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57-505: The Diplostraca or Cladocera , commonly known as water fleas , is a superorder of small, mostly freshwater crustaceans , most of which feed on microscopic chunks of organic matter, though some forms are predatory. Over 1000 species have been recognised so far, with many more undescribed . The oldest fossils of diplostracans date to the Jurassic , though their modern morphology suggests that they originated substantially earlier, during

114-461: A cohors (plural cohortes ). Some of the plant families still retain the names of Linnaean "natural orders" or even the names of pre-Linnaean natural groups recognized by Linnaeus as orders in his natural classification (e.g. Palmae or Labiatae ). Such names are known as descriptive family names. In the field of zoology , the Linnaean orders were used more consistently. That is,

171-521: A nauplius larval stage is absent in Diplostraca. Diplostraca are nested within the clam shrimp , being most closely related to the order Cyclestherida, the only living genus of which is Cyclestheria . Though several fossils from the Paleozoic have been claimed to represent fossils of diplostracans, none of these records can be confirmed. The oldest confirmed records of diplostracans are from

228-402: A taxonomist , as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with

285-509: A capital letter. For some groups of organisms, their orders may follow consistent naming schemes . Orders of plants , fungi , and algae use the suffix -ales (e.g. Dictyotales ). Orders of birds and fishes use the Latin suffix -iformes meaning 'having the form of' (e.g. Passeriformes ), but orders of mammals and invertebrates are not so consistent (e.g. Artiodactyla , Actiniaria , Primates ). For some clades covered by

342-622: A cataclysm known as " The Great Dying ", the third and most severe Phanerozoic mass extinction. The early Cambrian climate was probably moderate at first, becoming warmer over the course of the Cambrian, as the second-greatest sustained sea level rise in the Phanerozoic got underway. However, as if to offset this trend, Gondwana moved south, so that, in Ordovician time, most of West Gondwana (Africa and South America) lay directly over

399-570: A distinct rank of biological classification having its own distinctive name (and not just called a higher genus ( genus summum )) was first introduced by the German botanist Augustus Quirinus Rivinus in his classification of plants that appeared in a series of treatises in the 1690s. Carl Linnaeus was the first to apply it consistently to the division of all three kingdoms of nature (then minerals , plants , and animals ) in his Systema Naturae (1735, 1st. Ed.). For plants, Linnaeus' orders in

456-445: A foothold on land. These early plants were the forerunners of all plant life on land. During this time, there were four continents: Gondwana (Africa, South America, Australia, Antarctica, Siberia), Laurentia (North America), Baltica (Northern Europe), and Avalonia (Western Europe). The recent rise in sea levels allowed many new species to thrive in water. The Devonian spanned from 419–359 million years ago. Also known as "The Age of

513-469: Is angled downwards, and may be separated from the rest of the body by a "cervical sinus" or notch. It bears a single black compound eye, located on the animal's midline, in all but two genera, and often, a single ocellus is present. The head also bears two pairs of antennae – the first antennae are small, unsegmented appendages, while the second antennae are large, segmented, and branched, with powerful muscles. The first antennae bear olfactory setae , while

570-663: Is considered the first Phanerozoic mass extinction event, and the second deadliest. The Silurian spanned from 444–419 million years ago. The Silurian saw the rejuvenation of life as the Earth recovered from the previous glaciation. This period saw the mass evolution of fish, as jawless fish became more numerous, jawed fish evolved, and the first freshwater fish evolved, though arthropods, such as sea scorpions , were still apex predators . Fully terrestrial life evolved, including early arachnids, fungi, and centipedes. The evolution of vascular plants ( Cooksonia ) allowed plants to gain

627-406: Is lost, and oxygen taken up, through the body surface. With the exception of a few purely asexual species, the lifecycle of diplostracans is dominated by asexual reproduction, with occasional periods of sexual reproduction; this is known as cyclical parthenogenesis . When conditions are favourable, reproduction occurs by parthenogenesis for several generations, producing only female clones . As

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684-433: Is occasionally supplemented by sexual reproduction , which produces resting eggs that allow the species to survive harsh conditions and disperse to distant habitats. They are mostly 0.2–6.0 mm (0.01–0.24 in) long, with the exception of Leptodora , which can be up to 18 mm (0.71 in) long. The body is not obviously segmented and bears a folded carapace which covers the thorax and abdomen . The head

