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Anthocyanin

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59-460: Anthocyanins (from Ancient Greek ἄνθος ( ánthos )  'flower' and κυάνεος / κυανοῦς ( kuáneos/kuanoûs )  'dark blue'), also called anthocyans , are water-soluble vacuolar pigments that, depending on their pH , may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart named a chemical compound that gives flowers

118-431: A food colorant . Anthocyanins may be used as pH indicators because their color changes with pH; they are red or pink in acidic solutions (pH < 7), purple in neutral solutions (pH ≈ 7), greenish-yellow in alkaline solutions (pH > 7), and colorless in very alkaline solutions, where the pigment is completely reduced. Thus, more than five enzymes are required to synthesize these pigments, each working in concert. Even

177-543: A pitch accent . In Modern Greek, all vowels and consonants are short. Many vowels and diphthongs once pronounced distinctly are pronounced as /i/ ( iotacism ). Some of the stops and glides in diphthongs have become fricatives , and the pitch accent has changed to a stress accent . Many of the changes took place in the Koine Greek period. The writing system of Modern Greek, however, does not reflect all pronunciation changes. The examples below represent Attic Greek in

236-587: A 2010 review of scientific evidence concerning the possible health benefits of eating foods claimed to have "antioxidant properties" due to anthocyanins, the European Food Safety Authority concluded that 1) there was no basis for a beneficial antioxidant effect from dietary anthocyanins in humans, 2) there was no evidence of a cause-and-effect relationship between the consumption of anthocyanin-rich foods and protection of DNA , proteins , and lipids from oxidative damage , and 3) there

295-415: A blue color, Anthokyan, in his treatise " Die Farben der Blüthen " (English: The Colors of Flowers). Food plants rich in anthocyanins include the blueberry, raspberry, black rice , and black soybean, among many others that are red, blue, purple, or black. Some of the colors of autumn leaves are derived from anthocyanins. Anthocyanins belong to a parent class of molecules called flavonoids synthesized via

354-554: A breeding program using introgression lines of genetically modified organisms (but not incorporating them in the final purple tomato) to define the genetic basis of purple coloration in wild species that originally were from Chile and the Galapagos Islands . The variety known as "Indigo Rose" became available commercially to the agricultural industry and home gardeners in 2012. Investing tomatoes with high anthocyanin content doubles their shelf-life and inhibits growth of

413-477: A lack of contemporaneous evidence. Several theories exist about what Hellenic dialect groups may have existed between the divergence of early Greek-like speech from the common Proto-Indo-European language and the Classical period. They have the same general outline but differ in some of the detail. The only attested dialect from this period is Mycenaean Greek , but its relationship to the historical dialects and

472-419: A lesser degree. Pamphylian Greek , spoken in a small area on the southwestern coast of Anatolia and little preserved in inscriptions, may be either a fifth major dialect group, or it is Mycenaean Greek overlaid by Doric, with a non-Greek native influence. Regarding the speech of the ancient Macedonians diverse theories have been put forward, but the epigraphic activity and the archaeological discoveries in

531-503: A lipophilic colorant for foods. Although anthocyanins have been shown to have antioxidant properties in vitro , there is no evidence for antioxidant effects in humans after consuming foods rich in anthocyanins. Unlike controlled test-tube conditions, the fate of anthocyanins in vivo shows they are poorly conserved (less than 5%), with most of what is absorbed existing as chemically modified metabolites that are excreted rapidly. The increase in antioxidant capacity of blood seen after

590-400: A minor disruption in any of the mechanisms of these enzymes by either genetic or environmental factors, would halt anthocyanin production. While the biological burden of producing anthocyanins is relatively high, plants benefit significantly from the environmental adaptation, disease tolerance, and pest tolerance provided by anthocyanins. In anthocyanin biosynthetic pathway, L -phenylalanine

649-464: A post- harvest mold pathogen , Botrytis cinerea . Some tomatoes also have been modified genetically with transcription factors from snapdragons to produce high levels of anthocyanins in the fruits. Anthocyanins also may be found in naturally ripened olives , and are partly responsible for the red and purple colors of some olives. Content of anthocyanins in the leaves of colorful plant foods such as purple corn, blueberries, or lingonberries ,

