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63-716: See text The Aizoaceae ( / eɪ z oʊ ˈ eɪ s iː ˌ iː , - s i ˌ aɪ / ), or fig-marigold family , is a large family of dicotyledonous flowering plants containing 135 genera and about 1,800 species . Several genera are commonly known as ' ice plants ' or ' carpet weeds '. The Aizoaceae are also referred to as vygies in South Africa . Some of the unusual Southern African genera—such as Conophytum , Lithops , Titanopsis and Pleiospilos (among others)—resemble gemstones, rocks or pebbles, and are sometimes referred to as ' living stones ' or ' mesembs ' (short for mesembryanthemums ). The family Aizoaceae

126-415: A planetary body after the former is introduced by a space probe or spacecraft , either deliberately or unintentionally. It has also been hypothesized that the origin of life on earth is due to introductions of life from other planets billions of years ago, possibly by a sentient race. Projects have been proposed to introduce life to other lifeless but habitable planets in other star systems some time in

189-412: A family— or whether a described family should be acknowledged— is established and decided upon by active taxonomists . There are not strict regulations for outlining or acknowledging a family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to

252-554: A lack of widespread consensus within the scientific community for extended periods. The continual publication of new data and diverse opinions plays a crucial role in facilitating adjustments and ultimately reaching a consensus over time. The naming of families is codified by various international bodies using the following suffixes: The taxonomic term familia was first used by French botanist Pierre Magnol in his Prodromus historiae generalis plantarum, in quo familiae plantarum per tabulas disponuntur (1689) where he called

315-830: A native of East Asia, has been introduced into parts of Europe for ornamental reasons. Many plants have been introduced with the intent of aesthetically improving public recreation areas or private properties. The introduced Norway maple for example occupies a prominent status in many of Canada's parks. The transport of ornamental plants for landscaping use has and continues to be a source of many introductions. Some of these species have escaped horticultural control and become invasive. Notable examples include water hyacinth , salt cedar , and purple loosestrife . In other cases, species have been translocated for reasons of "cultural nostalgia", which refers to instances in which humans who have migrated to new regions have intentionally brought with them familiar organisms. Famous examples include

378-437: A new region may escape from captive or cultivated populations and subsequently establish independent breeding populations. Escaped organisms are included in this category because their initial transport to a new region is human motivated. The widespread phenomena of intentional introduction has also been described as biological globalization . Positive Introductions Although most introduced species have negative impacts on

441-584: A novel environment (e.g.: in Europe , house sparrows are well established since early Iron Age though they originated from Asia ). Immigrant species are species that travel, sometimes by themselves, but often with human help, between two habitats. Invasiveness is not a requirement. Introduction of a species outside its native range is all that is required to be qualified as an "introduced species". Such species might be termed naturalized , "established", or "wild non-native species". If they further spread beyond

504-463: A parasite or herbivore with it. Some become invasive, for example, the oleander aphid , accidentally introduced with the ornamental plant, oleander . Yet another unintentional pathway of introduction is during the delivery of humanitarian aid in the aftermath of natural disasters. This occurred during relief efforts for Hurricane Maria in Dominica , it was found that the common green iguana ,

567-467: A population decrease. This was seen in the Atlantic Salmon population when high levels of escape from Atlantic Salmon farms into the wild populations resulted in hybrids that had reduced survival. Potential positive effects include adding to the genetic diversity of the population which can increase the adaptation ability of the population and increase the number of healthy individuals within

630-758: A population. This was seen in the introduction of guppies in Trinidad to encourage population growth and introduce new alleles into the population. The results of this introduction included increased levels of heterozygosity and a larger population size . Wide-spread introductions of non-native iguanas are causing devastating effects on native Iguana populations in the Caribbean Lesser Antilles , as hybrids appear to have higher fitness than native iguanas, leading to competitive outcompetition and replacement. Numerous populations have already become extinct and hybridization continues to reduce

