flexible due to a resulting lack of sclerotization, particularly of the abdomen which often appears translucent. Pigmentation and sclerotization of

1716-601: A process of increasing autonomy from the United Kingdom, highlighted by the Statute of Westminster Adoption Act 1942 , and culminating in the Australia Acts of 1986. Australia is a federal parliamentary democracy and constitutional monarchy comprising six states and ten territories . Its population of more than 28 million is highly urbanised and heavily concentrated on the eastern seaboard. Canberra

1848-524: A series of moults to become winged adults . Larvae , which are defined as early nymph instars with absent wing buds, exhibit the highest developmental potentiality and are able to molt into Alates , Soldiers , Neotenics , or Workers . Workers are believed to have evolved from larvae, sharing many similarities to the extent that workers can be regarded as "larval", in that both lack wings, eyes, and functional reproductive organs while maintaining varying levels of developmental flexibility, although usually to

1980-581: A settlement was established in Van Diemen's Land (present-day Tasmania ), and in 1813, Gregory Blaxland , William Lawson and William Wentworth crossed the Blue Mountains west of Sydney, opening the interior to European settlement. The British claim extended to the whole Australian continent in 1827 when Major Edmund Lockyer established a settlement on King George Sound (modern-day Albany ). The Swan River Colony (present-day Perth )

2112-562: A short distance off the north-east coast and extends for more than 2,000 km (1,200 mi). Mount Augustus , claimed to be the world's largest monolith, is located in Western Australia. At 2,228 m (7,310 ft), Mount Kosciuszko is the highest mountain on the Australian mainland. Even taller are Mawson Peak , at 2,745 m (9,006 ft), on the remote Australian external territory of Heard Island , and, in

2244-441: A significantly lower level of oxidative damage, including oxidative DNA damage , than workers, soldiers and nymphs. The lower levels of damage appear to be due to increased catalase , an enzyme that protects against oxidative stress . Detritivore Plant tissues are made up of resilient molecules (e.g. cellulose , lignin , xylan ) that decay at a much lower rate than other organic molecules. The activity of detritivores

2376-403: A termite is in flight, its wings remain at a right angle, and when the termite is at rest, its wings remain parallel to the body. Due to termites being hemimetabolous insects , where the young go through multiple and gradual adultoid molts before becoming an adult, the advent of eusociality has significantly altered the developmental patterns of this group of insects of which, although similar,

2508-484: A wide variety of landscapes and climates including deserts in the interior and tropical rainforests along the coast . The ancestors of Aboriginal Australians began arriving from south-east Asia 50,000 to 65,000 years ago, during the last glacial period . By the time of British settlement, Aboriginal Australians spoke 250 distinct languages and had the oldest living culture in the world. Australia's written history commenced with Dutch exploration of most of

2640-411: A wide variety of landscapes, with tropical rainforests in the north-east, mountain ranges in the south-east, south-west and east, and desert in the centre. The desert or semi-arid land commonly known as the outback makes up by far the largest portion of land. Australia is the driest inhabited continent; its annual rainfall averaged over continental area is less than 500 mm. The population density

2772-436: Is 3.4 inhabitants per square kilometre, although the large majority of the population lives along the temperate south-eastern coastline. The population density exceeds 19,500 inhabitants per square kilometre in central Melbourne. In 2021 Australia had 10% of the global permanent meadows and pastureland. Forest cover is around 17% of Australia's total land area. The Great Barrier Reef , the world's largest coral reef, lies

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2904-413: Is a constitutional monarchy , a parliamentary democracy and a federation . The country has maintained its mostly unchanged constitution alongside a stable liberal democratic political system since Federation in 1901. It is one of the world's oldest federations, in which power is divided between the federal and state governments. The Australian system of government combines elements derived from

3036-728: Is believed to be from the Upper Cretaceous in West Texas , where the oldest known faecal pellets were also discovered. Claims that termites emerged earlier have faced controversy. For example, F. M. Weesner indicated that the Mastotermitidae termites may go back to the Late Permian , 251 million years ago, and fossil wings that have a close resemblance to the wings of Mastotermes of the Mastotermitidae,

3168-464: Is consistent with all four-winged insects. The mesothorax and metathorax have well-developed exoskeletal plates; the prothorax has smaller plates. Termites have a ten-segmented abdomen with two plates, the tergites and the sternites . The tenth abdominal segment has a pair of short cerci . There are ten tergites, of which nine are wide and one is elongated. The reproductive organs are similar to those in cockroaches but are more simplified. For example,

3300-557: Is divided into the following clade and family groups, showing the subfamilies in their respective classification: The Neoisoptera , literally meaning "newer termites" (in an evolutionary sense), are a recently coined clade that include families such as the Rhinotermitidae and Termitidae . Neoisopterans have a bifurcated caste development with true workers, and so notably lack pseudergates (except in some basal taxa such as Serritermitidae : see below ). All Neoisopterans have

3432-543: Is expected to increase over time as a consequence of climate change . Increased urbanization and connectivity is also predicted to expand the range of some pest termites. The infraorder name Isoptera is derived from the Greek words iso (equal) and ptera (winged), which refers to the nearly equal size of the fore and hind wings. "Termite" derives from the Latin and Late Latin word termes ("woodworm, white ant"), altered by

3564-490: Is found mainly in the derived taxa (i.e. Neoisoptera ), and is believed to have evolved in tandem with the sterile worker caste as species moved to foraging for food beyond their nests, as opposed to the nest also being the food (such as in obligate wood-dwellers). There are three main castes which are discussed below: Worker termites undertake the most labor within the colony, being responsible for foraging, food storage, and brood and nest maintenance. Workers are tasked with