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708-550: A prefix /e-/, called the augment . This was probably originally a separate word, meaning something like "then", added because tenses in PIE had primarily aspectual meaning. The augment is added to the indicative of the aorist, imperfect, and pluperfect, but not to any of the other forms of the aorist (no other forms of the imperfect and pluperfect exist). The two kinds of augment in Greek are syllabic and quantitative. The syllabic augment

767-423: A protective role in plants against extreme temperatures. Tomato plants protect against cold stress with anthocyanins countering reactive oxygen species, leading to a lower rate of cell death in leaves. The absorbance pattern responsible for the red color of anthocyanins may be complementary to that of green chlorophyll in photosynthetically active tissues such as young Quercus coccifera leaves. It may protect

826-608: A separate historical stage, though its earliest form closely resembles Attic Greek , and its latest form approaches Medieval Greek . There were several regional dialects of Ancient Greek; Attic Greek developed into Koine. Ancient Greek was a pluricentric language , divided into many dialects. The main dialect groups are Attic and Ionic , Aeolic , Arcadocypriot , and Doric , many of them with several subdivisions. Some dialects are found in standardized literary forms in literature , while others are attested only in inscriptions. There are also several historical forms. Homeric Greek

885-542: A single blooming flower and linear or broadly lanceolate leaves. This is a complex hybridized neo-species, and can also be called Tulipa × gesneriana . Most of the cultivars of tulip are derived from Tulipa gesneriana . It has become naturalised in parts of central and southern Europe and scattered locations in North America . This hybrid is widely believed to have originated in Tian Shan of China, from

944-608: A speculative frenzy now known as tulip mania was triggered between 1634 and 1637. Bulbs were exchanged for land, livestock, and houses, and the Dutch created futures markets where contracts to buy bulbs at the end of the season were bought and sold. A single bulb, the Semper Augustus , fetched 6,000 florins in Haarlem — at that time, a florin could purchase a bushel of wheat. The flower and bulb can cause dermatitis through

1003-630: A standard subject of study in educational institutions of the Western world since the Renaissance . This article primarily contains information about the Epic and Classical periods of the language, which are the best-attested periods and considered most typical of Ancient Greek. From the Hellenistic period ( c.  300 BC ), Ancient Greek was followed by Koine Greek , which is regarded as

1062-510: A vowel or /n s r/ ; final stops were lost, as in γάλα "milk", compared with γάλακτος "of milk" (genitive). Ancient Greek of the classical period also differed in both the inventory and distribution of original PIE phonemes due to numerous sound changes, notably the following: The pronunciation of Ancient Greek was very different from that of Modern Greek . Ancient Greek had long and short vowels ; many diphthongs ; double and single consonants; voiced, voiceless, and aspirated stops ; and

1121-570: Is a literary form of Archaic Greek (derived primarily from Ionic and Aeolic) used in the epic poems , the Iliad and the Odyssey , and in later poems by other authors. Homeric Greek had significant differences in grammar and pronunciation from Classical Attic and other Classical-era dialects. The origins, early form and development of the Hellenic language family are not well understood because of

1180-429: Is about ten times higher than in the edible kernels or fruit. The color spectrum of grape berry leaves may be analysed to evaluate the amount of anthocyanins. Fruit maturity, quality, and harvest time may be evaluated on the basis of the spectrum analysis. The reds, purples, and their blended combinations responsible for autumn foliage are derived from anthocyanins. Unlike carotenoids , anthocyanins are not present in

1239-418: Is added to stems beginning with consonants, and simply prefixes e (stems beginning with r , however, add er ). The quantitative augment is added to stems beginning with vowels, and involves lengthening the vowel: Some verbs augment irregularly; the most common variation is e → ei . The irregularity can be explained diachronically by the loss of s between vowels, or that of the letter w , which affected

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1298-666: Is called 'East Greek'. Arcadocypriot apparently descended more closely from the Mycenaean Greek of the Bronze Age. Boeotian Greek had come under a strong Northwest Greek influence, and can in some respects be considered a transitional dialect, as exemplified in the poems of the Boeotian poet Pindar who wrote in Doric with a small Aeolic admixture. Thessalian likewise had come under Northwest Greek influence, though to

1357-448: Is considered by some linguists to have been closely related to Greek . Among Indo-European branches with living descendants, Greek is often argued to have the closest genetic ties with Armenian (see also Graeco-Armenian ) and Indo-Iranian languages (see Graeco-Aryan ). Ancient Greek differs from Proto-Indo-European (PIE) and other Indo-European languages in certain ways. In phonotactics , ancient Greek words could end only in

