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Canadian French ( French : français canadien , pronounced [fʁãˈsɛ kanaˈd͡zjɛ̃] ) is the French language as it is spoken in Canada. It includes multiple varieties , the most prominent of which is Québécois ( Quebec French ). Formerly Canadian French referred solely to Quebec French and the closely related varieties of Ontario ( Franco-Ontarian ) and Western Canada —in contrast with Acadian French , which is spoken by Acadians in New Brunswick (including the Chiac dialect ) and some areas of Nova Scotia (including the dialect St. Marys Bay French ), Prince Edward Island and Newfoundland & Labrador (where Newfoundland French is also spoken).

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116-565: In 2011, the total number of native French speakers in Canada was around 7.3 million (22% of the entire population), while another 2 million spoke it as a second language. At the federal level, it has official status alongside English . At the provincial level, French is the sole official language of Quebec as well as one of two official languages of New Brunswick and jointly official (derived from its federal legal status) in Nunavut , Yukon and

232-553: A mixed language or a creole —a theory called the Middle English creole hypothesis . Although the great influence of these languages on the vocabulary and grammar of Modern English is widely acknowledged, most specialists in language contact do not consider English to be a true mixed language. English is classified as a Germanic language because it shares innovations with other Germanic languages including Dutch , German , and Swedish . These shared innovations show that

348-644: A 2012 official Eurobarometer poll (conducted when the UK was still a member of the EU), 38 percent of the EU respondents outside the countries where English is an official language said they could speak English well enough to have a conversation in that language. The next most commonly mentioned foreign language, French (which is the most widely known foreign language in the UK and Ireland), could be used in conversation by 12 percent of respondents. A working knowledge of English has become

464-728: A bacterial genome over three types of outbreak contact networks—homogeneous, super-spreading, and chain-like. They summarized the resulting phylogenies with five metrics describing tree shape. Figures 2 and 3 illustrate the distributions of these metrics across the three types of outbreaks, revealing clear differences in tree topology depending on the underlying host contact network. Super-spreader networks give rise to phylogenies with higher Colless imbalance, longer ladder patterns, lower Δw, and deeper trees than those from homogeneous contact networks. Trees from chain-like networks are less variable, deeper, more imbalanced, and narrower than those from other networks. Scatter plots can be used to visualize

580-499: A disproof of a previously widely accepted theory. During the late 19th century, Ernst Haeckel 's recapitulation theory , or "biogenetic fundamental law", was widely popular. It was often expressed as " ontogeny recapitulates phylogeny", i.e. the development of a single organism during its lifetime, from germ to adult, successively mirrors the adult stages of successive ancestors of the species to which it belongs. But this theory has long been rejected. Instead, ontogeny evolves  –

696-658: A distinct language from Modern English and is virtually impossible for 21st-century unstudied English-speakers to understand. Its grammar was similar to that of modern German: nouns, adjectives, pronouns, and verbs had many more inflectional endings and forms , and word order was much freer than in Modern English. Modern English has case forms in pronouns ( he , him , his ) and has a few verb inflections ( speak , speaks , speaking , spoke , spoken ), but Old English had case endings in nouns as well, and verbs had more person and number endings. Its closest relative

812-490: A foreign language, make up the "expanding circle". The distinctions between English as a first language, as a second language, and as a foreign language are often debatable and may change in particular countries over time. For example, in the Netherlands and some other countries of Europe, knowledge of English as a second language is nearly universal, with over 80 percent of the population able to use it, and thus English

928-536: A language as an evolutionary system. The evolution of human language closely corresponds with human's biological evolution which allows phylogenetic methods to be applied. The concept of a "tree" serves as an efficient way to represent relationships between languages and language splits. It also serves as a way of testing hypotheses about the connections and ages of language families. For example, relationships among languages can be shown by using cognates as characters. The phylogenetic tree of Indo-European languages shows

1044-520: A more standard version of English. They have many more speakers of English who acquire English as they grow up through day-to-day use and listening to broadcasting, especially if they attend schools where English is the medium of instruction. Varieties of English learned by non-native speakers born to English-speaking parents may be influenced, especially in their grammar, by the other languages spoken by those learners. Most of those varieties of English include words little used by native speakers of English in

1160-695: A number of other Anglic languages, including Scots and the extinct Fingallian dialect and Yola language of Ireland. Like Icelandic and Faroese , the development of English in the British Isles isolated it from the continental Germanic languages and influences, and it has since diverged considerably. English is not mutually intelligible with any continental Germanic language, as it differs in vocabulary , syntax , and phonology . However, some of these, such as Dutch or Frisian, do show strong affinities with English, especially with its earlier stages. Unlike Icelandic and Faroese, which were isolated,

1276-417: A phylogenetic tree can be living taxa or fossils , which represent the present time or "end" of an evolutionary lineage, respectively. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does not show the origin or "root" of the taxa in question or

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1392-684: A requirement in a number of occupations and professions such as medicine and computing. English has become so important in scientific publishing that more than 80 percent of all scientific journal articles indexed by Chemical Abstracts in 1998 were written in English, as were 90 percent of all articles in natural science publications by 1996 and 82 percent of articles in humanities publications by 1995. International communities such as international business people may use English as an auxiliary language , with an emphasis on vocabulary suitable for their domain of interest. This has led some scholars to develop