3696-422: Is not homologous to that of the eusocial Hymenoptera . Unlike ants, bees, and wasps which undergo a complete metamorphosis and as a result only exhibit developmental plasticity at the immobile larval stage , the mobile adultoid instars of termites remain developmentally flexible throughout all life stages up to the final molt , which has uniquely allowed for the evolution of distinct yet flexible castes amongst

3828-586: Is now Southeast Asia. It is uncertain how many waves of immigration may have contributed to these ancestors of modern Aboriginal Australians. The Madjedbebe rock shelter in Arnhem Land is possibly the oldest site showing the presence of humans in Australia. The oldest human remains found are the Lake Mungo remains , which have been dated to around 41,000 years ago. Aboriginal Australian culture

3960-629: Is one of only two pristine Archaean 3.6–2.7 Ga (billion years ago) crusts identified on the Earth. Having been part of all major supercontinents , the Australian continent began to form after the breakup of Gondwana in the Permian , with the separation of the continental landmass from the African continent and Indian subcontinent. It separated from Antarctica over a prolonged period beginning in

4092-466: Is one of the oldest continuous cultures on Earth. At the time of first European contact, Aboriginal Australians belonged to wide range of societies, with diverse economies spread across at least 250 different language groups . Estimates of the Aboriginal population before British settlement range from 300,000 to 3 million. Aboriginal Australians cultures were (and remain) deeply connected with

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4224-446: Is the nation's capital, while its most populous cities are Sydney and Melbourne , both with a population of more than 5 million. Australia's culture is diverse, and the country has one of the highest foreign-born populations in the world . It has a highly developed market economy and one of the highest per capita incomes globally . Its abundant natural resources and well-developed international trade relations are crucial to

4356-527: Is the reason why we do not see an accumulation of plant litter in nature. Detritivores are an important aspect of many ecosystems . They can live on any type of soil with an organic component, including marine ecosystems , where they are termed interchangeably with bottom feeders . Typical detritivorous animals include millipedes , springtails , woodlice , dung flies , slugs , many terrestrial worms , sea stars , sea cucumbers , fiddler crabs , and some sedentary marine Polychaetes such as worms of

4488-411: Is unique. About 85% of flowering plants, 84% of mammals, more than 45% of birds , and 89% of in-shore, temperate-zone fish are endemic . Australia has at least 755 species of reptile, more than any other country in the world. Besides Antarctica, Australia is the only continent that developed without feline species. Feral cats may have been introduced in the 17th century by Dutch shipwrecks, and later in

4620-521: The nymphal (winged) and apterous (wingless) lines. The bifurcation occurs early, either at the egg or the first two instars, and represents an irreversible and committed development to either the reproductive or non-reproductive lifestyles. As such, the apterous lineage consists mostly of wingless and truly altruistic sterile individuals (true workers, soldiers), whereas the nymphal lineage consists mainly of fertile individuals destined to become winged reproductives. The bifurcated developmental pathway

4752-475: The Archotermopsidae , which are able to retain high developmental plasticity even up to the late instars. In these basal taxa, the immatures are able to go through progressive ( nymph-to-imago ), regressive ( winged-to-wingless ) and stationary ( size increase, remains wingless ) molts, which typically indicates the developmental trajectory an individual follows. There is significant variation of

4884-479: The Australian continent , the island of Tasmania and numerous smaller islands . Australia has a total area of 7,688,287 km (2,968,464 sq mi), making it the sixth-largest country in the world and the largest country by area in Oceania . It is the world's oldest, flattest, and driest inhabited continent, with some of the least fertile soils. It is a megadiverse country , and its size gives it

5016-768: The Colonial Office that it be formally adopted. In 1824, the Admiralty agreed that the continent should be known officially by that name. The first official published use of the new name came with the publication in 1830 of The Australia Directory by the Hydrographic Office . Colloquial names for Australia include " Oz ", " Straya " and " Down Under ". Other epithets include "the Great Southern Land", "the Lucky Country" (from

5148-595: The Gulf Country —with their tropical climate—include forest, woodland, wetland, grassland, rainforest and desert. At the north-west corner of the continent are the sandstone cliffs and gorges of The Kimberley , and below that the Pilbara . The Victoria Plains tropical savanna lies south of the Kimberley and Arnhem Land savannas, forming a transition between the coastal savannas and the interior deserts. At

5280-551: The Indian Ocean Dipole and the El Niño–Southern Oscillation , which is correlated with periodic drought , and the seasonal tropical low-pressure system that produces cyclones in northern Australia. These factors cause rainfall to vary markedly from year to year. Much of the northern part of the country has a tropical, predominantly summer-rainfall ( monsoon ). The south-west corner of the country has

5412-539: The Iraq War from 2003 to 2009. The nation's trade relations also became increasingly oriented towards East Asia in the 21st century, with China becoming the nation's largest trading partner by a large margin. In 2020, during the COVID-19 pandemic , several of Australia's largest cities were locked down for extended periods and free movement across the national and state borders was restricted in an attempt to slow

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5544-664: The Menzies Government to ban the Communist Party of Australia , and a bitter split in the Labor Party in 1955. As a result of a 1967 referendum , the federal government gained the power to legislate with regard to Indigenous Australians, and Indigenous Australians were fully included in the census . Pre-colonial land interests (referred to as native title in Australia) was recognised in law for

5676-670: The Nasutes , which have a horn-like nozzle projection ( nasus ) on the head. These unique soldiers are able to spray noxious, sticky secretions containing diterpenes at their enemies. Nitrogen fixation plays an important role in Nasute nutrition. Soldiers are normally a committed sterile caste and so do not molt into anything else, but in certain basal taxa like the Archotermopsidae they are known to rarely molt into neotenic forms that develop functional sexual organs. In species with

