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138-400: Copenhagen Zoo was founded by the ornithologist Niels Kjærbølling in 1859. He was given the summer garden of "Prinsess Vilhelmines Have" (The garden of Princess Vilhelmine ) by the chief directorate of Copenhagen . The animals that the visitors could contemplate at the opening were eagles , chickens , ducks , owls , rabbits , a fox , a seal in a bathtub and a turtle in a bucket. In

276-630: A binomial name , categorising them into different genera. However, ornithology did not emerge as a specialised science until the Victorian era—with the popularization of natural history, and the collection of natural objects such as bird eggs and skins. This specialization led to the formation in Britain of the British Ornithologists' Union in 1858. In 1859, the members founded its journal The Ibis . The sudden spurt in ornithology

414-635: A menagerie and sponsored translations of Arabic texts, among which the popular Arabic work known as the Liber Moaminus by an unknown author which was translated into Latin by Theodore of Antioch from Syria in 1240-1241 as the De Scientia Venandi per Aves , and also Michael Scotus (who had removed to Palermo) translated Ibn Sīnā 's Kitāb al-Ḥayawān of 1027 for the Emperor, a commentary and scientific update of Aristotle's work which

552-429: A "difficult language to learn, acquire and understand", and some evidence shows that children are slower to acquire the phonological distinctions of Danish compared with other languages. The grammar is moderately inflective with strong (irregular) and weak (regular) conjugations and inflections. Nouns, adjectives, and demonstrative pronouns distinguish common and neutral gender. Like English, Danish only has remnants of

690-484: A 3-metre (9.8 ft) high wall, has been opened up so that people in the park can watch the elephants. Most of the "Africa" ( Danish : Afrika ) section is taken up by the "Savanna" ( Danish : Savannen ) area. This is an open field with a visitor bridge above containing species such as white rhinoceroses , giraffes , impalas , sable antelopes , ostriches and plains zebras . The hippopotamuses , which have their own separate enclosure, also have partial access to

828-453: A Hippopotamus House where the hippos can be watched underwater. The zoo has preserved many of its historical buildings. The oldest building still in use, a stable for yaks , was erected in 1872, and now houses the bactrian camels . A Herbivore House built in 1875 still houses herbivores, namely tapirs . An owl tower from 1885 is today left as a memorial commemorating how zoo animals were once kept. A notable and highly visible feature of

966-619: A Latin translation of Aristotle's work on animals from Arabic here around 1215, which was disseminated widely and was the first time in a millennium that this foundational text on zoology became available to Europeans. Falconry was popular in the Norman court in Sicily, and a number of works on the subject were written in Palermo . Emperor Frederick II of Hohenstaufen (1194–1250) learned about an falconry during his youth in Sicily and later built up

1104-697: A classification of birds, De Differentiis Avium (around 1572), that was based on structure and habits. Konrad Gesner wrote the Vogelbuch and Icones avium omnium around 1557. Like Gesner, Ulisse Aldrovandi , an encyclopedic naturalist, began a 14-volume natural history with three volumes on birds, entitled ornithologiae hoc est de avibus historiae libri XII , which was published from 1599 to 1603. Aldrovandi showed great interest in plants and animals, and his work included 3000 drawings of fruits, flowers, plants, and animals, published in 363 volumes. His Ornithology alone covers 2000 pages and included such aspects as

1242-577: A former case system , particularly in the pronouns. Unlike English, it has lost all person marking on verbs. Its word order is V2 , with the finite verb always occupying the second slot in the sentence. Danish is a Germanic language of the North Germanic branch . Other names for this group are the Nordic or Scandinavian languages. Along with Swedish, Danish descends from the Eastern dialects of

1380-417: A further giraffe into its bachelor herd in its state-of-the-art giraffe house built in 2012, and that contact had been made with Copenhagen Zoo but no reply had been received. YWP has previously received a male giraffe of the same age as Marius from Copenhagen Zoo. Bengt Holst stated that the capacity at YWP would be better utilised by a giraffe whose genes were less represented amongst EAZA zoos. The action

1518-467: A huge understanding of the anatomy of a giraffe that they wouldn't have had from watching a giraffe in a photo," Stenbaek Bro said in a telephone interview with the Associated Press. On 25 March 2014, despite the backlash from the giraffe cull, the zoo euthanized four lions (two adults and two ten-month-old cubs). The zoo stated that they were introducing a male lion and did not want to disrupt

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1656-402: A large contribution made by amateurs in terms of time, resources, and financial support. Studies on birds have helped develop key concepts in biology including evolution, behaviour and ecology such as the definition of species , the process of speciation , instinct , learning , ecological niches , guilds , island biogeography , phylogeography , and conservation . While early ornithology

1794-526: A literary language. Also in this period, Danish began to take on the linguistic traits that differentiate it from Swedish and Norwegian, such as the stød , the voicing of many stop consonants, and the weakening of many final vowels to /e/. The first printed book in Danish dates from 1495, the Rimkrøniken ( Rhyming Chronicle ), a history book told in rhymed verses. The first complete translation of

1932-448: A model for many studies in non-mammalian immunology. Studies in bird behaviour include the use of tamed and trained birds in captivity. Studies on bird intelligence and song learning have been largely laboratory-based. Field researchers may make use of a wide range of techniques such as the use of dummy owls to elicit mobbing behaviour, and dummy males or the use of call playback to elicit territorial behaviour and thereby to establish

2070-707: A number of Danes remained as a minority within German territories . After the occupation of Denmark by Germany in World War II, the 1948 orthography reform dropped the German-influenced rule of capitalizing nouns, and introduced the letter ⟨å⟩ . Three 20th-century Danish authors have become Nobel Prize laureates in Literature : Karl Gjellerup and Henrik Pontoppidan (joint recipients in 1917) and Johannes V. Jensen (awarded 1944). With

2208-593: A policy of the British Ornithologists' Union to keep out women. Unlike the RSPB, which was primarily conservation oriented, the British Trust for Ornithology was started in 1933 with the aim of advancing ornithological research. Members were often involved in collaborative ornithological projects. These projects have resulted in atlases which detail the distribution of bird species across Britain. In Canada, citizen scientist Elsie Cassels studied migratory birds and

2346-586: A resource for taxonomic studies. The use of bird skins to document species has been a standard part of systematic ornithology. Bird skins are prepared by retaining the key bones of the wings, legs, and skull along with the skin and feathers. In the past, they were treated with arsenic to prevent fungal and insect (mostly dermestid ) attack. Arsenic, being toxic, was replaced by less-toxic borax . Amateur and professional collectors became familiar with these skinning techniques and started sending in their skins to museums, some of them from distant locations. This led to