1508-407: A rich inflectional morphology and relatively free word order to a mostly analytic pattern with little inflection and a fairly fixed subject–verb–object word order . Modern English relies more on auxiliary verbs and word order for the expression of complex tenses , aspects and moods , as well as passive constructions , interrogatives , and some negation . The earliest form of English

1624-700: A shared evolutionary history. There are debates if increasing the number of taxa sampled improves phylogenetic accuracy more than increasing the number of genes sampled per taxon. Differences in each method's sampling impact the number of nucleotide sites utilized in a sequence alignment, which may contribute to disagreements. For example, phylogenetic trees constructed utilizing a more significant number of total nucleotides are generally more accurate, as supported by phylogenetic trees' bootstrapping replicability from random sampling. The graphic presented in Taxon Sampling, Bioinformatics, and Phylogenomics , compares

1740-758: A significant minority speaks English. The countries with the most native English speakers are, in descending order, the United States (at least 231 million), the United Kingdom (60 million), Canada (19 million), Australia (at least 17 million), South Africa (4.8 million), Ireland (4.2 million), and New Zealand (3.7 million). In these countries, children of native speakers learn English from their parents, and local people who speak other languages and new immigrants learn English to communicate in their neighbourhoods and workplaces. The inner-circle countries provide

1856-462: A significant source of error within phylogenetic analysis occurs due to inadequate taxon samples. Accuracy may be improved by increasing the number of genetic samples within its monophyletic group. Conversely, increasing sampling from outgroups extraneous to the target stratified population may decrease accuracy. Long branch attraction is an attributed theory for this occurrence, where nonrelated branches are incorrectly classified together, insinuating

1972-631: A single tree with true claim. The same process can be applied to texts and manuscripts. In Paleography , the study of historical writings and manuscripts, texts were replicated by scribes who copied from their source and alterations - i.e., 'mutations' - occurred when the scribe did not precisely copy the source. Phylogenetics has been applied to archaeological artefacts such as the early hominin hand-axes, late Palaeolithic figurines, Neolithic stone arrowheads, Bronze Age ceramics, and historical-period houses. Bayesian methods have also been employed by archaeologists in an attempt to quantify uncertainty in

2088-594: A small group of taxa to represent the evolutionary history of its broader population. This process is also known as stratified sampling or clade-based sampling. The practice occurs given limited resources to compare and analyze every species within a target population. Based on the representative group selected, the construction and accuracy of phylogenetic trees vary, which impacts derived phylogenetic inferences. Unavailable datasets, such as an organism's incomplete DNA and protein amino acid sequences in genomic databases, directly restrict taxonomic sampling. Consequently,

2204-592: A species to uncover either a higher abundance of important bioactive compounds (e.g., species of Taxus for taxol) or natural variants of known pharmaceuticals (e.g., species of Catharanthus for different forms of vincristine or vinblastine). Phylogenetic analysis has also been applied to biodiversity studies within the fungi family. Phylogenetic analysis helps understand the evolutionary history of various groups of organisms, identify relationships between different species, and predict future evolutionary changes. Emerging imagery systems and new analysis techniques allow for

2320-447: A verb ending ( present plural): From the 8th to the 11th centuries, Old English gradually transformed through language contact with Old Norse in some regions. The waves of Norse (Viking) colonisation of northern parts of the British Isles in the 8th and 9th centuries put Old English into intense contact with Old Norse , a North Germanic language. Norse influence was strongest in the north-eastern varieties of Old English spoken in

2436-563: Is Old Frisian , but even some centuries after the Anglo-Saxon migration, Old English retained considerable mutual intelligibility with other Germanic varieties. Even in the 9th and 10th centuries, amidst the Danelaw and other Viking invasions, there is historical evidence that Old Norse and Old English retained considerable mutual intelligibility, although probably the northern dialects of Old English were more similar to Old Norse than

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2552-550: Is "tree shape." These approaches, while computationally intensive, have the potential to provide valuable insights into pathogen transmission dynamics. The structure of the host contact network significantly impacts the dynamics of outbreaks, and management strategies rely on understanding these transmission patterns. Pathogen genomes spreading through different contact network structures, such as chains, homogeneous networks, or networks with super-spreaders, accumulate mutations in distinct patterns, resulting in noticeable differences in

2668-400: Is also the most widely learned second language in the world, with more second-language speakers than native speakers. English is either the official language or one of the official languages in 59 sovereign states (such as India , Ireland , and Canada ). In some other countries, it is the sole or dominant language for historical reasons without being explicitly defined by law (such as in

2784-466: Is also undergoing change under the influence of American English, fuelled by the strong presence of American English in the media and the prestige associated with the United States as a world power. As of 2016 , 400 million people spoke English as their first language , and 1.1 billion spoke it as a secondary language. English is the largest language by number of speakers . English

2900-610: Is also widely used in media and literature, and the number of English language books published annually in India is the third largest in the world after the US and UK. However, English is rarely spoken as a first language, numbering only around a couple hundred-thousand people, and less than 5% of the population speak fluent English in India. David Crystal claimed in 2004 that, combining native and non-native speakers, India now has more people who speak or understand English than any other country in

3016-407: Is an official language of countries populated by few descendants of native speakers of English. It has also become by far the most important language of international communication when people who share no native language meet anywhere in the world. The Indian linguist Braj Kachru distinguished countries where English is spoken with a three circles model . In his model, Kachru based his model on

