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Convention on Biological Diversity

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A multilateral treaty or multilateral agreement is a treaty to which two or more sovereign states are parties. Each party owes the same obligations to all other parties, except to the extent that they have stated reservations . Examples of multilateral treaties include the Convention Relating to the Status of Refugees , the United Nations Convention on the Law of the Sea , the Geneva Conventions , and the Rome Statute of the International Criminal Court .

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104-474: The Convention on Biological Diversity ( CBD ), known informally as the Biodiversity Convention , is a multilateral treaty . The Convention has three main goals: the conservation of biological diversity (or biodiversity ); the sustainable use of its components; and the fair and equitable sharing of benefits arising from genetic resources . Its objective is to develop national strategies for

208-409: A cell wall . Newly dead animals may be covered by an exoskeleton . Fragmentation processes, which break through these protective layers, accelerate the rate of microbial decomposition. Animals fragment detritus as they hunt for food, as does passage through the gut. Freeze-thaw cycles and cycles of wetting and drying also fragment dead material. The chemical alteration of the dead organic matter

312-495: A food chain . Real systems are much more complex than this—organisms will generally feed on more than one form of food, and may feed at more than one trophic level. Carnivores may capture some prey that is part of a plant-based trophic system and others that are part of a detritus-based trophic system (a bird that feeds both on herbivorous grasshoppers and earthworms, which consume detritus). Real systems, with all these complexities, form food webs rather than food chains which present

416-619: A habitat . Ecosystem ecology is the "study of the interactions between organisms and their environment as an integrated system". The size of ecosystems can range up to ten orders of magnitude , from the surface layers of rocks to the surface of the planet. The Hubbard Brook Ecosystem Study started in 1963 to study the White Mountains in New Hampshire . It was the first successful attempt to study an entire watershed as an ecosystem. The study used stream chemistry as

520-609: A central role over a wide range, for example, in the slow development of soil from bare rock and the faster recovery of a community from disturbance . Disturbance also plays an important role in ecological processes. F. Stuart Chapin and coauthors define disturbance as "a relatively discrete event in time that removes plant biomass". This can range from herbivore outbreaks, treefalls, fires, hurricanes, floods, glacial advances , to volcanic eruptions . Such disturbances can cause large changes in plant, animal and microbe populations, as well as soil organic matter content. Disturbance

624-507: A critical role in global nutrient cycling and ecosystem function. Phosphorus enters ecosystems through weathering . As ecosystems age this supply diminishes, making phosphorus-limitation more common in older landscapes (especially in the tropics). Calcium and sulfur are also produced by weathering, but acid deposition is an important source of sulfur in many ecosystems. Although magnesium and manganese are produced by weathering, exchanges between soil organic matter and living cells account for

728-517: A faster recovery. More severe and more frequent disturbance result in longer recovery times. From one year to another, ecosystems experience variation in their biotic and abiotic environments. A drought , a colder than usual winter, and a pest outbreak all are short-term variability in environmental conditions. Animal populations vary from year to year, building up during resource-rich periods and crashing as they overshoot their food supply. Longer-term changes also shape ecosystem processes. For example,

832-427: A function-based typology has been proposed to leverage the strengths of these different approaches into a unified system. Human activities are important in almost all ecosystems. Although humans exist and operate within ecosystems, their cumulative effects are large enough to influence external factors like climate. Ecosystems provide a variety of goods and services upon which people depend. Ecosystem goods include

936-698: A general level, for example, tropical forests , temperate grasslands , and arctic tundra . There can be any degree of subcategories among ecosystem types that comprise a biome, e.g., needle-leafed boreal forests or wet tropical forests. Although ecosystems are most commonly categorized by their structure and geography, there are also other ways to categorize and classify ecosystems such as by their level of human impact (see anthropogenic biome ), or by their integration with social processes or technological processes or their novelty (e.g. novel ecosystem ). Each of these taxonomies of ecosystems tends to emphasize different structural or functional properties. None of these