2484-482: A similarity in pronunciation, combined with the long tradition of having Danish as a written language, which has led to similarities in vocabulary. Among younger Danes, Copenhageners are worse at understanding Swedish than Danes from the provinces. In general, younger Danes are not as good at understanding the neighboring languages as the young in Norway and Sweden. The Danish philologist Johannes Brøndum-Nielsen divided

2622-586: A so-called multiethnolect in the urban areas, an immigrant Danish variety (also known as Perkerdansk ), combining elements of different immigrant languages such as Arabic, Turkish, and Kurdish, as well as English and Danish. Within the Danish Realm , Danish is the national language of Denmark and one of two official languages of the Faroe Islands (alongside Faroese ). There is a Faroese variant of Danish known as Gøtudanskt . Until 2009, Danish

2760-403: A spurt of bird studies in this area. The study of imprinting behaviour in ducks and geese by Konrad Lorenz and the studies of instinct in herring gulls by Nicolaas Tinbergen led to the establishment of the field of ethology . The study of learning became an area of interest and the study of bird songs has been a model for studies in neuroethology. The study of hormones and physiology in

2898-943: A tract on falconry, and introduced the words "incubation" and "oviparous" into the English language. Towards the late 18th century, Mathurin Jacques Brisson (1723–1806) and Comte de Buffon (1707–1788) began new works on birds. Brisson produced a six-volume work Ornithologie in 1760 and Buffon's included nine volumes (volumes 16–24) on birds Histoire naturelle des oiseaux (1770–1785) in his work on science Histoire naturelle générale et particulière (1749–1804). Jacob Temminck sponsored François Le Vaillant [1753–1824] to collect bird specimens in Southern Africa and Le Vaillant's six-volume Histoire naturelle des oiseaux d'Afrique (1796–1808) included many non-African birds. His other bird books produced in collaboration with

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3036-462: A transparent top and visible cues such as the direction of sunlight may be controlled using mirrors or the positions of the stars simulated in a planetarium . The entire genome of the domestic fowl ( Gallus gallus ) was sequenced in 2004, and was followed in 2008 by the genome of the zebra finch ( Taeniopygia guttata ). Such whole-genome sequencing projects allow for studies on evolutionary processes involved in speciation . Associations between

3174-462: A two-part article in The Auk , noting the tensions between amateurs and professionals, and suggested the possibility that the "vast army of bird lovers and bird watchers could begin providing the data scientists needed to address the fundamental problems of biology." The amateur ornithologist Harold F. Mayfield noted that the field was also funded by non-professionals. He noted that in 1975, 12% of

3312-625: A variant of Standard Danish, Southern Schleswig Danish , is spoken in the area. Since 2015, Schleswig-Holstein has officially recognized Danish as a regional language , just as German is north of the border. Furthermore, Danish is one of the official languages of the European Union and one of the working languages of the Nordic Council . Under the Nordic Language Convention , Danish-speaking citizens of

3450-407: A very large vowel inventory consisting of 27 phonemically distinctive vowels , and its prosody is characterized by the distinctive phenomenon stød , a kind of laryngeal phonation type . Due to the many pronunciation differences that set Danish apart from its neighboring languages, particularly the vowels, difficult prosody and "weakly" pronounced consonants, it is sometimes considered to be

3588-571: Is a North Germanic language from the Indo-European language family spoken by about six million people, principally in and around Denmark . Communities of Danish speakers are also found in Greenland , the Faroe Islands , and the northern German region of Southern Schleswig , where it has minority language status. Minor Danish-speaking communities are also found in Norway , Sweden ,

3726-700: Is a West Germanic language descended from Old English. Old Norse exerted a strong influence on Old English in the early medieval period. The shared Germanic heritage of Danish and English is demonstrated with many common words that are very similar in the two languages. For example, when written, commonly used Danish verbs, nouns, and prepositions such as have , over , under , for , give , flag , salt , and arm are easily recognizable to English speakers. Similarly, some other words are almost identical to their Scots equivalents, e.g. kirke (Scots kirk , i.e., 'church') or barn (Scots and northern English bairn , i.e. 'child'). In addition,

3864-536: Is also used as location in episode 84 of the DR television show Huset på Christianshavn and in the TV2 television Christmas calendar Brødrenes Mortensens jul . Ornithologist Ornithology is a branch of zoology that concerns the study of birds . Several aspects of ornithology differ from related disciplines, due partly to the high visibility and the aesthetic appeal of birds. It has also been an area with

4002-467: Is another technique that has the advantage of preserving stomach contents and anatomy, although it tends to shrink, making it less reliable for morphometrics. The study of birds in the field was helped enormously by improvements in optics. Photography made it possible to document birds in the field with great accuracy. High-power spotting scopes today allow observers to detect minute morphological differences that were earlier possible only by examination of

4140-447: Is commonplace knowledge to the mere collector, such as that hunting parties often travel more or less in circles. David Lack's studies on population ecology sought to find the processes involved in the regulation of population based on the evolution of optimal clutch sizes. He concluded that population was regulated primarily by density-dependent controls , and also suggested that natural selection produces life-history traits that maximize

4278-620: Is extinct and all extant ostrich races are today restricted to Africa . Other old writings such as the Vedas (1500–800 BC) demonstrate the careful observation of avian life histories and include the earliest reference to the habit of brood parasitism by the Asian koel ( Eudynamys scolopaceus ). Like writing, the early art of China, Japan, Persia, and India also demonstrate knowledge, with examples of scientifically accurate bird illustrations. Aristotle in 350 BC in his History of animals noted

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4416-510: Is for cows and calves and measures 45 by 23 metres (148 by 75 ft). The other is 30 by 15 metres (98 by 49 ft) and is for bulls, kept in separate pens during the mating season for fear of fights. The building also contain an exhibit space and a small lecture hall. The enclosures open out through mighty rusted steel doors into am almost 1 hectare big landscaped paddock with a pool 3 metres (9.8 ft) deep and 60 metres (200 ft) long. The paddock's border with Frederiksberg Gardens, once

4554-430: Is its closest relative. East Germanic languages West Germanic languages Icelandic Faroese Norwegian Danish Swedish Approximately 2,000 uncompounded Danish words are derived from Old Norse and ultimately from Proto Indo-European . Of these 2,000, 1,200 are nouns, 500 are verbs and 180 are adjectives. Danish has also absorbed many loanwords , most of which were borrowed from Low German of

4692-554: Is often regarded as the greatest ornithological work in history. The emergence of ornithology as a scientific discipline began in the 18th century, when Mark Catesby published his two-volume Natural History of Carolina, Florida, and the Bahama Islands , a landmark work which included 220 hand-painted engravings and was the basis for many of the species Carl Linnaeus described in the 1758 Systema Naturae . Linnaeus' work revolutionised bird taxonomy by assigning every species