693-449: A species is considered "introduced" when its transport into an area outside of its native range is human mediated. Introductions by humans can be described as either intentional or accidental. Intentional introductions have been motivated by individuals or groups who either (1) believe that the newly introduced species will be in some way beneficial to humans in its new location or, (2) species are introduced intentionally but with no regard to

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756-431: Is classified between order and genus . A family may be divided into subfamilies , which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae , but that family is commonly referred to as the "walnut family". The delineation of what constitutes

819-764: Is found in the Greater Cape Floristic Region , which is the most plant-diverse temperate region in the world. A few species are found in Australia and the Central Pacific area. Most fig-marigolds are herbaceous , rarely somewhat woody, with sympodial growth and stems either erect or prostrate. Leaves are simple, opposite or alternate, and more or less succulent with entire (or rarely toothed) margins. Flowers are perfect in most species (but unisexual in some), actinomorphic, and appear singularly or in few-flowered cymes developing from

882-528: Is highly variable. Some have a substantial negative effect on a local ecosystem (in which case they are also classified more specifically as an invasive species ), while other introduced species may have little or no negative impact (no invasiveness), and integrate well into the ecosystem they have been introduced to. Some species have been introduced intentionally to combat pests. They are called biocontrols and may be regarded as beneficial as an alternative to pesticides in agriculture for example. In some instances

945-488: Is likely to cause economic or environmental harm or harm to human health". The biological definition of invasive species, on the other hand, makes no reference to the harm they may cause, only to the fact that they spread beyond the area of original introduction. Some argue that "invasive" is a loaded word and harm is difficult to define. From a regulatory perspective, it is neither desirable nor practical to list as undesirable or outright ban all non-native species (although

1008-493: Is seldom understood in the beginning, and many non-native ornamentals languish in the trade for years before suddenly naturalizing and becoming invasive. Studies have shown that introduced species display a greater likeliness of naturalizing when there is an appropriate environmental match, the plant species are short lived herbs or cultivate from seeds. Peaches , for example, originated in China, and have been carried to much of

1071-575: Is the European rabbit in Australia . Thomas Austin , a British landowner, had rabbits released on his estate in Victoria because he missed hunting them. A more recent example is the introduction of the common wall lizard ( Podarcis muralis) to North America by a Cincinnati boy, George Rau, around 1950 after a family vacation to Italy . Intentional introductions have also been undertaken with

1134-593: Is the seaweed Caulerpa taxifolia . Caulerpa was first observed in the Mediterranean Sea in 1984, off the coast of Monaco . By 1997, it had covered some 50 km . It has a strong potential to overgrow natural biotopes , and represents a major risk for sublittoral ecosystems . The origin of the alga in the Mediterranean was thought to be either as a migration through the Suez Canal from

1197-859: Is the reintroduction of a species that has become locally endangered or extinct, done in the interests of conservation. Examples of successful reintroductions include wolves to Yellowstone National Park in the U.S., and the red kite to parts of England and Scotland. Introductions or translocations of species have also been proposed in the interest of genetic conservation , which advocates the introduction of new individuals into genetically depauperate populations of endangered or threatened species. Unintentional introductions occur when species are transported by human vectors. Increasing rates of human travel are providing accelerating opportunities for species to be accidentally transported into areas in which they are not considered native. For example, three species of rat (the black, Norway and Polynesian) have spread to most of

1260-551: Is the small Indian mongoose ( Urva auropunctata ). Originating in a region encompassing Iran and India , it was introduced to the West Indies and Hawaii in the late 1800s for pest control. Since then, it has thrived on prey unequipped to deal with its speed, nearly leading to the local extinction of a variety of species. In some cases, introduced animals may unintentionally promote the cause of rewilding . For example, escaped horses and donkeys that have gone feral in