1040-672: A means of monitoring ecosystem properties, and developed a detailed biogeochemical model of the ecosystem. Long-term research at the site led to the discovery of acid rain in North America in 1972. Researchers documented the depletion of soil cations (especially calcium) over the next several decades. Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Studies can be carried out at

1144-528: A more important role in moving nutrients around. This can be especially important as the soil thaws in the spring, creating a pulse of nutrients that become available. Decomposition rates are low under very wet or very dry conditions. Decomposition rates are highest in wet, moist conditions with adequate levels of oxygen. Wet soils tend to become deficient in oxygen (this is especially true in wetlands ), which slows microbial growth. In dry soils, decomposition slows as well, but bacteria continue to grow (albeit at

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1248-406: A number of common, non random properties in the topology of their network. The carbon and nutrients in dead organic matter are broken down by a group of processes known as decomposition. This releases nutrients that can then be re-used for plant and microbial production and returns carbon dioxide to the atmosphere (or water) where it can be used for photosynthesis. In the absence of decomposition,

1352-421: A process known as denitrification . Mycorrhizal fungi which are symbiotic with plant roots, use carbohydrates supplied by the plants and in return transfer phosphorus and nitrogen compounds back to the plant roots. This is an important pathway of organic nitrogen transfer from dead organic matter to plants. This mechanism may contribute to more than 70 Tg of annually assimilated plant nitrogen, thereby playing

1456-561: A signatory who had not yet ratified the treaty by 2010, produced one of the most thorough implementation programs through species recovery programs and other mechanisms long in place in the US for species conservation. Singapore established a detailed National Biodiversity Strategy and Action Plan . The National Biodiversity Centre of Singapore represents Singapore in the Convention for Biological Diversity. In accordance with Article 26 of

1560-696: A significant portion of ecosystem fluxes. Potassium is primarily cycled between living cells and soil organic matter. Biodiversity plays an important role in ecosystem functioning. Ecosystem processes are driven by the species in an ecosystem, the nature of the individual species, and the relative abundance of organisms among these species. Ecosystem processes are the net effect of the actions of individual organisms as they interact with their environment. Ecological theory suggests that in order to coexist, species must have some level of limiting similarity —they must be different from one another in some fundamental way, otherwise, one species would competitively exclude

1664-408: A slower rate) even after soils become too dry to support plant growth. Ecosystems are dynamic entities. They are subject to periodic disturbances and are always in the process of recovering from past disturbances. When a perturbation occurs, an ecosystem responds by moving away from its initial state. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance,

1768-464: A small effect on ecosystem function. Ecologically distinct species, on the other hand, have a much larger effect. Similarly, dominant species have a large effect on ecosystem function, while rare species tend to have a small effect. Keystone species tend to have an effect on ecosystem function that is disproportionate to their abundance in an ecosystem. An ecosystem engineer is any organism that creates, significantly modifies, maintains or destroys

1872-468: A variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Biomes are general classes or categories of ecosystems. However, there is no clear distinction between biomes and ecosystems. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of

1976-443: A variety of goods and services upon which people depend, and may be part of. Ecosystem goods include the "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like the maintenance of hydrological cycles , cleaning air and water,

2080-522: A variety of scales, ranging from whole-ecosystem studies to studying microcosms or mesocosms (simplified representations of ecosystems). American ecologist Stephen R. Carpenter has argued that microcosm experiments can be "irrelevant and diversionary" if they are not carried out in conjunction with field studies done at the ecosystem scale. In such cases, microcosm experiments may fail to accurately predict ecosystem-level dynamics. Biomes are general classes or categories of ecosystems. However, there

2184-431: Is a system that environments and their organisms form through their interaction. The biotic and abiotic components are linked together through nutrient cycles and energy flows. Ecosystems are controlled by external and internal factors . External factors such as climate , parent material which forms the soil and topography , control the overall structure of an ecosystem but are not themselves influenced by

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2288-400: Is a treaty between two states. A bilateral treaty may become a multilateral treaty when additional new parties succeed or accede to it. Pope Francis argues in his encyclical letter Fratelli tutti (2020) that "preference should be given to multilateral agreements between states, because, more than bilateral agreements, they guarantee the promotion of a truly universal common good and