4830-469: Is the best option". Images of the carcass being cut up in front of children and then fed to the zoo's lion population were circulated by the Associated Press . The zoo spokesperson, Tobias Stenbaek Bro, has been quoted as saying that visitors, including children, were invited to watch as the giraffe was skinned and parts fed to the lions. "I'm actually proud because I think we have given children

4968-584: Is the first such record of North American birds, significantly antedating Audubon. In the early 19th century, Lewis and Clark studied and identified many birds in the western United States. John James Audubon , born in 1785, observed and painted birds in France and later in the Ohio and Mississippi valleys. From 1827 to 1838, Audubon published The Birds of America , which was engraved by Robert Havell Sr. and his son Robert Havell Jr. Containing 435 engravings, it

5106-436: The skarre-R , the uvular R sound ( [ʁ] ), began spreading through Denmark, likely through influence from Parisian French and German. It affected all of the areas where Danish had been influential, including all of Denmark, Southern Sweden, and coastal southern Norway. In the 18th century, Danish philology was advanced by Rasmus Rask , who pioneered the disciplines of comparative and historical linguistics, and wrote

5244-623: The Bible in Danish, the Bible of Christian II translated by Christiern Pedersen , was published in 1550. Pedersen's orthographic choices set the de facto standard for subsequent writing in Danish. From around 1500, several printing presses were in operation in Denmark publishing in Danish and other languages. In the period after 1550, presses in Copenhagen dominated the publication of material in

5382-573: The Bulo Burti boubou ( Laniarius liberatus , no longer a valid species) and the Bugun liocichla ( Liocichla bugunorum ), using blood, DNA and feather samples as the holotype material, has now become possible. Other methods of preservation include the storage of specimens in spirit. Such wet specimens have special value in physiological and anatomical study, apart from providing better quality of DNA for molecular studies. Freeze drying of specimens

5520-570: The Late Middle Ages . Out of the 500 most frequently used Danish words, 100 are loans from Middle Low German; this is because Low German was the second official language of Denmark–Norway. In the 17th and 18th centuries, standard German and French superseded Low German influence, and in the 20th century, English became the main supplier of loanwords, especially after World War II . Although many old Nordic words remain, some were replaced with borrowed synonyms, for example æde (to eat)

5658-523: The Old Norse language ; Danish and Swedish are also classified as East Scandinavian or East Nordic languages. Scandinavian languages are often considered a dialect continuum , where no sharp dividing lines are seen between the different vernacular languages. Like Norwegian and Swedish, Danish was significantly influenced by Low German in the Middle Ages, and has been influenced by English since

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5796-1091: The United States , Canada , Brazil , and Argentina . Along with the other North Germanic languages, Danish is a descendant of Old Norse , the common language of the Germanic peoples who lived in Scandinavia during the Viking Era . Danish, together with Swedish, derives from the East Norse dialect group , while the Middle Norwegian language (before the influence of Danish) and Norwegian Bokmål are classified as West Norse along with Faroese and Icelandic . A more recent classification based on mutual intelligibility separates modern spoken Danish, Norwegian , and Swedish as "mainland (or continental ) Scandinavian", while Icelandic and Faroese are classified as "insular Scandinavian". Although

5934-544: The Zealand dialect Introductio ad lingvam Danicam puta selandicam ; and in 1685 the first Danish grammar written in Danish, Den Danske Sprog-Kunst ("The Art of the Danish Language") by Peder Syv . Major authors from this period are Thomas Kingo , poet and psalmist, and Leonora Christina Ulfeldt , whose novel Jammersminde ( Remembered Woes ) is considered a literary masterpiece by scholars. Orthography

6072-467: The chicken and poultry techniques. He used a number of traits including behaviour, particularly bathing and dusting, to classify bird groups. William Turner 's Historia Avium ( History of Birds ), published at Cologne in 1544, was an early ornithological work from England. He noted the commonness of kites in English cities where they snatched food out of the hands of children. He included folk beliefs such as those of anglers. Anglers believed that

6210-402: The introduction of the printing press , a standard language was developed which was based on the educated dialect of Copenhagen and Malmö . It spread through use in the education system and administration, though German and Latin continued to be the most important written languages well into the 17th century. Following the loss of territory to Germany and Sweden, a nationalist movement adopted

6348-421: The osprey emptied their fishponds and would kill them, mixing the flesh of the osprey into their fish bait. Turner's work reflected the violent times in which he lived, and stands in contrast to later works such as Gilbert White 's 1789 The Natural History and Antiquities of Selborne that were written in a tranquil era. In the 17th century, Francis Willughby (1635–1672) and John Ray (1627–1705) created

6486-469: The "Asia" ( Danish : Asien ) section, visitors can see Oriental small-clawed otters , red pandas , Malayan tapirs , Amur leopards and tigers , Asian elephants , and other animals. The new Elephant House, which opened in June 2008, is designed by Norman Foster in cooperation with the Danish landscape architect Stig L. Andersson . It houses Asian elephants, and contains two glass-domed enclosures. One

6624-506: The 1820s and 1830s, with pioneers such as J. Dovaston (who also pioneered in the use of bird feeders), but instruction manuals did not begin to insist on the use of optical aids such as "a first-class telescope" or "field glass" until the 1880s. The rise of field guides for the identification of birds was another major innovation. The early guides such as Thomas Bewick's two-volume guide and William Yarrell's three-volume guide were cumbersome, and mainly focused on identifying specimens in

6762-454: The 19th century. In 1901, Robert Ridgway wrote in the introduction to The Birds of North and Middle America that: There are two essentially different kinds of ornithology: systematic or scientific, and popular. The former deals with the structure and classification of birds, their synonymies, and technical descriptions. The latter treats of their habits, songs, nesting, and other facts pertaining to their life histories. This early idea that

6900-579: The 9th century with the younger futhark . Possibly as far back as the seventh century, the common Norse language began to undergo changes that did not spread to all of Scandinavia, resulting in the appearance of two dialect areas, Old West Norse ( Norway and Iceland ) and Old East Norse ( Denmark and Sweden ). Most of the changes separating East Norse from West Norse started as innovations in Denmark, that spread through Scania into Sweden and by maritime contact to southern Norway. A change that separated Old East Norse (Runic Swedish/Danish) from Old West Norse

7038-537: The Birds by Roger Tory Peterson in 1934, to Birds of the West Indies published in 1936 by Dr. James Bond - the same who inspired the amateur ornithologist Ian Fleming in naming his famous literary spy . The interest in birdwatching grew in popularity in many parts of the world, and the possibility for amateurs to contribute to biological studies was soon realized. As early as 1916, Julian Huxley wrote

