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43-527: 130, see text Aquilina Bubani, nom. illeg. Aquilegia (common names: granny's bonnet , columbine ) is a genus of about 130 species of perennial plants that are found in meadows , woodlands , and at higher elevations throughout the Northern Hemisphere, known for the spurred petals of their flowers . The genus name Aquilegia comes from the Latin "Aquila", or "eagle"; this

86-610: A geometer moth , also uses columbine as a larval food plant. The larvae of the Papaipema leucostigma also feed on columbine. Plants in the genus Aquilegia are a major food source for Bombus hortorum , a species of bumblebee. Specifically, they have been found to forage on species of Aquilegia vulgaris in Belgium and Aquilegia chrysantha in North America and Belgium. The bees do not show any preference in color of

129-557: A species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in the nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

172-459: A flat limb and spurred or saccate at the base. The spur is directed backwards and secretes nectar . Stamens are numerous (often more than 50) in whorls of 5, the innermost being scarious staminodes . There are ten membranaceous intrastaminal scales. There are five pistils and the carpels are free. The fruit has several (five to 15) follicles which are semi erect and slightly connate downwards. These hold many seeds and are formed at

215-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

258-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

301-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

344-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

387-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

430-460: A treatment for ulcers . However, the medical use of this plant is better avoided due to its high toxicity; columbine poisonings may be fatal. An acute toxicity test in mice has demonstrated that ethanol extract mixed with isocytisoside , the main flavonoid compound from the leaves and stems of Aquilegia vulgaris , can be classified as non-toxic, since a dose of 3000 mg/kg did not cause mortality. The Colorado blue columbine ( A. coerulea )

473-483: A variety of single colours and bi-colours, in single and double forms. Though perennial, cultivars may be short-lived and thus best treated as biennials . Spent flower-heads should be removed to prevent the plant going to seed. Cultivars include the Barlow series ('Nora Barlow', 'Black Barlow', 'Rose Barlow', 'Christa Barlow'), 'Pretty Bonnets'. Seeds may be sold as mixtures. The white flowering cultivar 'Nivea' has gained

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516-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

559-620: Is in obvious reference to the spurred, "hook" shapes within the blooms, that many gardeners say resemble an eagle's talons. Perennial herbs , with woody, erect stock, roots forming thick rhizomes . The basal leaves are compound, 1–3 ternate , blades 3-lobed -partite, and lobes lobulate and obtuse . The cauline leaves are similar to the basal ones, while the upper ones are bract like. The hermaphrodite (bisexual) flowers are terminal to stem and branches. They are usually pentamerous (with five spreading perianth petaloid sepal segments). Five tubular honey-leaves are semi erect with

602-526: Is native to Albania, Austria, Baltic States, Belarus, Belgium, Bulgaria, Central European Russia, Corse, Czechoslovakia, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Netherlands, Northwest European Russia, Poland, Portugal, Romania, Sicilia, Spain, Switzerland, Yugoslavia. It has been introduced to many other areas including parts of North and South America, and other parts of Europe and north eastern Asia. This species and various hybrids derived from it are popular garden flowers, available in

645-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

688-624: Is suggested to have driven these changes in nectar spur length. It was shown that this spur length diversity is achieved solely through changing cell shape, not cell number or cell size. This suggests that a simple microscopic change can result in a dramatic evolutionarily relevant morphological change. 130 columbine species are accepted. Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including

731-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

774-525: Is the official state flower of Colorado (see also Columbine, Colorado ). It is also used as a symbol of the former city of Scarborough in the Canadian province of Ontario. Columbines have been important in the study of evolution . It was found that the Sierra columbine ( A. pubescens ) and crimson columbine ( A. formosa ) each has adapted specifically to a pollinator . Bees and hummingbirds are