2392-525: Is also linkage with the EU Marine Strategy Framework Directive . There have been criticisms against CBD that its implementation has been weakened due to resistance of Western countries to the implementation of pro-South provisions of the Convention. CBD is also regarded as a case of a hard treaty gone soft in the implementation trajectory. The argument to enforce the treaty as a legally binding multilateral instrument with

2496-458: Is an international synthesis by over 1000 of the world's leading biological scientists that analyzes the state of the Earth's ecosystems and provides summaries and guidelines for decision-makers. The report identified four major categories of ecosystem services: provisioning, regulating, cultural and supporting services. It concludes that human activity is having a significant and escalating impact on

2600-442: Is consumed by animals while still alive and enters the plant-based trophic system. After plants and animals die, the organic matter contained in them enters the detritus-based trophic system. Ecosystem respiration is the sum of respiration by all living organisms (plants, animals, and decomposers) in the ecosystem. Net ecosystem production is the difference between gross primary production (GPP) and ecosystem respiration. In

2704-569: Is followed by succession, a "directional change in ecosystem structure and functioning resulting from biotically driven changes in resource supply." The frequency and severity of disturbance determine the way it affects ecosystem function. A major disturbance like a volcanic eruption or glacial advance and retreat leave behind soils that lack plants, animals or organic matter. Ecosystems that experience such disturbances undergo primary succession . A less severe disturbance like forest fires, hurricanes or cultivation result in secondary succession and

2808-554: Is governed by three sets of factors—the physical environment (temperature, moisture, and soil properties), the quantity and quality of the dead material available to decomposers, and the nature of the microbial community itself. Temperature controls the rate of microbial respiration; the higher the temperature, the faster the microbial decomposition occurs. Temperature also affects soil moisture, which affects decomposition. Freeze-thaw cycles also affect decomposition—freezing temperatures kill soil microorganisms, which allows leaching to play

2912-405: Is no clear distinction between biomes and ecosystems. Biomes are always defined at a very general level. Ecosystems can be described at levels that range from very general (in which case the names are sometimes the same as those of biomes) to very specific, such as "wet coastal needle-leafed forests". Biomes vary due to global variations in climate . Biomes are often defined by their structure: at

3016-531: Is primarily achieved through bacterial and fungal action. Fungal hyphae produce enzymes that can break through the tough outer structures surrounding dead plant material. They also produce enzymes that break down lignin , which allows them access to both cell contents and the nitrogen in the lignin. Fungi can transfer carbon and nitrogen through their hyphal networks and thus, unlike bacteria, are not dependent solely on locally available resources. Decomposition rates vary among ecosystems. The rate of decomposition

3120-485: Is termed its resistance . The capacity of a system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Resilience thinking also includes humanity as an integral part of the biosphere where we are dependent on ecosystem services for our survival and must build and maintain their natural capacities to withstand shocks and disturbances. Time plays

3224-422: Is the "best" classification. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of the definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Different approaches to ecological classifications have been developed in terrestrial, freshwater and marine disciplines, and

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3328-556: Is to create an international legally binding instrument (ILBI) involving area-based planning and decision-making under UNCLOS to support the conservation and sustainable use of marine biological diversity beyond areas of national jurisdiction (BBNJ treaty or High Seas Treaty ). The notion of an international convention on biodiversity was conceived at a United Nations Environment Programme (UNEP) Ad Hoc Working Group of Experts on Biological Diversity in November 1988. The subsequent year,

3432-579: The Cartagena Protocol and Nagoya Protocol . The Cartagena Protocol on Biosafety to the Convention on Biological Diversity is an international treaty governing the movements of living modified organisms (LMOs) resulting from modern biotechnology from one country to another. It was adopted on 29 January 2000 as a supplementary agreement to the CBD and entered into force on 11 September 2003. The Nagoya Protocol on Access to Genetic Resources and