1416-588: Is converted to naringenin by phenylalanine ammonialyase, cinnamate 4-hydroxylase, 4-coumarate CoA ligase, chalcone synthase, and chalcone isomerase. Then, the next pathway is catalyzed, resulting in the formation of complex aglycone and anthocyanin through composition by flavanone 3-hydroxylase, flavonoid 3'-hydroxylase, dihydroflavonol 4-reductase, anthocyanidin synthase , UDP-glucoside: flavonoid glucosyltransferase , and methyl transferase . The phenolic metabolic pathways and enzymes may be studied by mean of transgenesis of genes. The Arabidopsis regulatory gene in

1475-410: Is no conclusive evidence that anthocyanins have any effect on human biology or diseases. In flowers, the coloration that is provided by anthocyanin accumulation may attract a wide variety of animal pollinators, while in fruits, the same coloration may aid in seed dispersal by attracting herbivorous animals to the potentially-edible fruits bearing these red, blue, or purple colors. Anthocyanins may have

1534-759: The Greek region of Macedonia during the last decades has brought to light documents, among which the first texts written in Macedonian , such as the Pella curse tablet , as Hatzopoulos and other scholars note. Based on the conclusions drawn by several studies and findings such as Pella curse tablet , Emilio Crespo and other scholars suggest that ancient Macedonian was a Northwest Doric dialect , which shares isoglosses with its neighboring Thessalian dialects spoken in northeastern Thessaly . Some have also suggested an Aeolic Greek classification. The Lesbian dialect

1593-441: The allergen , tuliposide A, even though the bulbs may be consumed with little ill effect. The bulbs may be dried and pulverised and added to cereals or flour. The sweet-scented bisexual flowers appear during April and May. Bulbs are extremely resistant to frost and can tolerate temperatures well below freezing — a period of low temperature is necessary to induce proper growth and flowering, triggered by an increase in sensitivity to

1652-550: The phenylpropanoid pathway. They can occur in all tissues of higher plants, including leaves , stems , roots , flowers , and fruits . Anthocyanins are derived from anthocyanidins by adding sugars. They are odorless and moderately astringent . Although approved as food and beverage colorant in the European Union, anthocyanins are not approved for use as a food additive because they have not been verified as safe when used as food or supplement ingredients. There

1711-501: The present , future , and imperfect are imperfective in aspect; the aorist , present perfect , pluperfect and future perfect are perfective in aspect. Most tenses display all four moods and three voices, although there is no future subjunctive or imperative. Also, there is no imperfect subjunctive, optative or imperative. The infinitives and participles correspond to the finite combinations of tense, aspect, and voice. The indicative of past tenses adds (conceptually, at least)

1770-686: The seed coat of black soybean ( Glycine max L. Merr.) containing approximately 2 g per 100 g, in purple corn kernels and husks , and in the skins and pulp of black chokeberry ( Aronia melanocarpa L.) (see table). Due to critical differences in sample origin, preparation, and extraction methods determining anthocyanin content, the values presented in the adjoining table are not directly comparable. Nature, traditional agriculture methods, and plant breeding have produced various uncommon crops containing anthocyanins, including blue - or red-flesh potatoes and purple or red broccoli, cabbage, cauliflower, carrots, and corn. Garden tomatoes have been subjected to

1829-1031: The 5th century BC. Ancient pronunciation cannot be reconstructed with certainty, but Greek from the period is well documented, and there is little disagreement among linguists as to the general nature of the sounds that the letters represent. /oː/ raised to [uː] , probably by the 4th century BC. Greek, like all of the older Indo-European languages , is highly inflected. It is highly archaic in its preservation of Proto-Indo-European forms. In ancient Greek, nouns (including proper nouns) have five cases ( nominative , genitive , dative , accusative , and vocative ), three genders ( masculine , feminine , and neuter ), and three numbers (singular, dual , and plural ). Verbs have four moods ( indicative , imperative , subjunctive , and optative ) and three voices (active, middle, and passive ), as well as three persons (first, second, and third) and various other forms. Verbs are conjugated through seven combinations of tenses and aspect (generally simply called "tenses"):