1323-470: Is widely recognised by taxonomists. It once went by the botanical name " Ficoidaceae ", now disallowed. The APG II system of 2003 (unchanged from the APG system of 1998) also recognizes the family, and assigns it to the order Caryophyllales in the clade core eudicots . The APG II system also classes the former families Mesembryanthemaceae Fenzl , Sesuviaceae Horan. and Tetragoniaceae Link under

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1386-468: The Genera Plantarum of George Bentham and Joseph Dalton Hooker this word ordo was used for what now is given the rank of family. Families serve as valuable units for evolutionary, paleontological, and genetic studies due to their relatively greater stability compared to lower taxonomic levels like genera and species. Introduced species The impact of introduced species

1449-594: The American bison and the turkey are non-native species to North America. Collectively, non-native crops and livestock account for 98% of US food. These and other benefits from non-natives are so vast that, according to the Congressional Research Service, they probably exceed the costs. Other examples of species introduced for the purposes of benefiting agriculture , aquaculture or other economic activities are widespread. Eurasian carp

1512-684: The Australian Dung Beetle Project in an effort to reduce the impact of livestock manure. The timber industry promoted the introduction of Monterey pine ( Pinus radiata ) from California to Australia and New Zealand as a commercial timber crop. These examples represent only a small subsample of species that have been moved by humans for economic interests. The rise in the use of genetically modified organisms has added another potential economic advantage to introducing new/modified species into different environments. Companies such as Monsanto that earn much of their profit through

1575-553: The Cuban tree frog , and potentially the Venezuela snouted tree frog were introduced with the former two becoming established. Most accidentally or intentionally introduced species do not become invasive as the ones mentioned above. For instance, Some 179 coccinellid species have been introduced to the U.S. and Canada; about 27 of these non-native species have become established, and only a handful can be considered invasive, including

1638-560: The State of Hawaii has adopted an approach that comes close to this). Regulations require a definitional distinction between non-natives that are deemed especially onerous and all others. Introduced "pest" species, that are officially listed as invasive, best fit the definition of an invasive species. Early detection and rapid response is the most effective strategy for regulating a pest species and reducing economic and environmental impacts of an introduction. Management of invasion pathways are on

1701-524: The cane toad and red fox in Australia , nutria in North America , Eurasia , and Africa , and the common brushtail possum in New Zealand . In Taiwan , the success of introduced bird species was related to their native range size and body size; larger species with larger native range sizes were found to have larger introduced range sizes. One notoriously devastating introduced species

1764-573: The Americas may play ecological roles similar to those of the equids that became extinct there at the end of the Pleistocene . The exotic pet trade has also been a large source of introduced species. The species favored as pets have more general habitat requirements and larger distributions. Therefore, as these pets escape or are released, unintentionally or intentionally, they are more likely to survive and establish non-native populations in

1827-697: The Indian subcontinent and is impacting a range of wild species including urban birds. Most introduced species do not become invasive. Examples of introduced animals that have become invasive include the gypsy moth in eastern North America , the zebra mussel and alewife in the Great Lakes , the Canada goose and gray squirrel in Europe, the beaver in Tierra del Fuego , the muskrat in Europe and Asia ,

1890-433: The Mediterranean region, where it can form monocultures that threaten critical conservation habitats. Japanese knotweed grows profusely in many nations. Human beings introduced it into many places in the 19th century. It is a source of resveratrol , a dietary supplement. It can grow in building foundations, threatening their stability, and spreads quite quickly. Mesquite ( Prosopis juliflora ) has spread very widely across

1953-681: The Red Sea, or as an accidental introduction from an aquarium. Another troublesome plant species is the terrestrial plant Phyla canescens , which was intentionally introduced into many countries in North America, Europe, and Africa as an ornamental plant. This species has become invasive in Australia, where it threatens native rare plants and causes erosion and soil slumping around river banks. It has also become invasive in France where it has been listed as an invasive plant species of concern in