741-575: Is the sudden appearance of nearly all of the invertebrate animal phyla in great abundance at the beginning of the Cambrian. The first vertebrates appeared in the form of primitive fish, which greatly diversified in the Silurian and Devonian Periods. The first animals to venture onto dry land were the arthropods. Some fish had lungs, and powerful bony fins that in the late Devonian, 367.5 million years ago, allowed them to crawl onto land. The bones in their fins eventually evolved into legs and they became

798-605: The Appalachians , Caledonides , Ural Mountains , and mountains of Tasmania . The Cambrian spanned from 539–485 million years ago and is the first period of the Paleozoic Era of the Phanerozoic. The Cambrian marked a boom in evolution in an event known as the Cambrian explosion in which the largest number of creatures evolved in any single period of the history of the Earth. Creatures like algae evolved, but

855-531: The Carboniferous Rainforest Collapse . Gondwana was glaciated as much of it was situated around the south pole. The Permian spanned from 299–252 million years ago and was the last period of the Paleozoic Era. At the beginning of this period, all continents joined together to form the supercontinent Pangaea, which was encircled by one ocean called Panthalassa . The land mass was very dry during this time, with harsh seasons, as

912-645: The Early Palaeozoic Icehouse , culminating in the Hirnantian glaciation, 445  million years ago at the end of the Ordovician. The middle Paleozoic was a time of considerable stability. Sea levels had dropped coincident with the ice age, but slowly recovered over the course of the Silurian and Devonian. The slow merger of Baltica and Laurentia, and the northward movement of bits and pieces of Gondwana created numerous new regions of relatively warm, shallow sea floor. As plants took hold on

969-528: The International Code of Zoological Nomenclature , several additional classifications are sometimes used, although not all of these are officially recognized. In their 1997 classification of mammals , McKenna and Bell used two extra levels between superorder and order: grandorder and mirorder . Michael Novacek (1986) inserted them at the same position. Michael Benton (2005) inserted them between superorder and magnorder instead. This position

1026-425: The Paleozoic . Some have also adapted to a life in the ocean, the only members of Branchiopoda to do so, though several anostracans live in hypersaline lakes . Most are 0.2–6.0 mm (0.01–0.24 in) long, with a down-turned head with a single median compound eye , and a carapace covering the apparently unsegmented thorax and abdomen. Most species show cyclical parthenogenesis , where asexual reproduction

1083-571: The South Pole . The early Paleozoic climate was strongly zonal, with the result that the "climate", in an abstract sense, became warmer, but the living space of most organisms of the time – the continental shelf marine environment – became steadily colder. However, Baltica (Northern Europe and Russia) and Laurentia (eastern North America and Greenland) remained in the tropical zone, while China and Australia lay in waters which were at least temperate. The early Paleozoic ended, rather abruptly, with

1140-815: The Systema Naturae and the Species Plantarum were strictly artificial, introduced to subdivide the artificial classes into more comprehensible smaller groups. When the word ordo was first consistently used for natural units of plants, in 19th-century works such as the Prodromus Systematis Naturalis Regni Vegetabilis of Augustin Pyramus de Candolle and the Genera Plantarum of Bentham & Hooker, it indicated taxa that are now given

1197-550: The coal beds of Europe and eastern North America . Towards the end of the era, large, sophisticated synapsids and diapsids were dominant and the first modern plants ( conifers ) appeared. The Paleozoic Era ended with the largest extinction event of the Phanerozoic Eon , the Permian–Triassic extinction event . The effects of this catastrophe were so devastating that it took life on land 30 million years into

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1254-670: The Early Jurassic of Asia. Fossils from the Jurassic are assignable to modern as well as extinct groups, indicating that the initial radiation of the group occurred prior to the beginning of the Jurassic, likely during the late Paleozoic . A Devonian fossil, Ebullitiocaris , is tentatively placed as a diplostracan, however since it is only known from its carapace this is uncertain. Most diplostracan species live in fresh water and other inland water bodies, with only eight species being truly oceanic . The marine species are all in

1311-648: The Fish", the Devonian featured a huge diversification of fish, including armored fish like Dunkleosteus and lobe-finned fish which eventually evolved into the first tetrapods. On land, plant groups diversified rapidly in an event known as the Devonian explosion when plants made lignin , leading to taller growth and vascular tissue; the first trees and seeds evolved. These new habitats led to greater arthropod diversification. The first amphibians appeared and fish occupied

1368-658: The Mesozoic Era to recover. Recovery of life in the sea may have been much faster. The base of the Paleozoic is one of the major divisions in geological time representing the divide between the Proterozoic and Phanerozoic eons, the Paleozoic and Neoproterozoic eras and the Ediacaran and Cambrian periods. When Adam Sedgwick named the Paleozoic in 1835, he defined the base as the first appearance of complex life in