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1888-495: The Archaic period of ancient Greek (see Homeric Greek for more details): Μῆνιν ἄειδε, θεά, Πηληϊάδεω Ἀχιλῆος οὐλομένην, ἣ μυρί' Ἀχαιοῖς ἄλγε' ἔθηκε, πολλὰς δ' ἰφθίμους ψυχὰς Ἄϊδι προΐαψεν ἡρώων, αὐτοὺς δὲ ἑλώρια τεῦχε κύνεσσιν οἰωνοῖσί τε πᾶσι· Διὸς δ' ἐτελείετο βουλή· ἐξ οὗ δὴ τὰ πρῶτα διαστήτην ἐρίσαντε Ἀτρεΐδης τε ἄναξ ἀνδρῶν καὶ δῖος Ἀχιλλεύς. The beginning of Apology by Plato exemplifies Attic Greek from

1947-722: The Classical period of ancient Greek. (The second line is the IPA , the third is transliterated into the Latin alphabet using a modern version of the Erasmian scheme .) Ὅτι [hóti Hóti μὲν men mèn ὑμεῖς, hyːmêːs hūmeîs,   Tulipa gesneriana Tulipa gesneriana , the Didier's tulip or garden tulip , is a species of plant in the lily family , cultivated as an ornamental in many countries because of its large, showy flowers. This tall, late-blooming species has

2006-513: The European Union, Australia, and New Zealand, having colorant code E163. In 2013, a panel of scientific experts for the European Food Safety Authority concluded that anthocyanins from various fruits and vegetables have been insufficiently characterized by safety and toxicology studies to approve their use as food additives . Extending from a safe history of using red grape skin extract and blackcurrant extracts to color foods produced in Europe,

2065-471: The anthocyanins. Not all land plants contain anthocyanin; in the Caryophyllales (including cactus , beets , and amaranth ), they are replaced by betalains . Anthocyanins and betalains have never been found in the same plant. Sometimes bred purposely for high anthocyanin content, ornamental plants such as sweet peppers may have unusual culinary and aesthetic appeal. Anthocyanins occur in

2124-550: The aorist. Following Homer 's practice, the augment is sometimes not made in poetry , especially epic poetry. The augment sometimes substitutes for reduplication; see below. Almost all forms of the perfect, pluperfect, and future perfect reduplicate the initial syllable of the verb stem. (A few irregular forms of perfect do not reduplicate, whereas a handful of irregular aorists reduplicate.) The three types of reduplication are: Irregular duplication can be understood diachronically. For example, lambanō (root lab ) has

2183-419: The augment when it was word-initial. In verbs with a preposition as a prefix, the augment is placed not at the start of the word, but between the preposition and the original verb. For example, προσ(-)βάλλω (I attack) goes to προσ έ βαλoν in the aorist. However compound verbs consisting of a prefix that is not a preposition retain the augment at the start of the word: αὐτο(-)μολῶ goes to ηὐ τομόλησα in

2242-438: The center of Greek scholarship, this division of people and language is quite similar to the results of modern archaeological-linguistic investigation. One standard formulation for the dialects is: West vs. non-West Greek is the strongest-marked and earliest division, with non-West in subsets of Ionic-Attic (or Attic-Ionic) and Aeolic vs. Arcadocypriot, or Aeolic and Arcado-Cypriot vs. Ionic-Attic. Often non-West

2301-758: The collections of the sultan of the Ottoman Empire in Istanbul, as is the case with other species of tulips that came into Europe . In 1574, Sultan Selim II ordered the Kadi of A‘azāz in Syria to send him 50,000 tulip bulbs. However, Harvey points out several problems with this source, and there is also the possibility that tulips and hyacinth ( sümbüll ), originally Indian spikenard ( Nardostachys jatamansi ) have been confused. Sultan Selim also imported 300,000 bulbs of Kefe Lale (also known as Cafe-Lale, from

2360-720: The color and stability of anthocyanins. B-ring hydroxylation status and pH have been shown to mediate the degradation of anthocyanins to their phenolic acid and aldehyde constituents. Indeed, significant portions of ingested anthocyanins are likely to degrade to phenolic acids and aldehyde in vivo , following consumption. This characteristic confounds scientific isolation of specific anthocyanin mechanisms in vivo . Anthocyanins generally are degraded at higher pH. However, some anthocyanins, such as petanin (petunidin 3-[6- O -(4- O -( E )- p -coumaroyl- O -α- l -rhamnopyranosyl)-β- d -glucopyranoside]-5- O -β- d -glucopyranoside), are resistant to degradation at pH 8 and may be used effectively as