5808-798: The Pacific Community ; the Commonwealth of Nations ; and the defence and security organisations ANZUS , AUKUS , and the Five Eyes . It is also a major non-NATO ally of the United States. The name Australia (pronounced / ə ˈ s t r eɪ l i ə / in Australian English ) is derived from the Latin Terra Australis ( ' southern land ' ), a name used for a hypothetical continent in

5940-532: The Permian and continuing through to the Cretaceous . When the last glacial period ended in about 10,000 BC, rising sea levels formed Bass Strait , separating Tasmania from the mainland. Then between about 8,000 and 6,500 BC, the lowlands in the north were flooded by the sea, separating New Guinea, the Aru Islands , and the mainland of Australia. The Australian continent is moving toward Eurasia at

6072-686: The Territory of Papua and New Guinea in 1949 and gained independence from Australia in 1975. In 1914, Australia joined the Allies in fighting the First World War, and took part in many of the major battles fought on the Western Front . Of about 416,000 who served, about 60,000 were killed and another 152,000 were wounded. Many Australians regard the defeat of the Australian and New Zealand Army Corps (ANZAC) at Gallipoli in 1915 as

6204-528: The Torres Strait Islands . The Dutch charted the whole of the western and northern coastlines and named the island continent " New Holland " during the 17th century, and although no attempt at settlement was made, a number of shipwrecks left men either stranded or, as in the case of the Batavia in 1629, marooned for mutiny and murder, thus becoming the first Europeans to permanently inhabit

6336-632: The bombing of Darwin and other Japanese attacks on Australian soil , led to a widespread belief in Australia that a Japanese invasion was imminent , and a shift from the United Kingdom to the United States as Australia's principal ally and security partner. Since 1951, Australia has been allied with the United States under the ANZUS treaty. In the decades following World War II, Australia enjoyed significant increases in living standards, leisure time and suburban development. Governments encouraged

6468-400: The claws , which is present in species that climb on smooth surfaces but is absent in most termites. Unlike in ants, the hind-wings and fore-wings are of equal length. Most of the time, the alates are poor flyers; their technique is to launch themselves in the air and fly in a random direction. Studies show that in comparison to larger termites, smaller termites cannot fly long distances. When

6600-426: The emu and the kookaburra . Australia is home to many dangerous animals including some of the most venomous snakes in the world. The dingo was introduced by Austronesian people who traded with Indigenous Australians around 3000 BCE . Many animal and plant species became extinct soon after first human settlement, including the Australian megafauna ; others have disappeared since European settlement, among them

6732-595: The governor-general at the federal level and by the governors at the state level, who by section 63 of the Constitution and convention act on the advice of their ministers. Thus, in practice the governor-general acts as a legal figurehead for the actions of the prime minister and the Cabinet. The governor-general may in some situations exercise reserve powers : powers exercisable in the absence or contrary to ministerial advice. When these powers may be exercised

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6864-501: The intromittent organ is not present in male alates, and the sperm is either immotile or aflagellate. However, Mastotermitidae termites have multiflagellate sperm with limited motility . The genitals in females are also simplified. Unlike in other termites, Mastotermitidae females have an ovipositor , a feature strikingly similar to that in female cockroaches. The non-reproductive castes of termites are wingless and rely exclusively on their six legs for locomotion. The alates fly only for

6996-418: The social Hymenoptera (ants and various species of bees and wasps), but their differing evolutionary origins result in major differences in life cycle. In the eusocial Hymenoptera, the workers are exclusively female. Males (drones) are haploid and develop from unfertilised eggs, while females (both workers and the queen) are diploid and develop from fertilised eggs. In contrast, worker termites, which constitute

7128-661: The "baptism of fire" that forged the new nation's identity . The beginning of the campaign is commemorated annually on Anzac Day , a date which rivals Australia Day as the nation's most important. From 1939 to 1945, Australia joined the Allies in fighting the Second World War. Australia's armed forces fought in the Pacific , European and Mediterranean and Middle East theatres . The shock of Britain's defeat in Singapore in 1942, followed soon after by

7260-587: The "island continent" and is sometimes considered the world's largest island . Australia has 34,218 km (21,262 mi) of coastline (excluding all offshore islands), and claims an extensive exclusive economic zone of 8,148,250 square kilometres (3,146,060 sq mi). This exclusive economic zone does not include the Australian Antarctic Territory . Mainland Australia lies between latitudes 9° and 44° south , and longitudes 112° and 154° east . Australia's size gives it

7392-434: The "king" and "queen" that together form a lifelong monogamous pair. Also unlike ants, which undergo a complete metamorphosis , termites undergo an incomplete metamorphosis that proceeds through egg, nymph , and adult stages. Termite colonies are commonly described as superorganisms due to the collective behaviors of the individuals which form a self-governing entity: the colony itself. Their colonies range in size from

7524-408: The "workers" (pseudergates) essentially being late instar larvae that retain the ability to change into all other castes. The development of larvae into adults can take months; the time period depends on food availability and nutrition, temperature, and the size of the colony. Since larvae and nymphs are unable to feed themselves, workers must feed them, but workers also take part in the social life of

7656-428: The 18th century by European settlers. They are now considered a major factor in the decline and extinction of many vulnerable and endangered native species. Seafaring immigrants from Asia are believed to have brought the dingo to Australia sometime after the end of the last ice age—perhaps 4000 years ago—and Aboriginal people helped disperse them across the continent as pets, contributing to the demise of thylacines on

7788-539: The 1964 book of the same name ), "the Sunburnt Country", and "the Wide Brown Land". The latter two both derive from Dorothea Mackellar 's 1908 poem " My Country ". Indigenous Australians comprise two broad groups: Human habitation of the Australian continent is estimated to have begun 50,000 to 65,000 years ago, with the migration of people by land bridges and short sea crossings from what