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2016-436: The aim of ameliorating environmental problems. A number of fast spreading plants such as kudzu have been introduced as a means of erosion control. Other species have been introduced as biological control agents to control invasive species . This involves the purposeful introduction of a natural enemy of the target species with the intention of reducing its numbers or controlling its spread. A special case of introduction

2079-532: The broadest and most widely used sense, an introduced species is synonymous with "non-native" and therefore applies as well to most garden and farm organisms; these adequately fit the basic definition given above. However, some sources add to that basic definition "and are now reproducing in the wild", which means that species growing in a garden, farm, or house may not meet the criteria unless they escape and persist. There are many terms associated with introduced species that represent subsets of introduced species, and

2142-539: The economy. There have been calls from scientists to consider a species "invasive" only in terms of their spread and reproduction rather than the harm they may cause. According to a practical definition, an invasive species is one that has been introduced and become a pest in its new location, spreading (invading) by natural means. The term is used to imply both a sense of urgency and actual or potential harm. For example, U.S. Executive Order 13112 (1999) defines "invasive species" as "an alien species whose introduction does or

2205-585: The ecosystems they enter into, there are still some species that have affected the ecosystem in a positive way. For example, in New Hampshire invasive plants can provide some benefits to some species. Invasive species such as autumn olive, oriental bittersweet, and honeysuckle produce fruit that is used by a handful of fruit-eating bird species. The invasive plants can also be a source of pollen and nectar for many insects, such as bees. These invasive plants were able to help their ecosystem thriving, and increase

2268-528: The family Aizoaceae . The common Afrikaans name "vygie" meaning "small fig" refers to the fruiting capsule , which resembles the true fig. Glistening epidermal bladder cells give the family its common name "ice plants". Most species (96%, 1782 species in 132 genera) in this family are endemic to arid or semiarid parts of Southern Africa in the Succulent Karoo . Much of the Aizoaceae's diversity

2331-540: The family as a rank intermediate between order and genus was introduced by Pierre André Latreille in his Précis des caractères génériques des insectes, disposés dans un ordre naturel (1796). He used families (some of them were not named) in some but not in all his orders of "insects" (which then included all arthropods ). In nineteenth-century works such as the Prodromus of Augustin Pyramus de Candolle and

2394-473: The forefront of eliminating unwanted invasive species this would include preliminary steps; educating the public, cooperation from industries and government resources. In Great Britain , the Wildlife and Countryside Act 1981 prevents the introduction of any animal not naturally occurring in the wild or any of a list of both animals or plants introduced previously and proved to be invasive. By definition ,

2457-586: The intentionally introduced Harmonia axyridis , multicolored Asian lady beetle . However the small percentage of introduced species that become invasive can produce profound ecological changes. In North America, Harmonia axyridis has become the most abundant lady beetle and probably accounts for more observations than all the native lady beetles put together. Many non-native plants have been introduced into new territories, initially as either ornamental plants or for erosion control, stock feed, or forestry. Whether an exotic will become an invasive species

2520-849: The introduction of common starlings to North America by the American Eugene Schieffelin , a lover of the works of Shakespeare and the chairman of the American Acclimatization Society , who, it is rumoured, wanted to introduce all of the birds mentioned in Shakespeare's plays into the United States. He deliberately released eighty starlings into Central Park in New York City in 1890, and another forty in 1891. Yet another prominent example of an introduced species that became invasive

2583-489: The invasive comb jelly Mnemiopsis leidyi , the dangerous bacterium Vibrio cholerae , or the fouling zebra mussel . The Mediterranean and Black Seas, with their high volume shipping from exotic sources, are most impacted by this problem. Busy harbors are all potential hotspots as well: over 200 species have been introduced to the San Francisco Bay in this manner making it the most heavily invaded estuary in

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2646-403: The leaf axils. Sepals are typically five (3–8) and more or less connate (fused) below. True petals are absent. However, some species have numerous linear petals derived from staminodes. The seed capsules have one to numerous seeds per cell and are often hygrochastic , dispersing seeds by "jet action" when wet. The radiation of the Aizoaceae, specifically the subfamily Ruschioideae, was one of