3536-645: The Cook Islands , Niue , and the State of Palestine . The Holy See and the states with limited recognition are non-Parties. The US has signed but not ratified the treaty, because ratification requires a two-thirds majority in the Senate and is blocked by Republican Party senators. The European Union created the Cartagena Protocol (see below) in 2000 to enhance biosafety regulation and propagate

3640-477: The precautionary principle and allow developing nations to balance public health against economic benefits. It will, for example, let countries ban imports of a genetically modified organism if they feel there is not enough scientific evidence the product is safe and requires exporters to label shipments containing genetically modified commodities such as corn or cotton. The required number of 50 instruments of ratification/accession/approval/acceptance by countries

3744-588: The precautionary principle which demands that where there is a threat of significant reduction or loss of biological diversity, lack of full scientific certainty should not be used as a reason for postponing measures to avoid or minimize such a threat. The Convention acknowledges that substantial investments are required to conserve biological diversity . It argues, however, that conservation will bring us significant environmental, economic and social benefits in return. The Convention on Biological Diversity of 2010 banned some forms of geoengineering . As of April 2024,

3848-555: The resource inputs are generally controlled by external processes like climate and parent material, the availability of these resources within the ecosystem is controlled by internal factors like decomposition, root competition or shading. Other factors like disturbance, succession or the types of species present are also internal factors. Primary production is the production of organic matter from inorganic carbon sources. This mainly occurs through photosynthesis . The energy incorporated through this process supports life on earth, while

3952-609: The "precautionary principle" over the "sound science principle" defended by the United States. Whereas the impact of the Cartagena Protocol on domestic regulations has been substantial, its impact on international trade law remains uncertain. In 2006, the World Trade Organization (WTO) ruled that the European Union had violated international trade law between 1999 and 2003 by imposing a moratorium on

4056-435: The "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . They also include less tangible items like tourism and recreation, and genes from wild plants and animals that can be used to improve domestic species. Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like

4160-501: The Ad Hoc Working Group of Technical and Legal Experts was established for the drafting of a legal text which addressed the conservation and sustainable use of biological diversity, as well as the sharing of benefits arising from their utilization with sovereign states and local communities. In 1991, an intergovernmental negotiating committee was established, tasked with finalizing the Convention's text. A Conference for

4264-843: The Adoption of the Agreed Text of the Convention on Biological Diversity was held in Nairobi, Kenya, in 1992, and its conclusions were distilled in the Nairobi Final Act. The Convention's text was opened for signature on 5 June 1992 at the United Nations Conference on Environment and Development (the Rio "Earth Summit"). By its closing date, 4 June 1993, the Convention had received 168 signatures. It entered into force on 29 December 1993. The Convention recognized for

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4368-704: The Aichi Biodiversity Targets. The meetings of the Parties to the Convention are known as Conferences of the Parties (COP), with the first one (COP 1) held in Nassau, Bahamas , in 1994 and the most recent one (COP 16) in 2024 in Cali , Colombia . In the area of marine and coastal biodiversity CBD's focus at present is to identify Ecologically or Biologically Significant Marine Areas (EBSAs) in specific ocean locations based on scientific criteria. The aim

4472-885: The Conference of Parties reviewing the infractions and non-compliance is also gaining strength. Although the Convention explicitly states that all forms of life are covered by its provisions, examination of reports and of national biodiversity strategies and action plans submitted by participating countries shows that in practice this is not happening. The fifth report of the European Union, for example, makes frequent reference to animals (particularly fish) and plants, but does not mention bacteria , fungi or protists at all. The International Society for Fungal Conservation has assessed more than 100 of these CBD documents for their coverage of fungi using defined criteria to place each in one of six categories. No documents were assessed as good or adequate, less than 10% as nearly adequate or poor, and

4576-619: The Conference of the Parties, which may be endorsed in whole, in part or in modified form by the COPs. As of 2024 SBSTTA had met 26 times, with a 26th meeting taking place in Nairobi, Kenya in 2024. In 2014, the Conference of the Parties to the Convention on Biological Diversity established the Subsidiary Body on Implementation (SBI) to replace the Ad Hoc Open-ended Working Group on Review of Implementation of