7920-593: The 2,972 extant termite species currently classified, 1,000 are found in Africa , where mounds are extremely abundant in certain regions. Approximately 1.1 million active termite mounds can be found in the northern Kruger National Park alone. In Asia , there are 435 species of termites, which are mainly distributed in China . Within China, termite species are restricted to mild tropical and subtropical habitats south of

8052-543: The 2018 Environmental Performance Index . There are more than 1,800 animals and plants on Australia's threatened species list, including more than 500 animals. Paleontologists discovered a fossil site of a prehistoric rainforest in McGraths Flat , in South Australia, that presents evidence that this now arid desert and dry shrubland / grassland was once home to an abundance of life. Australia

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8184-656: The Australian Antarctic Territory, Mount McClintock and Mount Menzies , at 3,492 m (11,457 ft) and 3,355 m (11,007 ft) respectively. Eastern Australia is marked by the Great Dividing Range , which runs parallel to the coast of Queensland, New South Wales and much of Victoria. The name is not strictly accurate, because parts of the range consist of low hills, and the highlands are typically no more than 1,600 m (5,200 ft) in height. The coastal uplands and

8316-558: The Australian mainland. The first ship and crew to chart the Australian coast and meet with Aboriginal people was the Duyfken , captained by Dutch navigator Willem Janszoon . He sighted the coast of Cape York Peninsula in early 1606, and made landfall on 26 February 1606 at the Pennefather River near the modern town of Weipa on Cape York. Later that year, Spanish explorer Luís Vaz de Torres sailed through and navigated

8448-684: The Paleozoic and Triassic insects tentatively classified as termites are in fact unrelated to termites and should be excluded from the Isoptera. Other studies suggest that the origin of termites is more recent, having diverged from Cryptocercus sometime during the Early Cretaceous . The primitive giant northern termite ( Mastotermes darwiniensis ) exhibits numerous cockroach-like characteristics that are not shared with other termites, such as laying its eggs in rafts and having anal lobes on

8580-709: The Southern Hemisphere since ancient times. Several 16th-century cartographers used the word Australia on maps, but not to identify modern Australia. When Europeans began visiting and mapping Australia in the 17th century, the name Terra Australis was applied to the new territories. Until the early 19th century, Australia was best known as New Holland , a name first applied by the Dutch explorer Abel Tasman in 1644 (as Nieuw-Holland ) and subsequently anglicised. Terra Australis still saw occasional usage, such as in scientific texts. The name Australia

8712-791: The Yangtze River. In Australia , all ecological groups of termites (dampwood, drywood, subterranean) are endemic to the country, with over 360 classified species. Because termites are highly social and abundant, they represent a disproportionate amount of the world's insect biomass . Termites and ants comprise about 1% of insect species, but represent more than 50% of insect biomass. Due to their soft cuticles, termites do not inhabit cool or cold habitats. There are three ecological groups of termites: dampwood, drywood and subterranean. Dampwood termites are found only in coniferous forests, and drywood termites are found in hardwood forests; subterranean termites live in widely diverse areas. One species in

8844-527: The animal biomass , particularly in Africa which has the richest diversity with more than 1000 described species. They are important decomposers of decaying plant matter in the subtropical and tropical regions of the world, and their recycling of wood and plant matter is of considerable ecological importance. Many species are ecosystem engineers capable of altering soil characteristics such as hydrology , decomposition, nutrient cycling , vegetative growth, and consequently surrounding biodiversity through

8976-404: The capacity to digest lignin , and so large deposits of dead plant tissue accumulated during this period, later becoming the fossil fuels . By feeding on sediments directly to extract the organic component, some detritivores incidentally concentrate toxic pollutants . Australia Australia , officially the Commonwealth of Australia , is a country comprising the mainland of

9108-632: The centre and Phanerozoic sedimentary basins , metamorphic and igneous rocks in the east. The Australian mainland and Tasmania are situated in the middle of the tectonic plate and have no active volcanoes, but due to passing over the East Australia hotspot , recent volcanism has occurred during the Holocene , in the Newer Volcanics Province of western Victoria and south-eastern South Australia. Volcanism also occurs in

9240-489: The coastline in the 17th-century. British colonisation began in 1788 with the establishment of the penal colony of New South Wales . By the mid-19th century, most of the continent had been explored by European settlers and five additional self-governing British colonies were established, each gaining responsible government by 1890. The colonies federated in 1901, forming the Commonwealth of Australia. This continued

9372-484: The colonies was achieved after a decade of planning, constitutional conventions and referendums , resulting in the establishment of the Commonwealth of Australia as a nation under the new Australian Constitution . After the 1907 Imperial Conference , Australia and several other self-governing British settler colonies were given the status of self-governing dominions within the British Empire. Australia

9504-419: The colony and have certain other tasks to accomplish such as foraging, building or maintaining the nest or tending to the queen. Pheromones regulate the caste system in termite colonies, preventing all but a very few of the termites from becoming fertile queens. Queens of the eusocial termite Reticulitermes speratus are capable of a long lifespan without sacrificing fecundity . These long-lived queens have

9636-847: The continent. In 1770, Captain James Cook sailed along and mapped the east coast, which he named " New South Wales " and claimed for Great Britain. Following the loss of its American colonies in 1783, the British Government sent a fleet of ships, the First Fleet , under the command of Captain Arthur Phillip , to establish a new penal colony in New South Wales . A camp was set up and the Union Flag raised at Sydney Cove , Port Jackson , on 26 January 1788,

9768-594: The convict system culminated in its abolition in the eastern colonies by the 1850s. Initially a free colony, Western Australia practised penal transportation from 1850 to 1868. The six colonies individually gained responsible government between 1855 and 1890, thus becoming elective democracies managing most of their own affairs while remaining part of the British Empire . The Colonial Office in London retained control of some matters, notably foreign affairs. In