3132-461: Is believed to have resulted from a localized levelling of contact dialects between Québécois and Acadian settlers . There are two main sub-varieties of Canadian French. Joual is an informal variety of French spoken in working-class neighbourhoods in Quebec. Chiac is a blending of Acadian French syntax and vocabulary, with numerous lexical borrowings from English. The term "Canadian French"

3248-538: Is called Old English or Anglo-Saxon ( c.  450–1150 ). Old English developed from a set of West Germanic dialects, often grouped as Anglo-Frisian or North Sea Germanic , and originally spoken along the coasts of Frisia , Lower Saxony and southern Jutland by Germanic peoples known to the historical record as the Angles , Saxons , and Jutes . From the 5th century, the Anglo-Saxons settled Britain as

3364-507: Is endangered—both Quebec French and Acadian French are now more widely spoken among Newfoundland Francophones than the distinctive peninsular dialect. Brayon French is spoken in Madawaska County , New Brunswick, and, to a lesser extent, Aroostook County , Maine, and Beauce of Quebec. Although superficially a phonological descendant of Acadian French, analysis reveals it is morphosyntactically identical to Quebec French. It

3480-536: Is often arbitrarily defined as beginning with the conquest of England by William the Conqueror in 1066, but it developed further in the period from 1150 to 1500. With the Norman conquest of England in 1066, the now-Norsified Old English language was subject to another wave of intense contact, this time with Old French , in particular Old Norman French , influencing it as a superstrate. The Norman French spoken by

3596-424: Is routinely used to communicate with foreigners and often in higher education. In these countries, although English is not used for government business, its widespread use puts them at the boundary between the "outer circle" and "expanding circle". English is unusual among world languages in how many of its users are not native speakers but speakers of English as a second or foreign language. Many users of English in

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3712-517: Is spoken by communities on every continent and on islands in all the major oceans. The countries where English is spoken can be grouped into different categories according to how English is used in each country. The "inner circle" countries with many native speakers of English share an international standard of written English and jointly influence speech norms for English around the world. English does not belong to just one country, and it does not belong solely to descendants of English settlers. English

3828-889: Is spoken by over 350,000 Acadians in parts of the Maritime Provinces , Newfoundland, the Magdalen Islands , the Lower North Shore and the Gaspé Peninsula . St. Marys Bay French is a variety of Acadian French spoken in Nova Scotia. Métis French is spoken in Manitoba and Western Canada by the Métis , descendants of First Nations mothers and voyageur fathers during the fur trade . Many Métis spoke Cree in addition to French, and over

3944-626: Is spoken in Quebec. Closely related varieties are spoken by Francophone communities in Ontario, Western Canada and the New England region of the United States, differing only from Quebec French primarily by their greater linguistic conservatism . The term Laurentian French has limited applications as a collective label for all these varieties, and Quebec French has also been used for the entire dialect group. The overwhelming majority of francophone Canadians speak this dialect. Acadian French

4060-437: Is the identification, naming, and classification of organisms. Compared to systemization, classification emphasizes whether a species has characteristics of a taxonomic group. The Linnaean classification system developed in the 1700s by Carolus Linnaeus is the foundation for modern classification methods. Linnaean classification relies on an organism's phenotype or physical characteristics to group and organize species. With

4176-444: Is the study of the evolutionary history of life using genetics, which is known as phylogenetic inference . It establishes the relationship between organisms with the empirical data and observed heritable traits of DNA sequences, protein amino acid sequences, and morphology . The results are a phylogenetic tree —a diagram setting the hypothetical relationships between organisms and their evolutionary history. The tips of

4292-777: Is the third person pronoun group beginning with th- ( they, them, their ) which replaced the Anglo-Saxon pronouns with h- ( hie, him, hera ). Other core Norse loanwords include "give", "get", "sky", "skirt", "egg", and "cake", typically displacing a native Anglo-Saxon equivalent. Old Norse in this era retained considerable mutual intelligibility with some dialects of Old English, particularly northern ones. Englischmen þeyz hy hadde fram þe bygynnyng þre manner speche, Souþeron, Northeron, and Myddel speche in þe myddel of þe lond, ... Noþeles by comyxstion and mellyng, furst wiþ Danes, and afterward wiþ Normans, in menye þe contray longage ys asperyed, and som vseþ strange wlaffyng, chyteryng, harryng, and garryng grisbytting. Although, from

4408-628: The Office québécois de la langue française distinguishes between different kinds of anglicisms: Academic, colloquial , and pejorative terms are used in Canada to refer to the vernacular . Examples are des "sabirisation" (from sabir , " pidgin "), Franglais , Français québécois , and Canadian French. English language English is a West Germanic language in the Indo-European language family , whose speakers, called Anglophones , originated in early medieval England on

4524-485: The Augustinian canon Orrm , which highlights the blending of both Old English and Anglo-Norman elements in English for the first time. In Wycliff'e Bible of the 1380s, the verse Matthew 8:20 was written: Foxis han dennes, and briddis of heuene han nestis . Here the plural suffix -n on the verb have is still retained, but none of the case endings on the nouns are present. By the 12th century Middle English

4640-546: The Danelaw area around York, which was the centre of Norse colonisation; today these features are still particularly present in Scots and Northern English . The centre of Norsified English was in the Midlands around Lindsey . After 920 CE, when Lindsey was incorporated into the Anglo-Saxon polity, English spread extensively throughout the region. An element of Norse influence that continues in all English varieties today