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7176-594: The Danish language, and also started a period of homogenization, whereby the Copenhagen standard language gradually displaced the regional vernacular languages. Throughout the 19th century, Danes emigrated, establishing small expatriate communities in the Americas, particularly in the United States, Canada, and Argentina, where memory and some use of Danish remains today. After the Schleswig referendum in 1920 ,

7314-517: The Danish language. Herrer og Narre have frit Sprog . "Lords and jesters have free speech." Peder Syv , proverbs Following the first Bible translation, the development of Danish as a written language , as a language of religion, administration, and public discourse accelerated. In the second half of the 17th century, grammarians elaborated grammars of Danish, first among them Rasmus Bartholin 's 1657 Latin grammar De studio lingvæ danicæ ; then Laurids Olufsen Kock 's 1660 grammar of

7452-407: The Danish tongue." Heimskringla by Snorri Sturluson By the eighth century, the common Germanic language of Scandinavia, Proto-Norse , had undergone some changes and evolved into Old Norse . This language was generally called the "Danish tongue" ( Dǫnsk tunga ), or "Norse language" ( Norrœnt mál ). Norse was written in the runic alphabet , first with the elder futhark and from

7590-609: The Elder described birds, among other creatures, in his Historia Naturalis . The earliest record of falconry comes from the reign of Sargon II (722–705 BC) in Assyria . Falconry is thought to have made its entry to Europe only after AD 400, brought in from the east after invasions by the Huns and Alans . Starting from the eighth century, numerous Arabic works on the subject and general ornithology were written, as well as translations of

7728-544: The Mediterranean, and Pierre Belon , who described the fish and birds that he had seen in France and the Levant. Belon's Book of Birds (1555) is a folio volume with descriptions of some 200 species. His comparison of the skeleton of humans and birds is considered as a landmark in comparative anatomy . Volcher Coiter (1534–1576), a Dutch anatomist, made detailed studies of the internal structures of birds and produced

7866-710: The Old Norse word for "island". This monophthongization started in Jutland and spread eastward, having spread throughout Denmark and most of Sweden by 1100. Through Danish conquest, Old East Norse was once widely spoken in the northeast counties of England . Many words derived from Norse, such as "gate" ( gade ) for street, still survive in Yorkshire , the East Midlands and East Anglia, and parts of eastern England colonized by Danish Vikings . The city of York

8004-644: The Savanna. Other species in the Africa section are okapis , Abyssinian ground hornbills , Congo peafowl , caracals and meerkats . The "Tasmania" ( Danish : Tasmanien ) section is unique to Copenhagen Zoo. The section was started after the wedding of Frederik, the Crown Prince of Denmark and the Tasmanian native Mary Donaldson in 2004, where the Tasmanian government gifted four Tasmanian Devils to

8142-685: The South American pampas. The "Tropical Zoo" ( Danish : Tropezoo ) consists of a rainforest hall with one section for free-ranging birds, turtles and two-toed sloths and another for tropical butterflies and West African crocodiles . In between the two is a section for reptiles, amphibians, fish and the rare Java mouse-deer . In the Children's Zoo ( Danish : Børnezoo ), visitors can see farm animals and small domestic animals, such as llamas , goats , cows , pigs , horses , chickens and rabbits . A healthy young male giraffe, Marius ,

8280-486: The Victorian era observed the variations in bird forms and habits across geographic regions, noting local specialization and variation in widespread species. The collections of museums and private collectors grew with contributions from various parts of the world. The naming of species with binomials and the organization of birds into groups based on their similarities became the main work of museum specialists. The variations in widespread birds across geographical regions caused

8418-600: The area, eventually outnumbering the Danish speakers. The political loss of territory sparked a period of intense nationalism in Denmark, coinciding with the so-called " Golden Age " of Danish culture. Authors such as N.F.S. Grundtvig emphasized the role of language in creating national belonging. Some of the most cherished Danish-language authors of this period are existential philosopher Søren Kierkegaard and prolific fairy tale author Hans Christian Andersen . The influence of popular literary role models, together with increased requirements of education did much to strengthen

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8556-497: The artist Barraband are considered among the most valuable illustrated guides ever produced. Louis Pierre Vieillot (1748–1831) spent 10 years studying North American birds and wrote the Histoire naturelle des oiseaux de l'Amerique septentrionale (1807–1808?). Vieillot pioneered in the use of life histories and habits in classification. Alexander Wilson composed a nine-volume work, American Ornithology , published 1808-1814, which

8694-440: The behaviour, with many names being onomatopoeic , and still in use. Traditional knowledge may also involve the use of birds in folk medicine and knowledge of these practices are passed on through oral traditions (see ethno-ornithology ). Hunting of wild birds as well as their domestication would have required considerable knowledge of their habits. Poultry farming and falconry were practised from early times in many parts of

8832-530: The best genes were passed down to ensure the animals' long-term survival. He confirmed the zoo typically culls 20 to 30 animals every year. The death occurred despite offers from other animal institutions (EAZA member Yorkshire Wildlife Park (YWP) in the United Kingdom and Hoenderdael Park in The Netherlands) to take the giraffe. YWP released a statement that it had the capacity to accept

8970-431: The boundaries of bird territories. Studies of bird migration including aspects of navigation, orientation, and physiology are often studied using captive birds in special cages that record their activities. The Emlen funnel , for instance, makes use of a cage with an inkpad at the centre and a conical floor where the ink marks can be counted to identify the direction in which the bird attempts to fly. The funnel can have

9108-427: The categories of those that are applicable to specimens and those that are used in the field, but the classification is rough and many analysis techniques are usable both in the laboratory and field or may require a combination of field and laboratory techniques. The earliest approaches to modern bird study involved the collection of eggs, a practice known as oology . While collecting became a pastime for many amateurs,

9246-528: The control of behaviour has also been aided by bird models. These have helped in finding the proximate causes of circadian and seasonal cycles. Studies on migration have attempted to answer questions on the evolution of migration, orientation, and navigation. The growth of genetics and the rise of molecular biology led to the application of the gene-centered view of evolution to explain avian phenomena. Studies on kinship and altruism, such as helpers , became of particular interest. The idea of inclusive fitness

9384-561: The cooperation of people and organizations in different countries. Wild birds impact many human activities, while domesticated birds are important sources of eggs, meat, feathers, and other products. Applied and economic ornithology aim to reduce the ill effects of problem birds and enhance gains from beneficial species. Danish language Nordic Council Danish ( / ˈ d eɪ n ɪ ʃ / , DAY -nish ; endonym : dansk pronounced [ˈtænˀsk] , dansk sprog [ˈtænˀsk ˈspʁɔwˀ] )