9900-735: The country's economy. It ranks highly for quality of life, health, education, economic freedom, civil liberties and political rights. Australia is a middle power , and has the world's thirteenth-highest military expenditure . It is a member of international groups including the United Nations; the G20 ; the OECD ; the World Trade Organization ; Asia-Pacific Economic Cooperation ; the Pacific Islands Forum ;

10032-430: The cuticle correlates with life history , with species that spend more time in the surface in the open tending to have a more sclerotized and pigmented exoskeleton. Consistent with all insects, the anatomy of the termite thorax consists of three segments: the prothorax , the mesothorax and the metathorax . Each segment contains a pair of legs . On alates, the wings are located at the mesothorax and metathorax, which

10164-528: The descendants of the genus Cryptocercus . Some researchers have suggested a more conservative measure of retaining the termites as the Termitoidae, an epifamily within the cockroach order, which preserves the classification of termites at family level and below. Termites have long been accepted to be closely related to cockroaches and mantids , and they are classified in the same superorder ( Dictyoptera ). The oldest unambiguous termite fossils date to

10296-496: The developmental patterns in termites even across closely related taxa, but can typically be generalized into the following two patterns: The first is the linear developmental pathway , where all immatures are capable of developing into winged adults ( Alates ), exhibit high phenotypic potency, and where there exists no true sterile caste other than the soldier. The second is the bifurcated developmental pathway , where immatures diverge into two distinct developmental lineages known as

10428-417: The digestion of cellulose in food and are thus the most likely caste to be found in infested wood. The process of worker termites feeding other nestmates is known as trophallaxis . Trophallaxis is an effective nutritional tactic to convert and recycle nitrogenous components. It frees the parents from feeding all but the first generation of offspring, allowing for the group to grow much larger and ensuring that

10560-484: The drywood group is the West Indian drywood termite ( Cryptotermes brevis ) , which is an invasive species in Australia. Termites are usually small, measuring between 4 and 15 millimetres ( 3 ⁄ 16 and 9 ⁄ 16  in) in length. The largest of all extant termites are the queens of the species Macrotermes bellicosus , measuring up to over 10 centimetres (4 in) in length. Another giant termite,

10692-620: The early Cretaceous , but given the diversity of Cretaceous termites and early fossil records showing mutualism between microorganisms and these insects, they possibly originated earlier in the Jurassic or Triassic. Possible evidence of a Jurassic origin is the assumption that the extinct mammaliaform Fruitafossor from Morrison Formation consumed termites, judging from its morphological similarity to modern termite-eating mammals. Morrison Formation also yields social insect nest fossils close to that of termites. The oldest termite nest discovered

10824-459: The early 2010s after the 2000s Australian drought . Although most of Australia is semi-arid or desert, the continent includes a diverse range of habitats from alpine heaths to tropical rainforests . Fungi typify that diversity—an estimated 250,000 species—of which only 5% have been described—occur in Australia. Because of the continent's great age, extremely variable weather patterns, and long-term geographic isolation, much of Australia's biota

10956-678: The extinct Gyatermes styriensis , flourished in Austria during the Miocene and had a wingspan of 76 millimetres (3 in) and a body length of 25 millimetres (1 in). Most worker and soldier termites are completely blind as they do not have a pair of eyes. However, some species, such as Hodotermes mossambicus , have compound eyes which they use for orientation and to distinguish sunlight from moonlight. The alates (winged males and females) have eyes along with lateral ocelli . Lateral ocelli, however, are not found in all termites, absent in

11088-407: The families Hodotermitidae , Termopsidae , and Archotermopsidae . Like other insects, termites have a small tongue-shaped labrum and a clypeus ; the clypeus is divided into a postclypeus and anteclypeus. Termite antennae have a number of functions such as the sensing of touch, taste, odours (including pheromones), heat and vibration. The three basic segments of a termite antenna include a scape ,

11220-431: The family Terebellidae . Detritivores can be classified into more specific groups based on their size and biomes. Macrodetritivores are larger organisms such as millipedes, springtails, and woodlouse, while microdetritivores are smaller organisms such as bacteria. Scavengers are not typically thought to be detritivores, as they generally eat large quantities of organic matter, but both detritivores and scavengers are

11352-424: The family Termitidae . Termites comprise the infraorder Isoptera , or alternatively the epifamily Termitoidae , within the order Blattodea (along with cockroaches ). Termites were once classified in a separate order from cockroaches, but recent phylogenetic studies indicate that they evolved from cockroaches, as they are deeply nested within the group, and the sister group to wood eating cockroaches of

11484-1005: The first fully sequenced termite genome, of Zootermopsis nevadensis , which was published in the journal Nature Communications , consists of roughly 500Mb, while two subsequently published genomes, Macrotermes natalensis and Cryptotermes secundus , are considerably larger at around 1.3Gb. External phylogeny showing relationship of termites with other insect groups:  (Mantises) Blaberoidea Corydiodea Blattoidae Lamproblattidae Cryptocercidae (brown-hooded cockroaches) Isoptera (Termites) Internal phylogeny showing relationship of extant termite families: Mastotermitidae Stolotermitidae Hodotermopsidae Hodotermitidae Archotermopsidae Kalotermitidae Stylotermitidae Serritermitidae Rhinotermitidae Termitidae There are currently 3,173 living and fossil termite species recognised, classified in 12 families; reproductive and/or soldier castes are usually required for identification. The infraorder Isoptera

11616-772: The first time when the High Court of Australia held in Mabo v Queensland (No 2) that Australia was neither terra nullius ( ' land belonging to no one ' ) or "desert and uncultivated land" at the time of European settlement. Following the abolition of the last vestiges of the White Australia policy in 1973, Australia's demography and culture transformed as a result of a large and ongoing wave of non-European immigration, mostly from Asia. The late 20th century also saw an increasing focus on foreign policy ties with other Pacific Rim nations. The Australia Acts severed