4756-552: The European Free Trade Association , Association of Southeast Asian Nations (ASEAN), and Asia-Pacific Economic Cooperation (APEC) set English as their organisation's sole working language even though most members are not countries with a majority of native English speakers. While the European Union (EU) allows member states to designate any of the national languages as an official language of

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4872-536: The Germanic language branch, and as of 2021 , Ethnologue estimated that there were over 1.5 billion speakers worldwide. Old English emerged from a group of West Germanic dialects spoken by the Anglo-Saxons . Late Old English borrowed some grammar and core vocabulary from Old Norse , a North Germanic language . Then, Middle English borrowed vocabulary extensively from French dialects , which are

4988-767: The Low Saxon and Frisian languages . English is an Indo-European language and belongs to the West Germanic group of the Germanic languages . Old English originated from a Germanic tribal and linguistic continuum along the Frisian North Sea coast, whose languages gradually evolved into the Anglic languages in the British Isles , and into the Frisian languages and Low German /Low Saxon on

5104-623: The Northwest Territories . Government services are offered in French at select localities in Manitoba , Ontario (through the French Language Services Act ) and, to a lesser extent, elsewhere in the country, depending largely on the proximity to Quebec and/or French Canadian influence on any given region. In New Brunswick, all government services must be available in both official languages. Quebec French

5220-668: The United Nations at the end of World War II , English had become pre-eminent and is now the main worldwide language of diplomacy and international relations. It is one of six official languages of the United Nations. Many other worldwide international organisations, including the International Olympic Committee , specify English as a working language or official language of the organisation. Many regional international organisations such as

5336-518: The palatalisation of consonants that were velar consonants in Proto-Germanic (see Phonological history of Old English § Palatalization ). The earliest varieties of an English language, collectively known as Old English or "Anglo-Saxon", evolved from a group of North Sea Germanic dialects brought to Britain in the 5th century. Old English dialects were later influenced by Old Norse -speaking Viking invaders and settlers , starting in

5452-564: The "outer circle" countries are countries such as the Philippines , Jamaica , India , Pakistan , Singapore , Malaysia and Nigeria with a much smaller proportion of native speakers of English but much use of English as a second language for education, government, or domestic business, and its routine use for school instruction and official interactions with the government. Those countries have millions of native speakers of dialect continua ranging from an English-based creole to

5568-617: The 8th and 9th centuries. Middle English began in the late 11th century after the Norman Conquest of England, when a considerable amount of Old French vocabulary was incorporated into English over some three centuries. Early Modern English began in the late 15th century with the start of the Great Vowel Shift and the Renaissance trend of borrowing further Latin and Greek words and roots, concurrent with

5684-578: The Americas whereas Acadian French, Cajun French, and Newfoundland French are derivatives of non-koiné local dialects in France. The term anglicism ( anglicisme ) is related to the linguistic concepts of loanwords , barbarism , diglossia , and the macaronic mixture of the French and English languages. According to some, French spoken in Canada includes many anglicisms. The " Banque de dépannage linguistique " (Language Troubleshooting Database) by

5800-608: The Angles. English may have a small amount of substrate influence from Common Brittonic, and a number of possible Brittonicisms in English have been proposed, but whether most of these supposed Brittonicisms are actually a direct result of Brittonic substrate influence is disputed. Old English was divided into four dialects: the Anglian dialects ( Mercian and Northumbrian ) and the Saxon dialects ( Kentish and West Saxon ). Through

5916-524: The British Empire in the 1950s and 1960s, former colonies often did not reject English but rather continued to use it as independent countries setting their own language policies. For example, the view of the English language among many Indians has gone from associating it with colonialism to associating it with economic progress, and English continues to be an official language of India. English

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6032-547: The Early Modern period includes the works of William Shakespeare and the translation of the Bible commissioned by King James I . Even after the vowel shift the language still sounded different from Modern English: for example, the consonant clusters /kn ɡn sw/ in knight , gnat , and sword were still pronounced. Many of the grammatical features that a modern reader of Shakespeare might find quaint or archaic represent

6148-655: The English Language , which introduced standard spellings of words and usage norms. In 1828, Noah Webster published the American Dictionary of the English language to try to establish a norm for speaking and writing American English that was independent of the British standard. Within Britain, non-standard or lower class dialect features were increasingly stigmatised, leading to the quick spread of

6264-557: The English-speaking inner circle countries outside Britain helped level dialect distinctions and produce koineised forms of English in South Africa, Australia, and New Zealand. The majority of immigrants to the United States without British ancestry rapidly adopted English after arrival. Now the majority of the United States population are monolingual English speakers. English has ceased to be an "English language" in

6380-558: The German Phylogenie , introduced by Haeckel in 1866, and the Darwinian approach to classification became known as the "phyletic" approach. It can be traced back to Aristotle , who wrote in his Posterior Analytics , "We may assume the superiority ceteris paribus [other things being equal] of the demonstration which derives from fewer postulates or hypotheses." The modern concept of phylogenetics evolved primarily as

6496-554: The Roman economy and administration collapsed . By the 7th century, this Germanic language of the Anglo-Saxons became dominant in Britain , replacing the languages of Roman Britain (43–409): Common Brittonic , a Celtic language , and British Latin , brought to Britain by the Roman occupation. At this time, these dialects generally resisted influence from the then-local Brittonic and Latin languages. England and English (originally Ænglaland and Ænglisc ) are both named after