9522-412: The descriptions of bird species. These skin collections have been used in more recent times for studies on molecular phylogenetics by the extraction of ancient DNA . The importance of type specimens in the description of species make skin collections a vital resource for systematic ornithology. However, with the rise of molecular techniques, establishing the taxonomic status of new discoveries, such as

9660-478: The development of the discipline of landscape ecology . John Hurrell Crook studied the behaviour of weaverbirds and demonstrated the links between ecological conditions, behaviour, and social systems. Principles from economics were introduced to the study of biology by Jerram L. Brown in his work on explaining territorial behaviour. This led to more studies of behaviour that made use of cost-benefit analyses . The rising interest in sociobiology also led to

9798-559: The distribution patterns of birds. For Darwin, the problem was how species arose from a common ancestor, but he did not attempt to find rules for delineation of species. The species problem was tackled by the ornithologist Ernst Mayr , who was able to demonstrate that geographical isolation and the accumulation of genetic differences led to the splitting of species . Early ornithologists were preoccupied with matters of species identification. Only systematics counted as true science and field studies were considered inferior through much of

9936-417: The early years the zoo focused on showing as many different types of animals as possible, but as animal welfare later became an issue, the number of different species has dropped in favour of more space to each animal. In 1901 the zoo had a human display with 25 Indians - men, women and children- in an exhibition where the "brown exotic" people went about their daily lives in palm tree leaf huts constructed in

10074-436: The editorial policy of the journal, leading both to a unification of field and laboratory studies and a shift of research from museums to universities. Ornithology in the United States continued to be dominated by museum studies of morphological variations, species identities, and geographic distributions, until it was influenced by Stresemann's student Ernst Mayr . In Britain, some of the earliest ornithological works that used

10212-492: The environment. Camera traps have been found to be a useful tool for the detection and documentation of elusive species, nest predators and in the quantitative analysis of frugivory, seed dispersal and behaviour. Many aspects of bird biology are difficult to study in the field. These include the study of behavioural and physiological changes that require a long duration of access to the bird. Nondestructive samples of blood or feathers taken during field studies may be studied in

10350-508: The exclusive use of rigsdansk , the High Copenhagen Standard, in national broadcasting, the traditional dialects came under increased pressure. In the 20th century, they have all but disappeared, and the standard language has extended throughout the country. Minor regional pronunciation variation of the standard language, sometimes called regionssprog ("regional languages") remain, and are in some cases vital. Today,

10488-511: The expression of Bmp4 have been shown to be associated with changes in the growth and shape of the beak. The chicken has long been a model organism for studying vertebrate developmental biology . As the embryo is readily accessible, its development can be easily followed (unlike mice ). This also allows the use of electroporation for studying the effect of adding or silencing a gene. Other tools for perturbing their genetic makeup are chicken embryonic stem cells and viral vectors . With

10626-512: The expression of genes and behaviour may be studied using candidate genes. Variations in the exploratory behaviour of great tits ( Parus major ) have been found to be linked with a gene orthologous to the human gene DRD4 (Dopamine receptor D4) which is known to be associated with novelty-seeking behaviour. The role of gene expression in developmental differences and morphological variations have been studied in Darwin's finches . The difference in

10764-451: The first English-language grammar of Danish. Literary Danish continued to develop with the works of Ludvig Holberg , whose plays and historical and scientific works laid the foundation for the Danish literary canon. With the Danish colonization of Greenland by Hans Egede , Danish became the administrative and religious language there, while Iceland and the Faroe Islands had the status of Danish colonies with Danish as an official language until

10902-457: The first major system of bird classification that was based on function and morphology rather than on form or behaviour. Willughby's Ornithologiae libri tres (1676) completed by John Ray is sometimes considered to mark the beginning of scientific ornithology. Ray also worked on Ornithologia , which was published posthumously in 1713 as Synopsis methodica avium et piscium . The earliest list of British birds, Pinax Rerum Naturalium Britannicarum ,

11040-439: The fitness of individuals. Others, such as Wynne-Edwards , interpreted population regulation as a mechanism that aided the "species" rather than individuals . This led to widespread and sometimes bitter debate on what constituted the "unit of selection". Lack also pioneered the use of many new tools for ornithological research, including the idea of using radar to study bird migration. Birds were also widely used in studies of

11178-491: The formation of huge collections of bird skins in museums in Europe and North America. Many private collections were also formed. These became references for comparison of species, and the ornithologists at these museums were able to compare species from different locations, often places that they themselves never visited. Morphometrics of these skins, particularly the lengths of the tarsus, bill, tail, and wing became important in

11316-580: The habit of bird migration , moulting, egg laying, and lifespans, as well as compiling a list of 170 different bird species. However, he also introduced and propagated several myths, such as the idea that swallows hibernated in winter, although he noted that cranes migrated from the steppes of Scythia to the marshes at the headwaters of the Nile . The idea of swallow hibernation became so well established that even as late as in 1878, Elliott Coues could list as many as 182 contemporary publications dealing with

11454-476: The hand. The earliest of the new generation of field guides was prepared by Florence Merriam , sister of Clinton Hart Merriam , the mammalogist. This was published in 1887 in a series Hints to Audubon Workers: Fifty Birds and How to Know Them in Grinnell's Audubon Magazine . These were followed by new field guides, from the pioneering illustrated handbooks of Frank Chapman to the classic Field Guide to

11592-513: The hibernation of swallows and little published evidence to contradict the theory. Similar misconceptions existed regarding the breeding of barnacle geese . Their nests had not been seen, and they were believed to grow by transformations of goose barnacles , an idea that became prevalent from around the 11th century and noted by Bishop Giraldus Cambrensis ( Gerald of Wales ) in Topographia Hiberniae (1187). Around 77 AD, Pliny

11730-426: The history of Danish into a period from 800 AD to 1525 to be "Old Danish", which he subdivided into "Runic Danish" (800–1100), Early Middle Danish (1100–1350) and Late Middle Danish (1350–1525). Móðir Dyggva var Drótt, dóttir Danps konungs, sonar Rígs er fyrstr var konungr kallaðr á danska tungu . " Dyggvi 's mother was Drott, the daughter of king Danp, Ríg 's son, who was the first to be called king in

11868-439: The introduction of trinomial names. The search for patterns in the variations of birds was attempted by many. Friedrich Wilhelm Joseph Schelling (1775–1854), his student Johann Baptist von Spix (1781–1826), and several others believed that a hidden and innate mathematical order existed in the forms of birds. They believed that a "natural" classification was available and superior to "artificial" ones. A particularly popular idea