2419-440: The consumption of anthocyanin-rich foods may not be caused directly by the anthocyanins in the food, but instead by increased uric acid levels derived from metabolizing flavonoids (anthocyanin parent compounds) in the food. It is possible that metabolites of ingested anthocyanins are reabsorbed in the gastrointestinal tract from where they may enter the blood for systemic distribution and have effects as smaller molecules. In

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2478-615: The dialect of Sparta ), and Northern Peloponnesus Doric (including Corinthian ). All the groups were represented by colonies beyond Greece proper as well, and these colonies generally developed local characteristics, often under the influence of settlers or neighbors speaking different Greek dialects. After the conquests of Alexander the Great in the late 4th century BC, a new international dialect known as Koine or Common Greek developed, largely based on Attic Greek , but with influence from other dialects. This dialect slowly replaced most of

2537-536: The fact that they may be produced on flexible substrates, making them amenable to roll-to-roll printing processes. Anthocyanins fluoresce , enabling a tool for plant cell research to allow live cell imaging without a requirement for other fluorophores . Anthocyanin production may be engineered into genetically modified materials to enable their identification visually. Diglucosides: Others glycosides: Ancient Greek language Ancient Greek ( Ἑλληνῐκή , Hellēnikḗ ; [hellɛːnikɛ́ː] ) includes

2596-888: The flowers of many plants, such as the blue poppies of some Meconopsis species and cultivars. Anthocyanins have also been found in various tulip flowers, such as Tulipa gesneriana , Tulipa fosteriana and Tulipa eichleri . Plants rich in anthocyanins are Vaccinium species, such as blueberry , cranberry , and bilberry ; Rubus berries, including black raspberry , red raspberry , and blackberry ; blackcurrant , cherry , eggplant (aubergine) peel , black rice , ube , Okinawan sweet potato , Concord grape , muscadine grape , red cabbage , and violet petals. Red-fleshed peaches and apples contain anthocyanins. Anthocyanins are less abundant in banana , asparagus , pea , fennel , pear , and potato , and may be totally absent in certain cultivars of green gooseberries . The highest recorded amount appears to be specifically in

2655-673: The forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek ( c.  1400–1200 BC ), Dark Ages ( c.  1200–800 BC ), the Archaic or Epic period ( c.  800–500 BC ), and the Classical period ( c.  500–300 BC ). Ancient Greek was the language of Homer and of fifth-century Athenian historians, playwrights, and philosophers . It has contributed many words to English vocabulary and has been

2714-561: The historical Dorians . The invasion is known to have displaced population to the later Attic-Ionic regions, who regarded themselves as descendants of the population displaced by or contending with the Dorians. The Greeks of this period believed there were three major divisions of all Greek people – Dorians, Aeolians, and Ionians (including Athenians), each with their own defining and distinctive dialects. Allowing for their oversight of Arcadian, an obscure mountain dialect, and Cypriot, far from

2773-476: The historical circumstances of the times imply that the overall groups already existed in some form. Scholars assume that major Ancient Greek period dialect groups developed not later than 1120 BC, at the time of the Dorian invasions —and that their first appearances as precise alphabetic writing began in the 8th century BC. The invasion would not be "Dorian" unless the invaders had some cultural relationship to

2832-684: The leaf throughout the growing season, but are produced actively, toward the end of summer. They develop in late summer in the sap of leaf cells, resulting from complex interactions of factors inside and outside the plant. Their formation depends on the breakdown of sugars in the presence of light as the level of phosphate in the leaf is reduced. Orange leaves in autumn result from a combination of anthocyanins and carotenoids. Anthocyanins are present in approximately 10% of tree species in temperate regions, although in certain areas such as New England , up to 70% of tree species may produce anthocyanins. Anthocyanins are approved for use as food colorants in

2891-575: The leaves from attacks by herbivores that may be attracted by green color. Anthocyanins are found in the cell vacuole, mostly in flowers and fruits, but also in leaves, stems, and roots. In these parts, they are found predominantly in outer cell layers such as the epidermis and peripheral mesophyll cells. Most frequently occurring in nature are the glycosides of cyanidin , delphinidin , malvidin , pelargonidin , peonidin , and petunidin . Roughly 2% of all hydrocarbons fixed in photosynthesis are converted into flavonoids and their derivatives, such as