1425-610: The Middle Carboniferous). An important evolutionary development of the time was the evolution of amniotic eggs , which allowed amphibians to move farther inland and remain the dominant vertebrates for the duration of this period. Also, the first reptiles and synapsids evolved in the swamps. Throughout the Carboniferous, there was a cooling trend, which led to the Permo-Carboniferous glaciation or

1482-629: The Palaeozoic had very few facultatively motile animals that could easily adjust to disturbance, with such creatures composing 1% of its assemblages in contrast to 50% in Cenozoic faunal assemblages. Non-motile animals untethered to the substrate, extremely rare in the Cenozoic, were abundant in the Palaeozoic. Palaeozoic phytoplankton overall were both nutrient-poor themselves and adapted to nutrient-poor environmental conditions. This phytoplankton nutrient poverty has been cited as an explanation for

1539-459: The Paleozoic and Mesozoic eras and the Permian and Triassic periods is marked by the first occurrence of the conodont Hindeodus parvus . This is the first biostratigraphic event found worldwide that is associated with the beginning of the recovery following the end- Permian mass extinctions and environmental changes. In non-marine strata, the equivalent level is marked by the disappearance of

1596-660: The Permian Dicynodon tetrapods . This means events previously considered to mark the Permian-Triassic boundary, such as the eruption of the Siberian Traps flood basalts , the onset of greenhouse climate, ocean anoxia and acidification and the resulting mass extinction are now regarded as being of latest Permian in age. The GSSP is near Meishan , Zhejiang Province, southern China. Radiometric dating of volcanic clay layers just above and below

1653-529: The Silurian Period, about 420 million years ago, when they began to transition onto dry land. Terrestrial flora reached its climax in the Carboniferous, when towering lycopsid rainforests dominated the tropical belt of Euramerica . Climate change caused the Carboniferous Rainforest Collapse which fragmented this habitat, diminishing the diversity of plant life in the late Carboniferous and Permian periods. A noteworthy feature of Paleozoic life

1710-549: The basal Cambrian Global Stratotype Section and Point (GSSP) at the base of the Treptichnus pedum assemblage of trace fossils and immediately above the last occurrence of the Ediacaran problematica fossils Harlaniella podolica and Palaeopsacichnus . The base of the Phanerozoic, Paleozoic and Cambrian is dated at 538.8+/-0.2 Ma and now lies below both the first appearance of trilobites and SSF. The boundary between

1767-516: The boundary confine its age to a narrow range of 251.902+/-0.024 Ma. The beginning of the Paleozoic Era witnessed the breakup of the supercontinent of Pannotia and ended while the supercontinent Pangaea was assembling. The breakup of Pannotia began with the opening of the Iapetus Ocean and other Cambrian seas and coincided with a dramatic rise in sea level. Paleoclimatic studies and evidence of glaciers indicate that Central Africa

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1824-511: The climate of the interior of Pangaea was not regulated by large bodies of water. Diapsids and synapsids flourished in the new dry climate. Creatures such as Dimetrodon and Edaphosaurus ruled the new continent. The first conifers evolved, and dominated the terrestrial landscape. Near the end of the Permian, however, Pangaea grew drier. The interior was desert, and new taxa such as Scutosaurus and Gorgonopsids filled it. Eventually they disappeared, along with 95% of all life on Earth, in

1881-399: The conditions deteriorate, males are produced, and sexual reproduction occurs. This results in the production of long-lasting dormant eggs . These ephippial eggs can be transported over land by wind, and hatch when they reach favourable conditions, allowing many species to have very wide – even cosmopolitan – distributions . Except for the genus Leptodora, which has a metanauplius stage,

1938-561: The continental margins, oxygen levels increased and carbon dioxide dropped, although much less dramatically. The north–south temperature gradient also seems to have moderated, or metazoan life simply became hardier, or both. At any event, the far southern continental margins of Antarctica and West Gondwana became increasingly less barren. The Devonian ended with a series of turnover pulses which killed off much of middle Paleozoic vertebrate life, without noticeably reducing species diversity overall. There are many unanswered questions about

1995-507: The empty continent of Gondwana. By the end of the Ordovician, Gondwana was at the south pole, early North America had collided with Europe, closing the intervening ocean. Glaciation of Africa resulted in a major drop in sea level, killing off all life that had established along coastal Gondwana. Glaciation may have caused the Ordovician–Silurian extinction events , in which 60% of marine invertebrates and 25% of families became extinct, and