2709-465: The most recent among the angiosperms , occurring 1.13–6.49 Mya. It is also one of the fastest radiations ever described in the angiosperms, with a diversification rate of about 4.4 species per million years. This diversification was roughly contemporaneous with major radiations in two other succulent lineages, Cactaceae and Agave . The family includes many species that use crassulacean acid metabolism as pathway for carbon fixation . Some species in

2772-547: The native animal's chances of survival. Several introduced exotic trees served as nest sites for resident waterbird species in Udaipur city, India. Perhaps the most common motivation for introducing a species into a new place is that of economic gain. Non-native species can become such a common part of an environment, culture, and even diet that little thought is given to their geographic origin. For example, soybeans , kiwi fruit , wheat , honey bees , and all livestock except

2835-459: The new environment. Adventive species are often considered synonymous with "introduced species", but this term is sometimes applied exclusively to introduced species that are not permanently established. Naturalized species are often introduced species that do not need human help to reproduce and maintain their population in an area outside their native range (no longer adventive), but that also applies to populations migrating and establishing in

2898-517: The number of native iguanas on multiple islands. In plants, introduced species have been observed to undergo rapid evolutionary change to adapt to their new environments, with changes in plant height, size, leaf shape, dispersal ability, reproductive output, vegetative reproduction ability, level of dependence on the mycorrhizal network , and level of phenotype plasticity appearing on timescales of decades to centuries. It has been hypothesized that invasive species of microbial life could contaminate

2961-431: The place of introduction and cause damage to nearby species, they are called " invasive species ". The transition from introduction, to establishment and to invasion has been described in the context of plants. Introduced species are essentially "non-native" species. Invasive species are those introduced species that spread widely or quickly and cause harm, be that to the environment, human health, other valued resources, or

3024-778: The populated world. Tomatoes are native to the Andes . Squash ( pumpkins ), maize (corn), and tobacco are native to the Americas , but were introduced to the Old World . Many introduced species require continued human intervention to survive in the new environment. Others may become feral, but do not seriously compete with natives, but simply increase the biodiversity of the area. One example would be Dandelions in North America , which have become an essential source of early season nectar for both native and introduced pollinators, and do not meaningfully compete with native grasses or flowers. A very troublesome marine species in southern Europe

3087-572: The potential for being beneficial or detrimental in the long run remains unknown. The effects of introduced species on natural environments have gained much scrutiny from scientists, governments, farmers and others. The formal definition of an introduced species from the United States Environmental Protection Agency is "A species that has been intentionally or inadvertently brought into a region or area. Also called an exotic or non-native species". In

3150-436: The potential impact. Unintentional or accidental introductions are most often a byproduct of human movements and are thus unbound to human motivations. Subsequent range expansion of introduced species may or may not involve human activity. Species that humans intentionally transport to new regions can subsequently become successfully established in two ways. In the first case, organisms are purposely released for establishment in

3213-464: The purposes of sport fishing and hunting. The introduced amphibian ( Ambystoma tigrinum ) that threatens the endemic California salamander ( A. californiense ) was introduced to California as a source of bait for fishermen. Pet animals have also been frequently transported into new areas by humans, and their escapes have resulted in several introductions, such as feral cats , parrots , and pond slider . Lophura nycthemera ( silver pheasant ),

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3276-519: The selling of genetically modified seeds has added to the controversy surrounding introduced species. The effect of genetically modified organisms varies from organism to organism and is still being researched today, however, the rise of genetically modified organisms has added complexity to the conversations surrounding introduced species. Introductions have also been important in supporting recreation activities or otherwise increasing human enjoyment. Numerous fish and game animals have been introduced for