11748-553: The forehead that lead to a gland which exudes defensive secretions, are a feature of the clade Neoisoptera and are present in all extant taxa such as Rhinotermitidae. The majority of termite species have mandibulate soldiers which are easily identified by the disproportionately large sclerotized head and mandibles. Among certain termites, the soldier caste has evolved globular (phragmotic) heads to block their narrow tunnels such as seen in Cryptotermes . Amongst mandibulate soldiers,

11880-699: The fungus Termitomyces ; in contrast, basal Neoisopterans and all other Euisoptera have flagellates and prokaryotes in their hindguts. Extant families and subfamilies are organized as follows: Termites are found on all continents except Antarctica . The diversity of termite species is low in North America and Europe (10 species known in Europe and 50 in North America), but is high in South America , where over 400 species are known. Of

12012-695: The genus Cryptocercus . Previous estimates suggested the divergence took place during the Jurassic or Triassic . More recent estimates suggest that they have an origin during the Late Jurassic , with the first fossil records in the Early Cretaceous . Similarly to ants and some bees and wasps from the separate order Hymenoptera , most termites have an analogous "worker" and "soldier" caste system consisting of mostly sterile individuals which are physically and behaviorally distinct. Unlike ants, most colonies begin from sexually mature individuals known as

12144-412: The heart of the country are the uplands of central Australia . Prominent features of the centre and south include Uluru (also known as Ayers Rock), the famous sandstone monolith, and the inland Simpson , Tirari and Sturt Stony , Gibson , Great Sandy, Tanami , and Great Victoria deserts, with the famous Nullarbor Plain on the southern coast. The Western Australian mulga shrublands lie between

12276-638: The highest phenotypic potency and can be described as totipotent ( able to molt into all alternative phenotypes ), whereas following instars can be pluripotent ( able to molt into reproductives and non-reproductives but cannot molt into at least one phenotype ), to multipotent ( able to molt into either reproductive or non-reproductive phenotypes ), to unipotent ( able to molt into developmentally close phenotypes ), and then finally committed ( no longer able to change phenotype, functionally an adult.) In most termites, phenotypic potency decreases with every successive molt. Notable exceptions are basal taxa such as

12408-436: The immatures. As a result the caste system of termites consists mostly of neotenous or juvenile individuals that undertake the most labor in the colony, which is in contrast to the eusocial Hymenoptera where work is strictly undertaken by the adults. The developmental plasticity in termites can be described similarly to cell potency , where each molt offers a varying level of phenotypic potency. Early instars typically exhibit

12540-545: The influence of Latin terere ("to rub, wear, erode") from the earlier word tarmes . A termite nest is also known as a termitary or termitarium (plural termitaria or termitariums ). The word was first used in English in 1781. Earlier attested designations were "wood ants" or "white ants", though these may never have been in wide use as termites do not exist in the British Isles . Termites were formerly placed in

12672-479: The interior deserts and Mediterranean-climate Southwest Australia . Lying on the Indo-Australian Plate , the mainland of Australia is the lowest and most primordial landmass on Earth with a relatively stable geological history. The landmass includes virtually all known rock types and from all geological time periods spanning more than 3.8 billion years of the Earth's history. The Pilbara Craton

12804-417: The island of New Guinea (considered geologically as part of the Australian continent), and in the Australian external territory of Heard Island and McDonald Islands . Seismic activity in the Australian mainland and Tasmania is also low, with the greatest number of fatalities having occurred in the 1989 Newcastle earthquake . The climate of Australia is significantly influenced by ocean currents, including

12936-543: The king will continue to mate with the queen throughout their lives. In some species, the abdomen of the queen swells up dramatically to increase fecundity , a characteristic known as physogastrism . Depending on the species, the queen starts producing reproductive alates at a certain time of the year, and huge swarms emerge from the colony when nuptial flight begins. These swarms attract a wide variety of predators. The queens can be particularly long-lived for insects, with some reportedly living as long as 30 or 50 years. In both

13068-620: The land and the environment, with stories of The Dreaming maintained through oral tradition , songs , dance and paintings. Certain groups engaged in fire-stick farming , fish farming , and built semi-permanent shelters . These practices have variously been characterised as " hunter-gatherer ", " agricultural ", "natural cultivation" and "intensification". Torres Strait Islander people first settled their islands at least 2,500 years ago. Culturally and linguistically distinct from mainland Aboriginal peoples, they were seafarers and obtained their livelihood from seasonal horticulture and

13200-576: The large mounds constructed by certain species. Termites have several impacts on humans. They are a delicacy in the diet of some human cultures such as the Makiritare in the Alto Orinoco province of Venezuela , where they are commonly used as a spice. They are also used in traditional medicinal treatments of various diseases and ailments, such as influenza, asthma, bronchitis, etc. Termites are most famous for being structural pests; however,

13332-829: The level of developmental plasticity even between closely related taxa, with many species having true workers that can molt into the other apterous castes such as ergatoids (worker reproductive; apterous neotenics), soldiers, or the other worker castes. Pseudergates sensu stricto are individuals which arise from the linear developmental pathway that have regressively molted and lost their wing buds, and are regarded as totipotent immatures. They are capable of performing work but are overall less involved in labor and considered more cooperative than truly altruistic. Pseudergates sensu lato , otherwise known as false workers , are most represented in basal lineages ( Kalotermitidae , Archotermopsidae , Hodotermopsidae , Serritermitidae ) and closely resemble true workers in which they also perform most of