12006-476: The labels associated with these early egg collections made them unreliable for the serious study of bird breeding. To preserve eggs, a tiny hole was made and the contents extracted. This technique became standard with the invention of the blow drill around 1830. Egg collection is no longer popular; however, historic museum collections have been of value in determining the effects of pesticides such as DDT on physiology. Museum bird collections continue to act as

12144-534: The laboratory and out in the field, and innovations are constantly made. Most biologists who recognise themselves as "ornithologists" study specific biology research areas, such as anatomy , physiology , taxonomy , ecology , or behaviour . The word "ornithology" comes from the late 16th-century Latin ornithologia meaning "bird science" from the Greek ὄρνις ornis ("bird") and λόγος logos ("theory, science, thought"). The history of ornithology largely reflects

12282-510: The laboratory. For instance, the variation in the ratios of stable hydrogen isotopes across latitudes makes establishing the origins of migrant birds possible using mass spectrometric analysis of feather samples. These techniques can be used in combination with other techniques such as ringing. The first attenuated vaccine developed by Louis Pasteur , for fowl cholera, was tested on poultry in 1878. Anti-malarials were tested on birds which harbour avian-malarias. Poultry continues to be used as

12420-424: The language as a token of Danish identity, and the language experienced a strong surge in use and popularity, with major works of literature produced in the 18th and 19th centuries. Today, traditional Danish dialects have all but disappeared, though regional variants of the standard language exist. The main differences in language are between generations, with youth language being particularly innovative. Danish has

12558-547: The language of the courts. Since 1997, public authorities have been obliged to follow the official spelling system laid out in the Orthography Law. In the 21st century, discussions have been held with a view to create a law that would make Danish the official language of Denmark. In addition, a noticeable community of Danish speakers is in Southern Schleswig , the portion of Germany bordering Denmark, and

12696-407: The major varieties of Standard Danish are High Copenhagen Standard, associated with elderly, well to-do, and well educated people of the capital, and low Copenhagen speech traditionally associated with the working class, but today adopted as the prestige variety of the younger generations. Also, in the 21st century, the influence of immigration has had linguistic consequences, such as the emergence of

12834-422: The mid-20th century. Moders navn er vort Hjertesprog, kun løs er al fremmed Tale. Det alene i mund og bog, kan vække et folk af dvale. "Mother's name is our hearts' tongue, only idle is all foreign speech It alone, in mouth or in book, can rouse a people from sleep." N.F.S. Grundtvig , "Modersmaalet" Following the loss of Schleswig to Germany, a sharp influx of German speakers moved into

12972-533: The middle of the zoo. One of the most notable animals kept there was a male slow worm that lived there from 1892 to 1946 (for 54 years, which is a record among lizards ). Starting in the early 1980s, Copenhagen Zoo has been undergoing a renovations aimed at replacing cages with enclosures which recreate animals' natural environments, giving a better lifestyle to the animals, and a more realistic experience to visitors. The Elephant House and 1.5-hectare (3.7-acre) Savanna are results of these efforts. The Savanna includes

13110-454: The niche hypothesis and Georgii Gause 's competitive exclusion principle. Work on resource partitioning and the structuring of bird communities through competition were made by Robert MacArthur . Patterns of biodiversity also became a topic of interest. Work on the relationship of the number of species to area and its application in the study of island biogeography was pioneered by E. O. Wilson and Robert MacArthur . These studies led to

13248-463: The oldest indications of an interest in birds. Birds were perhaps important as food sources, and bones of as many as 80 species have been found in excavations of early Stone Age settlements. Waterbird and seabird remains have also been found in shell mounds on the island of Oronsay off the coast of Scotland . Cultures around the world have rich vocabularies related to birds. Traditional bird names are often based on detailed knowledge of

13386-504: The order within the crow family. Where he failed to find five genera, he left a blank insisting that a new genus would be found to fill these gaps. These ideas were replaced by more complex "maps" of affinities in works by Hugh Edwin Strickland and Alfred Russel Wallace . A major advance was made by Max Fürbringer in 1888, who established a comprehensive phylogeny of birds based on anatomy, morphology, distribution, and biology. This

13524-689: The papers in American ornithology journals were written by persons who were not employed in biology related work. Organizations were started in many countries, and these grew rapidly in membership, most notable among them being the Royal Society for the Protection of Birds (RSPB) in Britain and the Audubon Society in the US, which started in 1885. Both these organizations were started with

13662-418: The part of the zoo called "The Nordics " ( Danish : Norden ), visitors can see species such as harbour seals , snowy owls , reindeer , musk oxen , brown bears , arctic foxes and grey wolves . The "Arctic Ring" ( Danish : Den Arktiske Ring ), which opened in 2013, has an exhibit for polar bears (including a tunnel allowing underwater views) and an aviary for North Atlantic seabirds ( alcids ). In

13800-438: The pride's natural structure and behaviour. The lions were not culled or dissected in public. Copenhagen Zoo is used as location in the films Københavnere (1933), Wienerbarnet (1941), Hold fingrene fra mor (1951), Far til fire på landet (1955), Vi er allesammen tossede (1959), Rikki og mændene (1962), Den store badedag (1991), Jungledyret Hugo (1993), and Far til fire på japansk (2010). The zoo

13938-468: The primary objective of conservation. The RSPB, born in 1889, grew from a small Croydon -based group of women, including Eliza Phillips , Etta Lemon , Catherine Hall and Hannah Poland . Calling themselves the "Fur, Fin, and Feather Folk", the group met regularly and took a pledge "to refrain from wearing the feathers of any birds not killed for the purpose of food, the ostrich only exempted." The organization did not allow men as members initially, avenging

14076-505: The relative lengths of wing feathers in warblers) are vital in establishing identity. Captured birds are often marked for future recognition. Rings or bands provide long-lasting identification, but require capture for the information on them to be read. Field-identifiable marks such as coloured bands, wing tags, or dyes enable short-term studies where individual identification is required. Mark and recapture techniques make demographic studies possible. Ringing has traditionally been used in

14214-472: The shared written standard language remained). With the introduction of absolutism in 1660, the Danish state was further integrated, and the language of the Danish chancellery, a Zealandic variety with German and French influence, became the de facto official standard language , especially in writing—this was the original so-called rigsdansk ("Danish of the Realm"). Also, beginning in the mid-18th century,

14352-1032: The specimen "in the hand". The capture and marking of birds enable detailed studies of life history. Techniques for capturing birds are varied and include the use of bird liming for perching birds, mist nets for woodland birds, cannon netting for open-area flocking birds, the bal-chatri trap for raptors, decoys and funnel traps for water birds. The bird in the hand may be examined and measurements can be made, including standard lengths and weights. Feather moult and skull ossification provide indications of age and health. Sex can be determined by examination of anatomy in some sexually nondimorphic species. Blood samples may be drawn to determine hormonal conditions in studies of physiology, identify DNA markers for studying genetics and kinship in studies of breeding biology and phylogeography. Blood may also be used to identify pathogens and arthropod-borne viruses . Ectoparasites may be collected for studies of coevolution and zoonoses . In many cryptic species, measurements (such as