6612-494: The Union, in practice English is the main working language of EU organisations. Although in most countries English is not an official language, it is currently the language most often taught as a foreign language . In the countries of the EU, English is the most widely spoken foreign language in nineteen of the twenty-five member states where it is not an official language (that is, the countries other than Ireland and Malta ). In

6728-519: The United States and United Kingdom ). It is a co-official language of the United Nations , the European Union , and many other international and regional organisations. It has also become the de facto lingua franca of diplomacy, science , technology, international trade, logistics, tourism, aviation, entertainment, and the Internet . English accounts for at least 70% of total speakers of

6844-645: The absence of genetic recombination . Phylogenetics can also aid in drug design and discovery. Phylogenetics allows scientists to organize species and can show which species are likely to have inherited particular traits that are medically useful, such as producing biologically active compounds - those that have effects on the human body. For example, in drug discovery, venom -producing animals are particularly useful. Venoms from these animals produce several important drugs, e.g., ACE inhibitors and Prialt ( Ziconotide ). To find new venoms, scientists turn to phylogenetics to screen for closely related species that may have

6960-407: The base from which English spreads to other countries in the world. Estimates of the numbers of second language and foreign-language English speakers vary greatly from 470 million to more than 1 billion, depending on how proficiency is defined. Linguist David Crystal estimates that non-native speakers now outnumber native speakers by a ratio of 3 to 1. In Kachru's three-circles model,

7076-453: The basis for the required controlled natural languages Seaspeak and Airspeak, used as international languages of seafaring and aviation. English used to have parity with French and German in scientific research, but now it dominates that field. It achieved parity with French as a language of diplomacy at the Treaty of Versailles negotiations in 1919. By the time of the foundation of

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7192-414: The basis of a computational classifier used to analyze real-world outbreaks. Computational predictions of transmission dynamics for each outbreak often align with known epidemiological data. Different transmission networks result in quantitatively different tree shapes. To determine whether tree shapes captured information about underlying disease transmission patterns, researchers simulated the evolution of

7308-400: The beginning, Englishmen had three manners of speaking, southern, northern and midlands speech in the middle of the country, ... Nevertheless, through intermingling and mixing, first with Danes and then with Normans, amongst many the country language has arisen, and some use strange stammering, chattering, snarling, and grating gnashing. John Trevisa , c.  1385 Middle English

7424-436: The branching pattern and "degree of difference" to find a compromise between them. Usual methods of phylogenetic inference involve computational approaches implementing the optimality criteria and methods of parsimony , maximum likelihood (ML), and MCMC -based Bayesian inference . All these depend upon an implicit or explicit mathematical model describing the evolution of characters observed. Phenetics , popular in

7540-460: The characteristics of species to interpret their evolutionary relationships and origins. Phylogenetics focuses on whether the characteristics of a species reinforce a phylogenetic inference that it diverged from the most recent common ancestor of a taxonomic group. In the field of cancer research, phylogenetics can be used to study the clonal evolution of tumors and molecular chronology , predicting and showing how cell populations vary throughout

7656-638: The consensus of educated English speakers around the world, without any oversight by any government or international organisation. American listeners readily understand most British broadcasting, and British listeners readily understand most American broadcasting. Most English speakers around the world can understand radio programmes, television programmes, and films from many parts of the English-speaking world. Both standard and non-standard varieties of English can include both formal or informal styles, distinguished by word choice and syntax and use both technical and non-technical registers. The settlement history of

7772-701: The continent. The Frisian languages, which together with the Anglic languages form the Anglo-Frisian languages , are the closest living relatives of English. Low German/Low Saxon is also closely related, and sometimes English, the Frisian languages, and Low German are grouped together as the North Sea Germanic languages, though this grouping remains debated. Old English evolved into Middle English , which in turn evolved into Modern English. Particular dialects of Old and Middle English also developed into

7888-400: The correctness of phylogenetic trees generated using fewer taxa and more sites per taxon on the x-axis to more taxa and fewer sites per taxon on the y-axis. With fewer taxa, more genes are sampled amongst the taxonomic group; in comparison, with more taxa added to the taxonomic sampling group, fewer genes are sampled. Each method has the same total number of nucleotide sites sampled. Furthermore,

8004-413: The data distribution. They may be used to quickly identify differences or similarities in the transmission data. Phylogenetic tools and representations (trees and networks) can also be applied to philology , the study of the evolution of oral languages and written text and manuscripts, such as in the field of quantitative comparative linguistics . Computational phylogenetics can be used to investigate

8120-476: The development of English was influenced by a long series of invasions of the British Isles by other peoples and languages, particularly Old Norse and French dialects . These left a profound mark of their own on the language, so that English shows some similarities in vocabulary and grammar with many languages outside its linguistic clades —but it is not mutually intelligible with any of those languages either. Some scholars have argued that English can be considered

8236-426: The direction of inferred evolutionary transformations. In addition to their use for inferring phylogenetic patterns among taxa, phylogenetic analyses are often employed to represent relationships among genes or individual organisms. Such uses have become central to understanding biodiversity , evolution, ecology , and genomes . Phylogenetics is a component of systematics that uses similarities and differences of