2052-560: The end of the Permian period. In late middle Permian the pareiasaurs originated, successful herbivores and the only sauropsids that could reach sizes comparable to some of the largest synapsids. The Palaeozoic marine fauna was notably lacking in predators relative to the present day. Predators made up about 4% of the fauna in Palaeozoic assemblages while making up 17% of temperate Cenozoic assemblages and 31% of tropical ones. Infaunal animals made up 4% of soft substrate Palaeozoic communities but about 47% of Cenozoic communities. Additionally,

2109-570: The family Podonidae , except for the genus Penilia . Some diplostracans inhabit leaf litter . According to the World Registry of Marine Species, Cladocera is a synonym of the superorder Diplostraca, which is included in the class Branchiopoda . Both names are currently in use. The superorder forms a monophyletic group of 7 orders, about 24 families, and more than 11,000 species. Many more species remain undescribed . The genus Daphnia alone contains around 150 species. Many groups of

2166-461: The first tetrapods, 390  million years ago , and began to develop lungs. Amphibians were the dominant tetrapods until the mid-Carboniferous, when climate change greatly reduced their diversity, allowing amniotes to take over. Amniotes would split into two clades shortly after their origin in the Carboniferous; the synapsids, which was the dominant group, and the sauropsids . The synapsids continued to prosper and increase in number and variety till

2223-548: The late Paleozoic. The Mississippian (early Carboniferous Period) began with a spike in atmospheric oxygen, while carbon dioxide plummeted to new lows. This destabilized the climate and led to one, and perhaps two, ice ages during the Carboniferous. These were far more severe than the brief Late Ordovician ice age; but, this time, the effects on world biota were inconsequential. By the Cisuralian Epoch, both oxygen and carbon dioxide had recovered to more normal levels. On

2280-504: The majority of Ediacaran to Cambrian rock sequences are composed of siliciclastic rocks where skeletal fossils are rarely preserved. This led the International Commission on Stratigraphy (ICS) to use trace fossils as an indicator of complex life. Unlike later in the fossil record, Cambrian trace fossils are preserved in a wide range of sediments and environments, which aids correlation between different sites around

2337-434: The most rapid and widespread diversification of life in Earth's history, known as the Cambrian explosion , in which most modern phyla first appeared. Arthropods , molluscs , fish , amphibians , reptiles , and synapsids all evolved during the Paleozoic. Life began in the ocean but eventually transitioned onto land, and by the late Paleozoic, great forests of primitive plants covered the continents, many of which formed

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2394-603: The most ubiquitous of that period were the armored arthropods, like trilobites. Almost all marine phyla evolved in this period. During this time, the supercontinent Pannotia begins to break up, most of which later became the supercontinent Gondwana. The Ordovician spanned from 485–444 million years ago. The Ordovician was a time in Earth's history in which many of the biological classes still prevalent today evolved, such as primitive fish, cephalopods, and coral. The most common forms of life, however, were trilobites, snails and shellfish. The first arthropods went ashore to colonize

2451-708: The orders in the zoology part of the Systema Naturae refer to natural groups. Some of his ordinal names are still in use, e.g. Lepidoptera (moths and butterflies) and Diptera (flies, mosquitoes, midges, and gnats). In virology , the International Committee on Taxonomy of Viruses 's virus classification includes fifteen taxomomic ranks to be applied for viruses , viroids and satellite nucleic acids : realm , subrealm , kingdom , subkingdom, phylum , subphylum , class, subclass, order, suborder, family, subfamily , genus, subgenus , and species. There are currently fourteen viral orders, each ending in

2508-520: The other hand, the assembly of Pangaea created huge arid inland areas subject to temperature extremes. The Lopingian Epoch is associated with falling sea levels, increased carbon dioxide and general climatic deterioration, culminating in the devastation of the Permian extinction. While macroscopic plant life appeared early in the Paleozoic Era and possibly late in the Neoproterozoic Era of the earlier eon, plants mostly remained aquatic until

2565-564: The precursor of the currently used International Code of Nomenclature for algae, fungi, and plants . In the first international Rules of botanical nomenclature from the International Botanical Congress of 1905, the word family ( familia ) was assigned to the rank indicated by the French famille , while order ( ordo ) was reserved for a higher rank, for what in the 19th century had often been named

2622-502: The rank of family (see ordo naturalis , ' natural order '). In French botanical publications, from Michel Adanson 's Familles naturelles des plantes (1763) and until the end of the 19th century, the word famille (plural: familles ) was used as a French equivalent for this Latin ordo . This equivalence was explicitly stated in the Alphonse Pyramus de Candolle 's Lois de la nomenclature botanique (1868),