3339-575: The seventy-six groups of plants he recognised in his tables families ( familiae ). The concept of rank at that time was not yet settled, and in the preface to the Prodromus Magnol spoke of uniting his families into larger genera , which is far from how the term is used today. In his work Philosophia Botanica published in 1751, Carl Linnaeus employed the term familia to categorize significant plant groups such as trees , herbs , ferns , palms , and so on. Notably, he restricted

3402-584: The subfamily Sesuvioideae instead use C 4 carbon fixation , which might have evolved multiple times in the group. Because of the hyperdiversity of the Aizoaceae and the young age of the clade, many generic and species boundaries are uncertain. Genera: Genera: Genera: Genera: * no longer recognised This subfamily includes a number of C 4 species. Genera: Include; Several genera are cultivated. Lithops , or "living stones", are popular as novelty house plants because of their stone-like appearance. Some species are edible, including: C. edulis

3465-628: The terminology associated with introduced species is now in flux for various reasons. Examples of these terms are "invasive", "acclimatized", "adventive", "naturalized", and "immigrant" species. The term "invasive" is used to describe introduced species that cause ecological, economic, or other damage to the area in which they were introduced. Acclimatized species are introduced species that have changed physically and/or behaviorally in order to adjust to their new environment. Acclimatized species are not necessarily optimally adjusted to their new environment and may just be physically/behaviorally sufficient for

3528-549: The use of this term solely within the book's morphological section, where he delved into discussions regarding the vegetative and generative aspects of plants. Subsequently, in French botanical publications, from Michel Adanson 's Familles naturelles des plantes (1763) and until the end of the 19th century, the word famille was used as a French equivalent of the Latin ordo (or ordo naturalis ). In zoology ,

3591-619: The wild. Among the popular exotic pets that have become alien or invasive species are parrots, frogs, terrapins, and iguanas. Some species, such as the Western honey bee , brown rat , house sparrow , ring-necked pheasant , and European starling , have been introduced very widely. In addition there are some agricultural and pet species that frequently become feral ; these include rabbits , dogs , ducks , snakes , goats , fish , pigs , and cats . Many water fleas such as Daphnia , Bosmina and Bythotrephes have introduced around

3654-423: The wild. It is sometimes difficult to predict whether a species will become established upon release, and if not initially successful, humans have made repeated introductions to improve the probability that the species will survive and eventually reproduce in the wild. In these cases, it is clear that the introduction is directly facilitated by human desires. In the second case, species intentionally transported into

3717-427: The world as hitchhikers on ships, and arachnids such as scorpions and exotic spiders are sometimes transported to areas far beyond their native range by riding in shipments of tropical fruit. This was seen during the introduction of Steatoda nobilis (Noble false widow) worldwide through banana shipments. Further there are numerous examples of marine organisms being transported in ballast water , among them

3780-431: The world, causing dramatic changes in native freshwater ecosystems. When a new species is introduced, the species could potentially breed with members of native species, producing hybrids. The effect of the creating of hybrids can range from having little effect, a negative effect, to having devastating effects on native species. Potential negative effects include hybrids that are less fit for their environment resulting in

3843-603: The world. There is also the accidental release of the Africanized honey bees (AHB), known colloquially as "killer bees") or Africanized bee to Brazil in 1957 and the Asian carp to the United States. The insect commonly known as the brown marmorated stink bug ( Halyomorpha halys ) was introduced accidentally in Pennsylvania. Another form of unintentional introductions is when an intentionally introduced plant carries

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3906-460: Was introduced to California in the early 1900s to stabilize soil along railroad tracks and has become invasive. In southern California, ice plants are sometimes used as firewalls; however, they do burn if not carefully maintained. Family (biology) Family ( Latin : familia , pl. : familiae ) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy . It

3969-534: Was first introduced to the United States as a potential food source. The apple snail was released in Southeast Asia with the intent that it be used as a protein source, and subsequently to places like Hawaii to establish a food industry. In Alaska, foxes were introduced to many islands to create new populations for the fur trade. About twenty species of African and European dung beetles have established themselves in Australia after deliberate introduction by

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