8352-597: The discovery of more genetic relationships in biodiverse fields, which can aid in conservation efforts by identifying rare species that could benefit ecosystems globally. Whole-genome sequence data from outbreaks or epidemics of infectious diseases can provide important insights into transmission dynamics and inform public health strategies. Traditionally, studies have combined genomic and epidemiological data to reconstruct transmission events. However, recent research has explored deducing transmission patterns solely from genomic data using phylodynamics , which involves analyzing

8468-476: The distinct characteristics of Early Modern English. In the 1611 King James Version of the Bible, written in Early Modern English, Matthew 8:20 says, "The Foxes haue holes and the birds of the ayre haue nests." This exemplifies the loss of case and its effects on sentence structure (replacement with subject–verb–object word order, and the use of of instead of the non-possessive genitive), and

8584-488: The dotted line represents a 1:1 accuracy between the two sampling methods. As seen in the graphic, most of the plotted points are located below the dotted line, which indicates gravitation toward increased accuracy when sampling fewer taxa with more sites per taxon. The research performed utilizes four different phylogenetic tree construction models to verify the theory; neighbor-joining (NJ), minimum evolution (ME), unweighted maximum parsimony (MP), and maximum likelihood (ML). In

8700-401: The early period of Old English were written using a runic script . By the 6th century, a Latin alphabet was adopted, written with half-uncial letterforms . It included the runic letters wynn ⟨ ƿ ⟩ and thorn ⟨ þ ⟩ , and the modified Latin letters eth ⟨ ð ⟩ , and ash ⟨ æ ⟩ . Old English is essentially

8816-551: The educational reforms of King Alfred in the 9th century and the influence of the kingdom of Wessex , the West Saxon dialect became the standard written variety . The epic poem Beowulf is written in West Saxon, and the earliest English poem, Cædmon's Hymn , is written in Northumbrian. Modern English developed mainly from Mercian, but the Scots language developed from Northumbrian. A few short inscriptions from

8932-695: The elite in England eventually developed into the Anglo-Norman language . Because Norman was spoken primarily by the elites and nobles, while the lower classes continued speaking English, the main influence of Norman was the introduction of a wide range of loanwords related to politics, legislation and prestigious social domains. Middle English also greatly simplified the inflectional system, probably in order to reconcile Old Norse and Old English, which were inflectionally different but morphologically similar. The distinction between nominative and accusative cases

9048-668: The emergence of biochemistry , organism classifications are now usually based on phylogenetic data, and many systematists contend that only monophyletic taxa should be recognized as named groups. The degree to which classification depends on inferred evolutionary history differs depending on the school of taxonomy: phenetics ignores phylogenetic speculation altogether, trying to represent the similarity between organisms instead; cladistics (phylogenetic systematics) tries to reflect phylogeny in its classifications by only recognizing groups based on shared, derived characters ( synapomorphies ); evolutionary taxonomy tries to take into account both

9164-408: The expanding circle use it to communicate with other people from the expanding circle, so that interaction with native speakers of English plays no part in their decision to use the language. Non-native varieties of English are widely used for international communication, and speakers of one such variety often encounter features of other varieties. Very often today a conversation in English anywhere in

9280-721: The growing economic and cultural influence of the United States and its status as a superpower following the Second World War has, along with worldwide broadcasting in English by the BBC and other broadcasters, caused the language to spread across the planet much faster. In the 21st century, English is more widely spoken and written than any language has ever been. As Modern English developed, explicit norms for standard usage were published, and spread through official media such as public education and state-sponsored publications. In 1755, Samuel Johnson published his A Dictionary of

9396-479: The history of English was Early Modern English (1500–1700). Early Modern English was characterised by the Great Vowel Shift (1350–1700), inflectional simplification, and linguistic standardisation. The Great Vowel Shift affected the stressed long vowels of Middle English. It was a chain shift , meaning that each shift triggered a subsequent shift in the vowel system. Mid and open vowels were raised , and close vowels were broken into diphthongs . For example,

9512-407: The history of how English spread in different countries, how users acquire English, and the range of uses English has in each country. The three circles change membership over time. Countries with large communities of native speakers of English (the inner circle) include Britain, the United States, Australia, Canada, Ireland, and New Zealand, where the majority speaks English, and South Africa, where

9628-403: The inner-circle countries, and they may show grammatical and phonological differences from inner-circle varieties as well. The standard English of the inner-circle countries is often taken as a norm for use of English in the outer-circle countries. In the three-circles model, countries such as Poland, China, Brazil, Germany, Japan, Indonesia, Egypt, and other countries where English is taught as

9744-531: The introduction of loanwords from French ( ayre ) and word replacements ( bird originally meaning "nestling" had replaced OE fugol ). By the late 18th century, the British Empire had spread English through its colonies and geopolitical dominance. Commerce, science and technology, diplomacy, art, and formal education all contributed to English becoming the first truly global language. English also facilitated worldwide international communication. English

9860-475: The introduction of the printing press to London. This era notably culminated in the King James Bible and the works of William Shakespeare . The printing press greatly standardised English spelling, which has remained largely unchanged since then, despite a wide variety of later sound shifts in English dialects. Modern English has spread around the world since the 17th century as a consequence of

9976-489: The island of Great Britain . The namesake of the language is the Angles , one of the ancient Germanic peoples that migrated to Britain . It is the most spoken language in the world, primarily due to the global influences of the former British Empire (succeeded by the Commonwealth of Nations ) and the United States . English is the third-most spoken native language , after Standard Chinese and Spanish ; it