14490-616: The study of living birds was merely recreation held sway until ecological theories became the predominant focus of ornithological studies. The study of birds in their habitats was particularly advanced in Germany with bird ringing stations established as early as 1903. By the 1920s, the Journal für Ornithologie included many papers on the behaviour, ecology, anatomy, and physiology, many written by Erwin Stresemann . Stresemann changed

14628-499: The study of migration. In recent times, satellite transmitters provide the ability to track migrating birds in near-real time. Techniques for estimating population density include point counts , transects , and territory mapping. Observations are made in the field using carefully designed protocols and the data may be analysed to estimate bird diversity, relative abundance, or absolute population densities. These methods may be used repeatedly over large timespans to monitor changes in

14766-486: The three languages can often understand the others fairly well, though studies have shown that the mutual intelligibility is asymmetric: Norwegian speakers generally understand both Danish and Swedish far better than Swedes or Danes understand each other. Concomitantly, Swedes and Danes understand Norwegian better than they understand each other's languages. Norwegian occupies the middle position in terms of intelligibility because of its shared border with Sweden, resulting in

14904-457: The trends in the history of biology , as well as many other scientific disciplines, including ecology , anatomy , physiology , paleontology , and more recently, molecular biology. Trends include the move from mere descriptions to the identification of patterns, thus towards elucidating the processes that produce these patterns. Humans have had an observational relationship with birds since prehistory , with some stone-age drawings being amongst

15042-445: The turn of the 20th century. Danish itself can be divided into three main dialect areas: Jutlandic (West Danish), Insular Danish (including the standard variety), and East Danish (including Bornholmian and Scanian ). According to the view that Scandinavian is a dialect continuum, East Danish can be considered intermediary between Danish and Swedish, while Scanian can be considered a Swedified East Danish dialect, and Bornholmian

15180-411: The vernacular language to be accessible also to those who were not Latinate. The Jutlandic Law and Scanian Law were written in vernacular Danish in the early 13th century. Beginning in 1350, Danish began to be used as a language of administration, and new types of literature began to be written in the language, such as royal letters and testaments. The orthography in this period was not standardized nor

15318-469: The vernacular, such as the plural form of verbs, should be conserved in writing (i.e. han er "he is" vs. de ere "they are"). The East Danish provinces were lost to Sweden after the Second Treaty of Brömsebro (1645) after which they were gradually Swedified; just as Norway was politically severed from Denmark, beginning also a gradual end of Danish influence on Norwegian (influence through

15456-516: The whore-bed with another man's wife and he comes away alive..." Jutlandic Law, 1241 In the medieval period, Danish emerged as a separate language from Swedish. The main written language was Latin, and the few Danish-language texts preserved from this period are written in the Latin alphabet, although the runic alphabet seems to have lingered in popular usage in some areas. The main text types written in this period are laws, which were formulated in

15594-1084: The widespread interest in birds, use of a large number of people to work on collaborative ornithological projects that cover large geographic scales has been possible. These citizen science projects include nationwide projects such as the Christmas Bird Count , Backyard Bird Count, the North American Breeding Bird Survey , the Canadian EPOQ or regional projects such as the Asian Waterfowl Census and Spring Alive in Europe. These projects help to identify distributions of birds, their population densities and changes over time, arrival and departure dates of migration, breeding seasonality, and even population genetics. The results of many of these projects are published as bird atlases . Studies of migration using bird ringing or colour marking often involve

15732-492: The word by , meaning ‘village’ or ‘town’, occurs in many English place-names, such as Whitby and Selby , as remnants of the Viking occupation . During that period English adopted ‘are’, the third person plural form of the verb ‘to be’, as well as the personal pronouns ‘they’, ‘them’ and ‘their’ from contemporary Old Norse. Danish is largely mutually intelligible with Norwegian and Swedish . A proficient speaker of any of

15870-463: The word ecology appeared in 1915. The Ibis , however, resisted the introduction of these new methods of study, and no paper on ecology appeared until 1943. The work of David Lack on population ecology was pioneering. Newer quantitative approaches were introduced for the study of ecology and behaviour, and this was not readily accepted. For instance, Claud Ticehurst wrote: Sometimes it seems that elaborate plans and statistics are made to prove what

16008-566: The works of ancient writers from Greek and Syriac . In the 12th and 13th centuries, crusades and conquest had subjugated Islamic territories in southern Italy, central Spain, and the Levant under European rule, and for the first time translations into Latin of the great works of Arabic and Greek scholars were made with the help of Jewish and Muslim scholars, especially in Toledo , which had fallen into Christian hands in 1085 and whose libraries had escaped destruction. Michael Scotus from Scotland made

16146-576: The world. Artificial incubation of poultry was practised in China around 246 BC and around at least 400 BC in Egypt. The Egyptians also made use of birds in their hieroglyphic scripts, many of which, though stylized, are still identifiable to species. Early written records provide valuable information on the past distributions of species. For instance, Xenophon records the abundance of the ostrich in Assyria (Anabasis, i. 5); this subspecies from Asia Minor

16284-489: The written languages are compatible, spoken Danish is distinctly different from Norwegian and Swedish and thus the degree of mutual intelligibility with either is variable between regions and speakers . Until the 16th century, Danish was a continuum of dialects spoken from Southern Jutland and Schleswig to Scania with no standard variety or spelling conventions. With the Protestant Reformation and

16422-595: The zoo is the wooden observation tower. 43.5 metres (142.7 ft) high, it offers views of the surrounding parklands and city. The tower was built in 1905 and is one of the tallest observation towers built of wood in the world. Its base is similar to that of Eiffel Tower . Animals exhibited at the zoo that are not housed in any of the main areas include bactrian camels , American flamingos , scarlet ibises , roseate spoonbills , Dalmatian pelicans , turkey vultures , Humboldt penguins , California sea lions , black-capped squirrel monkeys , chimpanzees and lions . In

16560-557: The zoo. These were the first Tasmanian devils to live outside of Australia. The area has since been expanded with a combined area for red-necked wallabies , Eastern Grey Kangaroos and common wombats . Parts of this area is also accessible by visitors, allowing kangaroos and people to cross paths. In the "South America" ( Danish : Sydamerika ) section, visitors can see species such as capybaras , guanacos , greater rheas , Patagonian maras , southern screamers and giant anteaters . They all have access to one common area, modeled after