817-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

860-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

903-527: The Ranunculaceae , all parts of the plant contain toxic alkaloids and are poisonous raw, although the flowers are edible in small quantities. The seeds and root are particularly toxic. The leaves may be detoxified by thorough boiling in multiple changes of water. The acute toxicity test in mice showed that ethanol extract and the main flavonoid compound isocytisoside from the leaves and stems of Aquilegia vulgaris can be classified as nontoxic since

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946-533: The Royal Horticultural Society 's Award of Garden Merit . Aphids and sawfly larvae may attack the plant. In traditional herbalism columbine was considered sacred to Venus ; carrying a posy of it was said to arouse the affections of a loved one. Nicholas Culpeper recommended the seeds taken in wine to speed the process of childbirth. In modern herbal medicine it is used as an astringent and diuretic . Like many other members of

989-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

1032-473: The French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or the scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of

1075-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

1118-723: The collection started to succumb to Aquilegia Downy Mildew Peronospora aquilegiicola which was at the time an emerging disease to which the plants had no resistance. By 2018 the entire collection had been lost. Aquilegia can be grown from seeds or rhizomes. The flowers of various species of columbine were consumed in moderation by Native Americans as a condiment with other fresh greens, and are reported to be very sweet, and safe if consumed in small quantities. The plant's seeds and roots, however, are highly poisonous and contain cardiogenic toxins which cause both severe gastroenteritis and heart palpitations if consumed as food. Native Americans used very small amounts of Aquilegia root as

1161-800: The end of the pistils. The nectar is mainly consumed by long-beaked birds such as hummingbirds. Almost all Aquilegia species have a ring of staminodia around the base of the stigma, which may help protect against insects. Chromosome number is x=7. Columbines are closely related to plants in the genera Actaea (baneberries) and Aconitum (wolfsbanes/monkshoods), which like Aquilegia produce cardiogenic toxins . They are used as food plants by some Lepidoptera (butterfly and moth) caterpillars . These are mainly of noctuid moths – noted for feeding on many poisonous plants without harm – such as cabbage moth ( Mamestra brassicae ), dot moth ( Melanchra persicariae ) and mouse moth ( Amphipyra tragopoginis ). The engrailed ( Ectropis crepuscularia ),

1204-465: The flowers. Columbine is a hardy perennial , which propagates by seed. It will grow to a height of 40–50 cm (15–20 in). It will grow in full sun; however, it prefers growing in partial shade and well drained soil, and is able to tolerate average soils and dry soil conditions. Columbine is rated at hardiness zone 3 in the United States so does not require mulching or protection in

1247-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

1290-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

1333-432: The idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of the same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but

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1376-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

1419-790: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Aquilegia vulgaris Aquilegia vulgaris is a species of columbine native to Europe with common names that include: European columbine , common columbine , granny's nightcap , and granny's bonnet . It is a flowering herbaceous perennial plant growing to 1.2 m tall, with branched, thinly hairy stems. The leaves are biternate; each leaf has three groups of three leaflets. The flowers, in various shades of purple, blue, pink and white, are pendent or horizontal with strongly hooked spurs, and appear in early summer. The Latin specific epithet vulgaris means "common". The species

1462-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

1505-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

1548-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

1591-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

1634-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

1677-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

1720-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

1763-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

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1806-462: The visitors to A. formosa , while hawkmoths would only visit A. pubescens when given a choice. Such a " pollination syndrome ", being due to flower color and orientation controlled by their genetics , ensures reproductive isolation and can be a cause of speciation . Aquilegia petals show an enormous range of petal spur length diversity ranging from a centimeter to the 15 cm spurs of Aquilegia longissima . Selection from pollinator shifts

1849-496: The winter. Large numbers of hybrids are available for the garden, since the European A. vulgaris was hybridized with other European and North American varieties. Aquilegia species are very interfertile, and will self-sow. Some varieties are short-lived so are better treated as biennials . The British National Collection of Aquilegia s was held by Mrs Carrie Thomas at Killay near Swansea . Some time during or before 2014

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