10092-431: The languages have descended from a single common ancestor called Proto-Germanic . Some shared features of Germanic languages include the division of verbs into strong and weak classes, the use of modal verbs , and the sound changes affecting Proto-Indo-European consonants, known as Grimm's and Verner's laws . English is classified as an Anglo-Frisian language because Frisian and English share other features, such as

10208-475: The majority of models, sampling fewer taxon with more sites per taxon demonstrated higher accuracy. Generally, with the alignment of a relatively equal number of total nucleotide sites, sampling more genes per taxon has higher bootstrapping replicability than sampling more taxa. However, unbalanced datasets within genomic databases make increasing the gene comparison per taxon in uncommonly sampled organisms increasingly difficult. The term "phylogeny" derives from

10324-621: The mid-20th century but now largely obsolete, used distance matrix -based methods to construct trees based on overall similarity in morphology or similar observable traits (i.e. in the phenotype or the overall similarity of DNA , not the DNA sequence ), which was often assumed to approximate phylogenetic relationships. Prior to 1950, phylogenetic inferences were generally presented as narrative scenarios. Such methods are often ambiguous and lack explicit criteria for evaluating alternative hypotheses. In phylogenetic analysis, taxon sampling selects

10440-528: The phylogenetic history of a species cannot be read directly from its ontogeny, as Haeckel thought would be possible, but characters from ontogeny can be (and have been) used as data for phylogenetic analyses; the more closely related two species are, the more apomorphies their embryos share. One use of phylogenetic analysis involves the pharmacological examination of closely related groups of organisms. Advances in cladistics analysis through faster computer programs and improved molecular techniques have increased

10556-516: The precision of phylogenetic determination, allowing for the identification of species with pharmacological potential. Historically, phylogenetic screens for pharmacological purposes were used in a basic manner, such as studying the Apocynaceae family of plants, which includes alkaloid-producing species like Catharanthus , known for producing vincristine , an antileukemia drug. Modern techniques now enable researchers to study close relatives of

10672-426: The prestige varieties among the middle classes. In modern English, the loss of grammatical case is almost complete (it is now only found in pronouns, such as he and him , she and her , who and whom ), and SVO word order is mostly fixed. Some changes, such as the use of do-support , have become universalised. (Earlier English did not use the word "do" as a general auxiliary as Modern English does; at first it

10788-415: The progression of the disease and during treatment, using whole genome sequencing techniques. The evolutionary processes behind cancer progression are quite different from those in most species and are important to phylogenetic inference; these differences manifest in several areas: the types of aberrations that occur, the rates of mutation , the high heterogeneity (variability) of tumor cell subclones, and

10904-418: The properties of pathogen phylogenies. Phylodynamics uses theoretical models to compare predicted branch lengths with actual branch lengths in phylogenies to infer transmission patterns. Additionally, coalescent theory , which describes probability distributions on trees based on population size, has been adapted for epidemiological purposes. Another source of information within phylogenies that has been explored

11020-678: The reign of Henry V . Around 1430, the Court of Chancery in Westminster began using English in its official documents , and a new standard form of Middle English, known as Chancery Standard , developed from the dialects of London and the East Midlands . In 1476, William Caxton introduced the printing press to England and began publishing the first printed books in London, expanding the influence of this form of English. Literature from

11136-543: The relationship between two variables in pathogen transmission analysis, such as the number of infected individuals and the time since infection. These plots can help identify trends and patterns, such as whether the spread of the pathogen is increasing or decreasing over time, and can highlight potential transmission routes or super-spreader events. Box plots displaying the range, median, quartiles, and potential outliers datasets can also be valuable for analyzing pathogen transmission data, helping to identify important features in

11252-410: The relationships between several of the languages in a timeline, as well as the similarity between words and word order. There are three types of criticisms about using phylogenetics in philology, the first arguing that languages and species are different entities, therefore you can not use the same methods to study both. The second being how phylogenetic methods are being applied to linguistic data. And

11368-578: The relationships between viruses e.g., all viruses are descendants of Virus A. HIV forensics uses phylogenetic analysis to track the differences in HIV genes and determine the relatedness of two samples. Phylogenetic analysis has been used in criminal trials to exonerate or hold individuals. HIV forensics does have its limitations, i.e., it cannot be the sole proof of transmission between individuals and phylogenetic analysis which shows transmission relatedness does not indicate direction of transmission. Taxonomy

11484-485: The same useful traits. The phylogenetic tree shows which species of fish have an origin of venom, and related fish they may contain the trait. Using this approach in studying venomous fish, biologists are able to identify the fish species that may be venomous. Biologist have used this approach in many species such as snakes and lizards. In forensic science , phylogenetic tools are useful to assess DNA evidence for court cases. The simple phylogenetic tree of viruses A-E shows

11600-468: The sense of belonging only to people who are ethnically English . Use of English is growing country-by-country internally and for international communication. Most people learn English for practical rather than ideological reasons. Many speakers of English in Africa have become part of an "Afro-Saxon" language community that unites Africans from different countries. As decolonisation proceeded throughout

11716-484: The shape of phylogenetic trees, as illustrated in Fig. 1. Researchers have analyzed the structural characteristics of phylogenetic trees generated from simulated bacterial genome evolution across multiple types of contact networks. By examining simple topological properties of these trees, researchers can classify them into chain-like, homogeneous, or super-spreading dynamics, revealing transmission patterns. These properties form