16698-492: Was a territory ruled by Denmark–Norway , one of whose official languages was Danish. Though Danish ceased to be an official language in Iceland in 1944, it is still widely used and is a mandatory subject in school, taught as a second foreign language after English. No law stipulates an official language for Denmark, making Danish the de facto official language only. The Code of Civil Procedure does, however, lay down Danish as

16836-446: Was also due in part to colonialism . At 100 years later, in 1959, R. E. Moreau noted that ornithology in this period was preoccupied with the geographical distributions of various species of birds. No doubt the preoccupation with widely extended geographical ornithology, was fostered by the immensity of the areas over which British rule or influence stretched during the 19th century and for some time afterwards. The bird collectors of

16974-457: Was also one of two official languages of Greenland (alongside Greenlandic ). Danish now acts as a lingua franca in Greenland, with a large percentage of native Greenlanders able to speak Danish as a second language (it was introduced into the education system as a compulsory language in 1928). About 10% of the population speaks Danish as their first language , due to immigration. Iceland

17112-514: Was criticized by Stine Jensen from Denmark's Organisation Against the Suffering of Animals, who repudiated the action as unethical. She said, "This situation should not have occurred at all. It just shows that the zoo is in fact not the ethical institution that it wants to portray itself as being, because here you have a waste product - that being Marius. Here we have a zoo which thinks that putting this giraffe down instead of thinking of alternatives

17250-400: Was developed further by Hans Gadow and others. The Galapagos finches were especially influential in the development of Charles Darwin 's theory of evolution. His contemporary Alfred Russel Wallace also noted these variations and the geographical separations between different forms leading to the study of biogeography . Wallace was influenced by the work of Philip Lutley Sclater on

17388-706: Was involved in establishing Gaetz Lakes bird sanctuary. In the United States, the Breeding Bird Surveys , conducted by the United States Geological Survey , have also produced atlases with information on breeding densities and changes in the density and distribution over time. Other volunteer collaborative ornithology projects were subsequently established in other parts of the world. The tools and techniques of ornithology are varied, and new inventions and approaches are quickly incorporated. The techniques may be broadly dealt under

17526-470: Was killed on 9 February 2014 on the recommendation of the European Association of Zoos and Aquaria . His meat was fed to the zoo's lions, while other parts were transferred to a scientific institution. The scientific director at Copenhagen Zoo, Bengt Holst defended the culling, saying that the giraffes at the zoo bred very well, and where this was the case, giraffes had to be killed to ensure

17664-488: Was mostly supplanted by the Low German spise . As well as loanwords, new words can be freely formed by compounding existing words. In standard texts of contemporary Danish, Middle Low German loans account for about 16–17% of the vocabulary, Graeco-Latin loans 4–8%, French 2–4% and English about 1%. Danish and English are both Germanic languages. Danish is a North Germanic language descended from Old Norse, and English

17802-627: Was once the Viking settlement of Jorvik. Several other English words derive from Old East Norse, for example "knife" ( kniv ), "husband" ( husbond ), and "egg" ( æg ). The suffix "-by" for 'town' is common in place names in Yorkshire and the east Midlands, for example Selby, Whitby, Derby, and Grimsby. The word "dale" meaning valley is common in Yorkshire and Derbyshire placenames. Fangær man saar i hor seng mæth annæns mansz kunæ. oc kumær han burt liuænd... . "If one catches someone in

17940-491: Was part of Ibn Sīnā's massive Kitāb al-Šifāʾ . Frederick II eventually wrote his own treatise on falconry, the De arte venandi cum avibus , in which he related his ornithological observations and the results of the hunts and experiments his court enjoyed performing. Several early German and French scholars compiled old works and conducted new research on birds. These included Guillaume Rondelet , who described his observations in

18078-668: Was pioneered by Charles Sibley and Jon Edward Ahlquist , resulting in what is called the Sibley–Ahlquist taxonomy . These early techniques have been replaced by newer ones based on mitochondrial DNA sequences and molecular phylogenetics approaches that make use of computational procedures for sequence alignment , construction of phylogenetic trees , and calibration of molecular clocks to infer evolutionary relationships. Molecular techniques are also widely used in studies of avian population biology and ecology. The use of field glasses or telescopes for bird observation began in

18216-445: Was principally concerned with descriptions and distributions of species, ornithologists today seek answers to very specific questions, often using birds as models to test hypotheses or predictions based on theories. Most modern biological theories apply across life forms, and the number of scientists who identify themselves as "ornithologists" has therefore declined. A wide range of tools and techniques are used in ornithology, both inside

18354-463: Was still not standardized and the principles for doing so were vigorously discussed among Danish philologists. The grammar of Jens Pedersen Høysgaard was the first to give a detailed analysis of Danish phonology and prosody, including a description of the stød . In this period, scholars were also discussing whether it was best to "write as one speaks" or to "speak as one writes", including whether archaic grammatical forms that had fallen out of use in

18492-535: Was the Quinarian system popularised by Nicholas Aylward Vigors (1785–1840), William Sharp Macleay (1792–1865), William Swainson , and others. The idea was that nature followed a "rule of five" with five groups nested hierarchically. Some had attempted a rule of four, but Johann Jakob Kaup (1803–1873) insisted that the number five was special, noting that other natural entities such as the senses also came in fives. He followed this idea and demonstrated his view of

18630-469: Was the change of the diphthong æi (Old West Norse ei ) to the monophthong e , as in stæin to sten . This is reflected in runic inscriptions where the older read stain and the later stin . Also, a change of au as in dauðr into ø as in døðr occurred. This change is shown in runic inscriptions as a change from tauþr into tuþr . Moreover, the øy (Old West Norse ey ) diphthong changed into ø , as well, as in

18768-474: Was the spoken language, and the regional laws demonstrate the dialectal differences between the regions in which they were written. Throughout this period, Danish was in contact with Low German , and many Low German loan words were introduced in this period. With the Protestant Reformation in 1536, Danish also became the language of religion, which sparked a new interest in using Danish as

18906-426: Was used to interpret observations on behaviour and life history, and birds were widely used models for testing hypotheses based on theories postulated by W. D. Hamilton and others. The new tools of molecular biology changed the study of bird systematics, which changed from being based on phenotype to the underlying genotype . The use of techniques such as DNA–DNA hybridization to study evolutionary relationships

19044-436: Was written by Christopher Merrett in 1667, but authors such as John Ray considered it of little value. Ray did, however, value the expertise of the naturalist Sir Thomas Browne (1605–82), who not only answered his queries on ornithological identification and nomenclature, but also those of Willoughby and Merrett in letter correspondence. Browne himself in his lifetime kept an eagle, owl, cormorant, bittern, and ostrich, penned

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