13464-470: The limited vegetation available in the desert, desert detritivores adapted and evolved ways to feed in the extreme conditions of the desert. Detritivore feeding behaviour is affected by rainfall; moist soil increases detritivore feeding and excretion. Fungi, acting as decomposers, are important in today's terrestrial environment. During the Carboniferous period , fungi and bacteria had yet to evolve

13596-441: The linear and bifurcated developmental pathways, the primary reproductives only develop from winged immatures (nymphs). These winged immatures are capable of regressively molting into a form known as brachypterous neotenics ( nymphoids ), which retain juvenile and adult characteristics. BN 's can be found in both the derived and basal termite taxa, and generally serve as supplementary reproductives. Termites are often compared with

13728-437: The linear developmental pathway, soldiers develop from apterous immatures and constitute the only true sterile caste in these taxa. The primary reproductive caste of a colony consists of the fertile adult ( imago ) female and male individuals, colloquially known as the queen and king. The queen of the colony is responsible for egg production of the colony. Unlike in ants, the male and female reproductives form lifelong pairs where

13860-494: The location for the future federal capital of Canberra . While it was being constructed, Melbourne served as the temporary capital from 1901 to 1927. The Northern Territory was transferred from the control of South Australia to the Commonwealth in 1911. Australia became the colonial ruler of the Territory of Papua (which had initially been annexed by Queensland in 1883) in 1902 and of the Territory of New Guinea (formerly German New Guinea ) in 1920. The two were unified as

13992-448: The mainland. Australia is also one of 17 megadiverse countries. Australian forests are mostly made up of evergreen species, particularly eucalyptus trees in the less arid regions; wattles replace them as the dominant species in drier regions and deserts. Among well-known Australian animals are the monotremes (the platypus and echidna ); a host of marsupials , including the kangaroo , koala , and wombat , and birds such as

14124-649: The majority in a colony, are diploid individuals of both sexes and develop from fertilised eggs. Depending on species, male and female workers may have different roles in a termite colony. The life cycle of a termite begins with an egg , but is different from that of a bee or ant in that it goes through a developmental process called incomplete metamorphosis , going through multiple gradual pre-adult molts that are highly developmentally plastic before becoming an adult. Unlike in other hemimetabolous insects, nymphs are more strictly defined in termites as immature young with visible wing buds, which often invariably go through

14256-443: The mandibles have been adapted for a variety of defensive strategies: Biting/crushing ( Incisitermes ), slashing ( Cubitermes ), slashing/snapping ( Dentispicotermes ), symmetrical snapping ( Termes ), asymmetrical snapping ( Neocapritermes ), and piercing ( Armitermes ). In the more derived termite taxa, the soldier caste can be polymorphic and include minor and major forms. Other morphologically specialized soldiers includes

14388-474: The mid-19th century, explorers such as Burke and Wills charted Australia's interior. A series of gold rushes beginning in the early 1850s led to an influx of new migrants from China , North America and continental Europe, as well as outbreaks of bushranging and civil unrest; the latter peaked in 1854 when Ballarat miners launched the Eureka Rebellion against gold license fees. The 1860s saw

14520-411: The most basal. True workers are individuals which irreversibly develop from the apterous lineage and have completely forgo development into a winged adult. They display altruistic behaviors and either have terminal molts or exhibit a low level of phenotypical potency. True workers across different termite taxa ( Mastotermitidae , Hodotermitidae , Rhinotermitidae & Termitidae) can widely vary in

14652-544: The most primitive living termite, have been discovered in the Permian layers in Kansas. It is even possible that the first termites emerged during the Carboniferous . The folded wings of the fossil wood roach Pycnoblattina , arranged in a convex pattern between segments 1a and 2a, resemble those seen in Mastotermes , the only living insect with the same pattern. Kumar Krishna et al. , though, consider that all of

14784-459: The necessary gut symbionts are transferred from one generation to another. Workers are believed to have evolved from older wingless immatures ( Larvae ) that evolved cooperative behaviors; and indeed in some basal taxa the late instar larvae are known to undertake the role of workers without differentiating as a true separate caste. Workers can either be male or female, although in some species with polymorphic workers either sex may be restricted to

14916-602: The next two decades, social and economic reforms, together with the establishment of a Legislative Council and Supreme Court , saw the penal colony transition to a civil society. The indigenous population declined for 150 years following European settlement, mainly due to infectious disease. British colonial authorities did not sign any treaties with Aboriginal groups . As settlement expanded, tens of thousands of Indigenous people and thousands of settlers were killed in frontier conflicts while settlers dispossessed surviving Indigenous peoples of most of their land. In 1803,

15048-467: The order Isoptera. As early as 1934 suggestions were made that they were closely related to wood-eating cockroaches (genus Cryptocercus , the woodroach) based on the similarity of their symbiotic gut flagellates. In the 1960s additional evidence supporting that hypothesis emerged when F. A. McKittrick noted similar morphological characteristics between some termites and Cryptocercus nymphs . In 2008 DNA analysis from 16S rRNA sequences supported

15180-428: The political systems of the United Kingdom (a fused executive , constitutional monarchy and strong party discipline ) and the United States ( federalism , a written constitution and strong bicameralism with an elected upper house), resulting in a distinct hybrid. Federal government power is partially separated between three groups: Charles III reigns as King of Australia and is represented in Australia by

15312-578: The position of termites being nested within the evolutionary tree containing the order Blattodea , which included the cockroaches. The cockroach genus Cryptocercus shares the strongest phylogenetical similarity with termites and is considered to be a sister-group to termites. Termites and Cryptocercus share similar morphological and social features: for example, most cockroaches do not exhibit social characteristics, but Cryptocercus takes care of its young and exhibits other social behaviour such as trophallaxis and allogrooming . Termites are thought to be