11832-483: The source of approximately 28% of Modern English words , and from Latin , which is the source of an additional 28% . As such, though most of its total vocabulary comes from Romance languages , Modern English's grammar, phonology, and most commonly used words in everyday use keep it genealogically classified under the Germanic branch. It exists on a dialect continuum with Scots and is then most closely related to

11948-474: The southern dialects. Theoretically, as late as the 900s AD, a commoner from certain (northern) parts of England could hold a conversation with a commoner from certain parts of Scandinavia. Research continues into the details of the myriad tribes in peoples in England and Scandinavia and the mutual contacts between them. The translation of Matthew 8:20 from 1000 shows examples of case endings ( nominative plural, accusative plural, genitive singular) and

12064-455: The study of English as an auxiliary language. The trademarked Globish uses a relatively small subset of English vocabulary (about 1500 words, designed to represent the highest use in international business English) in combination with the standard English grammar. Other examples include Simple English . The increased use of the English language globally has had an effect on other languages, leading to some English words being assimilated into

12180-421: The third, discusses the types of data that is being used to construct the trees. Bayesian phylogenetic methods, which are sensitive to how treelike the data is, allow for the reconstruction of relationships among languages, locally and globally. The main two reasons for the use of Bayesian phylogenetics are that (1) diverse scenarios can be included in calculations and (2) the output is a sample of trees and not

12296-515: The vocabularies of other languages. This influence of English has led to concerns about language death , and to claims of linguistic imperialism , and has provoked resistance to the spread of English; however the number of speakers continues to increase because many people around the world think that English provides them with opportunities for better employment and improved lives. Phylogenetics In biology , phylogenetics ( / ˌ f aɪ l oʊ dʒ ə ˈ n ɛ t ɪ k s , - l ə -/ )

12412-473: The word bite was originally pronounced as the word beet is today, and the second vowel in the word about was pronounced as the word boot is today. The Great Vowel Shift explains many irregularities in spelling since English retains many spellings from Middle English, and it also explains why English vowel letters have very different pronunciations from the same letters in other languages. English began to rise in prestige, relative to Norman French, during

12528-785: The world may include no native speakers of English at all, even while including speakers from several different countries. This is particularly true of the shared vocabulary of mathematics and the sciences. English is a pluricentric language , which means that no one national authority sets the standard for use of the language. Spoken English, including English used in broadcasting, generally follows national pronunciation standards that are established by custom rather than by regulation. International broadcasters are usually identifiable as coming from one country rather than another through their accents , but newsreader scripts are also composed largely in international standard written English . The norms of standard written English are maintained purely by

12644-539: The world, but the number of English speakers in India is uncertain, with most scholars concluding that the United States still has more speakers of English than India. Modern English, sometimes described as the first global lingua franca , is also regarded as the first world language . English is the world's most widely used language in newspaper publishing, book publishing, international telecommunications, scientific publishing, international trade, mass entertainment, and diplomacy. English is, by international treaty,

12760-438: The worldwide influence of the British Empire and the United States. Through all types of printed and electronic media in these countries, English has become the leading language of international discourse and the lingua franca in many regions and professional contexts such as science, navigation , and law. Its modern grammar is the result of a gradual change from a dependent-marking pattern typical of Indo-European with

12876-563: The years they developed a unique mixed language called Michif by combining Métis French nouns, numerals, articles and adjectives with Cree verbs, demonstratives , postpositions , interrogatives and pronouns. Both the Michif language and the Métis dialect of French are severely endangered . Newfoundland French is spoken by a small population on the Port au Port Peninsula of Newfoundland. It

12992-405: Was adopted in parts of North America, parts of Africa, Oceania, and many other regions. When they obtained political independence, some of the newly independent states that had multiple indigenous languages opted to continue using English as the official language to avoid the political and other difficulties inherent in promoting any one indigenous language above the others. In the 20th century

13108-443: Was formerly used to refer specifically to Quebec French and the closely related varieties of Ontario and Western Canada descended from it. This is presumably because Canada and Acadia were distinct parts of New France , and also of British North America , until 1867. The term is no longer usually deemed to exclude Acadian French. Phylogenetically , Quebec French, Métis French and Brayon French are representatives of koiné French in

13224-518: Was fully developed, integrating both Norse and French features; it continued to be spoken until the transition to early Modern English around 1500. Middle English literature includes Geoffrey Chaucer 's The Canterbury Tales , and Thomas Malory 's Le Morte d'Arthur . In the Middle English period, the use of regional dialects in writing proliferated, and dialect traits were even used for effect by authors such as Chaucer. The next period in

13340-505: Was lost except in personal pronouns, the instrumental case was dropped, and the use of the genitive case was limited to indicating possession . The inflectional system regularised many irregular inflectional forms, and gradually simplified the system of agreement, making word order less flexible. The transition from Old to Middle English can be placed during the writing of the Ormulum . The oldest Middle English texts that were written by

13456-533: Was only used in question constructions, and even then was not obligatory. Now, do-support with the verb have is becoming increasingly standardised.) The use of progressive forms in -ing , appears to be spreading to new constructions, and forms such as had been being built are becoming more common. Regularisation of irregular forms also slowly continues (e.g. dreamed instead of dreamt ), and analytical alternatives to inflectional forms are becoming more common (e.g. more polite instead of politer ). British English

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