2950-591: The medieval name Kaffa, probably Tulipa schrenkii ) from the port of Kefe in Crimea , for his gardens in the Topkapı Sarayı in Istanbul . They are hybridized with other species present in the collections. Tulipa schrenkii is genetically very closely related to Tulipa gesneriana , and sometimes classified in the same species. Tulipa gesneriana was introduced to western Europe from Constantinople in 1554. It

3009-608: The number at more than 550 different anthocyanins. The difference in chemical structure that occurs in response to changes in pH, is the reason why anthocyanins often are used as pH indicators, as they change from red in acids to blue in bases through a process called halochromism . Anthocyanins are thought to be subject to physiochemical degradation in vivo and in vitro . Structure, pH, temperature, light, oxygen, metal ions, intramolecular association, and intermolecular association with other compounds (copigments, sugars, proteins, degradation products, etc.) generally are known to affect

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3068-508: The older dialects, although the Doric dialect has survived in the Tsakonian language , which is spoken in the region of modern Sparta. Doric has also passed down its aorist terminations into most verbs of Demotic Greek . By about the 6th century AD, the Koine had slowly metamorphosed into Medieval Greek . Phrygian is an extinct Indo-European language of West and Central Anatolia , which

3127-611: The panel concluded that these extract sources were exceptions to the ruling and were sufficiently shown to be safe. Anthocyanin extracts are not specifically listed among approved color additives for foods in the United States; however, grape juice , red grape skin and many fruit and vegetable juices, which are approved for use as colorants, are rich in naturally occurring anthocyanins. No anthocyanin sources are included among approved colorants for drugs or cosmetics . When esterified with fatty acids, anthocyanins can be used as

3186-487: The perfect stem eilēpha (not * lelēpha ) because it was originally slambanō , with perfect seslēpha , becoming eilēpha through compensatory lengthening. Reduplication is also visible in the present tense stems of certain verbs. These stems add a syllable consisting of the root's initial consonant followed by i . A nasal stop appears after the reduplication in some verbs. The earliest extant examples of ancient Greek writing ( c.  1450 BC ) are in

3245-411: The production of anthocyanin pigment 1 ( AtPAP1 ) may be expressed in other plant species. Anthocyanins have been used in organic solar cells because of their ability to convert light energy into electrical energy. The many benefits to using dye-sensitized solar cells instead of traditional p-n junction silicon cells, include lower purity requirements and abundance of component materials, as well as

3304-517: The syllabic script Linear B . Beginning in the 8th century BC, however, the Greek alphabet became standard, albeit with some variation among dialects. Early texts are written in boustrophedon style, but left-to-right became standard during the classic period. Modern editions of ancient Greek texts are usually written with accents and breathing marks , interword spacing , modern punctuation , and sometimes mixed case , but these were all introduced later. The beginning of Homer 's Iliad exemplifies

3363-480: Was Aeolic. For example, fragments of the works of the poet Sappho from the island of Lesbos are in Aeolian. Most of the dialect sub-groups listed above had further subdivisions, generally equivalent to a city-state and its surrounding territory, or to an island. Doric notably had several intermediate divisions as well, into Island Doric (including Cretan Doric ), Southern Peloponnesus Doric (including Laconian ,

3422-529: Was first described in 1559 by Conrad Gesner . When the tulip originally arrived in Europe from the Ottoman Empire, its popularity soared and it quickly became a status symbol for the newly wealthy merchants of the Dutch Golden Age . As a mosaic virus began to infect bulbs, producing rare and spectacular effects in the bloom but weakening and destroying the already limited number of bulbs,

3481-474: Was no evidence generally for consumption of anthocyanin-rich foods having any "antioxidant", " anti-cancer ", " anti-aging ", or "healthy aging" effects. The anthocyanins, anthocyanidins with sugar group(s), are mostly 3- glucosides of the anthocyanidins. The anthocyanins are subdivided into the sugar -free anthocyanidin aglycones and the anthocyanin glycosides. As of 2003, more than 400 anthocyanins had been reported, while later literature in early 2006, puts

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