15444-594: The protection of threatened species. Numerous protected areas have been created under the National Strategy for the Conservation of Australia's Biological Diversity to protect and preserve unique ecosystems; 65 wetlands are listed under the Ramsar Convention , and 16 natural World Heritage Sites have been established. Australia was ranked 21st out of 178 countries in the world on

15576-480: The rate of 6 to 7 centimetres a year. The Australian mainland's continental crust , excluding the thinned margins, has an average thickness of 38   km, with a range in thickness from 24 km to 59 km. Australia's geology can be divided into several main sections, showcasing that the continent grew from west to east: the Archaean cratonic shields found mostly in the west, Proterozoic fold belts in

15708-545: The remaining constitutional ties between Australia and the United Kingdom while maintaining the monarch in her independent capacity as Queen of Australia . In a 1999 constitutional referendum , 55% of voters rejected abolishing the monarchy and becoming a republic. Following the September 11 attacks on the United States, Australia joined the United States in fighting the Afghanistan War from 2001 to 2021 and

15840-419: The resources of their reefs and seas. Agriculture also developed on some islands and villages appeared by the 1300s. By the mid-18th century in northern Australia, contact, trade and cross-cultural engagement had been established between local Aboriginal groups and Makassan trepangers , visiting from present-day Indonesia. The Dutch are the first Europeans that recorded sighting and making landfall on

15972-491: The rise of blackbirding , where South Sea Islanders were coerced or abducted into indentured labour, mainly by Queensland colonists. From 1886, Australian colonial governments began removing many Aboriginal children from their families and communities, justified on the grounds of child protection and forced assimilation policies. The Second Boer War (1899–1902) marked the largest overseas deployment of Australia's colonial forces . On 1 January 1901, federation of

16104-485: The same type of cases of consumer-resource systems . The consumption of wood, whether alive or dead, is known as xylophagy . The activity of animals feeding only on dead wood is called sapro -xylophagy and those animals, sapro-xylophagous. Detritivores play an important role as recyclers in the ecosystem's energy flow and biogeochemical cycles . Alongside decomposers, they reintroduce vital elements such as carbon, nitrogen, phosphorus, calcium, and potassium back into

16236-416: The soil, allowing plants to take in these elements and use them for growth. They shred the dead plant matter which releases the trapped nutrients in the plant tissues. An abundance of detritivores in the soil allows the ecosystem to efficiently recycle nutrients. Many detritivores live in mature woodland , though the term can be applied to certain bottom-feeders in wet environments . These organisms play

16368-741: The spread of the SARS-CoV-2 virus . Surrounded by the Indian and Pacific oceans, Australia is separated from Asia by the Arafura and Timor seas, with the Coral Sea lying off the Queensland coast, and the Tasman Sea lying between Australia and New Zealand. The world's smallest continent and sixth-largest country by total area , Australia—owing to its size and isolation—is often dubbed

16500-406: The thylacine. Many of Australia's ecoregions, and the species within those regions, are threatened by human activities and introduced animal, chromistan , fungal and plant species. All these factors have led to Australia's having the highest mammal extinction rate of any country in the world. The federal Environment Protection and Biodiversity Conservation Act 1999 is the legal framework for

16632-465: The vast majority of termite species are innocuous, with the regional numbers of economically significant species being: North America , 9; Australia , 16; Indian subcontinent , 26; tropical Africa , 24; Central America and the West Indies , 17. Of known pest species, 28 of the most invasive and structurally damaging belong to the genus Coptotermes . The distribution of most known pest species

16764-442: The wings. It has been proposed that the Isoptera and Cryptocercidae be grouped in the clade " Xylophagodea ". Termites are sometimes called "white ants", but the only resemblance to the ants is due to their sociality which is due to convergent evolution with termites being the first social insects to evolve a caste system more than 100 million years ago. Termite genomes are generally relatively large compared to those of other insects;

16896-570: The work and are similarly altruistic, however differ in developing from the linear developmental pathway where they exist in a stationary molt; i.e they have halted development before the growth of wing buds, and are regarded as pluripotent immatures. The soldier caste is the most anatomically and behaviorally specialized, and their sole purpose is to defend the colony. Many soldiers have large heads with highly modified powerful jaws so enlarged that they cannot feed themselves. Instead, like juveniles, they are fed by workers. Fontanelles , simple holes in

17028-615: Was Australia's warmest recorded year, and the 2019–2020 bushfire season was the country's worst on record . Australia's greenhouse gas emissions per capita are among the highest in the world. Water restrictions are frequently in place in many regions and cities of Australia in response to chronic shortages due to urban population increases and localised drought. Throughout much of the continent, major flooding regularly follows extended periods of drought, flushing out inland river systems, overflowing dams and inundating large inland flood plains, as occurred throughout Eastern Australia in

17160-413: Was established in 1829, evolving into the largest Australian colony by area, Western Australia . In accordance with population growth, separate colonies were carved from New South Wales: Tasmania in 1825, South Australia in 1836, New Zealand in 1841, Victoria in 1851, and Queensland in 1859. South Australia was founded as a free colony—it never accepted transported convicts. Growing opposition to

17292-579: Was one of the founding members of the League of Nations in 1920, and the United Nations in 1945. The Statute of Westminster 1931 formally ended the ability of the UK to pass federal laws without Australia's consent. Australia adopted it in 1942, but it was backdated to 1939 to confirm the validity of legislation passed during World War II. The Australian Capital Territory was formed in 1911 as

17424-470: Was popularised by the explorer Matthew Flinders , who said it was "more agreeable to the ear, and an assimilation to the names of the other great portions of the Earth". The first time that Australia appears to have been officially used was in April 1817, when Governor Lachlan Macquarie acknowledged the receipt of Flinders' charts of Australia from Lord Bathurst . In December 1817, Macquarie recommended to

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