2679-401: The rock record as shown by the presence of trilobite -dominated fauna. Since then evidence of complex life in older rock sequences has increased and by the second half of the 20th century, the first appearance of small shelly fauna (SSF), also known as early skeletal fossils, were considered markers for the base of the Paleozoic. However, whilst SSF are well preserved in carbonate sediments,

2736-546: The second are used for swimming by most species. The pattern of setae on the second antennae is useful for identification. The part of the head which projects in front of the first antennae is known as the rostrum or "beak". The mouthparts are small, and consist of an unpaired labrum, a pair of mandibles, a pair of maxillae, and an unpaired labium. They are used to eat "organic detritus of all kinds" and bacteria . The thorax bears five or six pairs of lobed, leaf-like appendages, each with numerous hairs or setae. Carbon dioxide

2793-479: The short, but apparently severe, late Ordovician ice age. This cold spell caused the second-greatest mass extinction of the Phanerozoic Eon. Over time, the warmer weather moved into the Paleozoic Era. The Ordovician and Silurian were warm greenhouse periods, with the highest sea levels of the Paleozoic (200 m above today's); the warm climate was interrupted only by a 30 million year cool period,

2850-646: The start of the Mesozoic Era. The Paleozoic is subdivided into six geologic periods (from oldest to youngest), Cambrian , Ordovician , Silurian , Devonian , Carboniferous and Permian . Some geological timescales divide the Paleozoic informally into early and late sub-eras: the Early Paleozoic consisting of the Cambrian, Ordovician and Silurian; the Late Paleozoic consisting of the Devonian, Carboniferous and Permian. The name Paleozoic

2907-613: The suffix -virales . Paleozoic The Paleozoic ( / ˌ p æ l i . ə ˈ z oʊ . ɪ k , - i . oʊ -, ˌ p eɪ -/ PAL-ee-ə-ZOH-ik , -⁠ee-oh- , PAY- ; or Palaeozoic ) Era is the first of three geological eras of the Phanerozoic Eon. Beginning 538.8 million years ago (Ma), it succeeds the Neoproterozoic (the last era of the Proterozoic Eon) and ends 251.9 Ma at

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2964-554: The top of the food chain. Earth's second Phanerozoic mass extinction event (a group of several smaller extinction events), the Late Devonian extinction , ended 70% of existing species. The Carboniferous is named after the large coal deposits laid down during the period. It spanned from 359–299 million years ago. During this time, average global temperatures were exceedingly high; the early Carboniferous averaged at about 20 degrees Celsius (but cooled to 10 °C during

3021-465: The water fleas are cryptic species or species flocks. The following families are recognised: Superorder Diplostraca Gerstaecker, 1866 (=Cladocera) The word "Cladocera" derives via Neo-Latin from the Ancient Greek κλάδος ( kládos , "branch") and κέρας ( kéras , "horn"). Superorder (biology) What does and does not belong to each order is determined by

3078-521: The world. Trace fossils reflect the complexity of the body plan of the organism that made them. Ediacaran trace fossils are simple, sub-horizontal feeding traces. As more complex organisms evolved, their more complex behaviour was reflected in greater diversity and complexity of the trace fossils they left behind. After two decades of deliberation, the ICS chose Fortune Head , Burin Peninsula, Newfoundland as

3135-418: Was adopted by Systema Naturae 2000 and others. In botany , the ranks of subclass and suborder are secondary ranks pre-defined as respectively above and below the rank of order. Any number of further ranks can be used as long as they are clearly defined. The superorder rank is commonly used, with the ending -anae that was initiated by Armen Takhtajan 's publications from 1966 onwards. The order as

3192-469: Was first used by Adam Sedgwick (1785–1873) in 1838 to describe the Cambrian and Ordovician periods. It was redefined by John Phillips (1800–1874) in 1840 to cover the Cambrian to Permian periods. It is derived from the Greek palaiós (παλαιός, "old") and zōḗ (ζωή, "life") meaning "ancient life". The Paleozoic was a time of dramatic geological, climatic, and evolutionary change. The Cambrian witnessed

3249-539: Was most likely in the polar regions during the early Paleozoic. The breakup of Pannotia was followed by the assembly of the huge continent Gondwana ( 510  million years ago ). By the mid-Paleozoic, the collision of North America and Europe produced the Acadian-Caledonian uplifts, and a subducting plate uplifted eastern Australia . By the late Paleozoic, continental collisions formed the supercontinent of Pangaea and created great mountain chains, including

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