4680-452: The Convention at the national level. The Convention requires that countries prepare a national biodiversity strategy and to ensure that this strategy is included in planning for activities in all sectors where diversity may be impacted. As of early 2012, 173 Parties had developed NBSAPs. The United Kingdom, New Zealand and Tanzania carried out elaborate responses to conserve individual species and specific habitats. The United States of America,

4784-548: The Convention include: The Convention's governing body is the Conference of the Parties (COP), consisting of all governments (and regional economic integration organizations) that have ratified the treaty. This ultimate authority reviews progress under the Convention, identifies new priorities, and sets work plans for members. The COP can also make amendments to the Convention, create expert advisory bodies, review progress reports by member nations, and collaborate with other international organizations and agreements. The Conference of

4888-589: The Convention, Parties prepare national reports on the status of implementation of the Convention. The Cartagena Protocol on Biosafety, also known as the Biosafety Protocol, was adopted in January 2000, after a CBD Open-ended Ad Hoc Working Group on Biosafety had met six times between July 1996 and February 1999. The Working Group submitted a draft text of the Protocol for consideration by Conference of

4992-582: The Convention. The four functions and core areas of work of SBI are: (a) review of progress in implementation; (b) strategic actions to enhance implementation; (c) strengthening means of implementation; and (d) operations of the Convention and the Protocols. The first meeting of the SBI was held on 2–6 May 2016 and the second meeting was held on 9–13 July 2018, both in Montreal, Canada. The latest (fifth) meeting of

5096-559: The EBSA process based on internationally agreed scientific criteria. This is aimed at creating an international legally binding instrument (ILBI) under UNCLOS to support the conservation and sustainable use of marine biological diversity beyond areas of national jurisdiction (BBNJ or High Seas Treaty ). The central mechanism is area-based planning and decision-making. It integrates EBSAs, Vulnerable Marine Ecosystems (VMEs) and High Seas ( Marine Protected Areas ) with Blue Growth scenarios. There

5200-503: The Fair and Equitable Sharing of Benefits Arising from their Utilization (ABS) to the Convention on Biological Diversity is another supplementary agreement to the CBD. It provides a transparent legal framework for the effective implementation of one of the three objectives of the CBD: the fair and equitable sharing of benefits arising out of the utilization of genetic resources. The Nagoya Protocol

5304-470: The Fair and Equitable Sharing of Benefits Arising from their Utilization to the Convention on Biological Diversity was adopted on 29 October 2010 in Nagoya, Aichi Prefecture , Japan, at the tenth meeting of the Conference of the Parties , and entered into force on 12 October 2014. The protocol is a supplementary agreement to the Convention on Biological Diversity, and provides a transparent legal framework for

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5408-503: The Parties at its first extraordinary meeting, which was convened for the express purpose of adopting a protocol on biosafety to the Convention on Biological Diversity. After a few delays, the Cartagena Protocol was eventually adopted on 29 January 2000. The Biosafety Protocol seeks to protect biological diversity from the potential risks posed by living modified organisms resulting from modern biotechnology. The Biosafety Protocol makes clear that products from new technologies must be based on

5512-758: The Parties to the Convention took place in May 1998, in Bratislava , Slovakia . The First Extraordinary Meeting of the Conference of the Parties took place in February 1999, in Cartagena, Colombia . A series of meetings led to the adoption of the Cartagena Protocol on Biosafety in January 2000, effective from 2003. The fifth ordinary meeting of the Parties to the Convention took place in May 2000, in Nairobi , Kenya . Multilateral treaty A bilateral treaty

5616-609: The Parties to the Convention took place in November and December 1994, in Nassau, Bahamas . The International Coral Reef Initiative (ICRI) was launched at this first COP for the Convention on Biological Diversity. The second ordinary meeting of the Parties to the Convention took place in November 1995, in Jakarta , Indonesia . The third ordinary meeting of the Parties to the Convention took place in November 1996, in Buenos Aires , Argentina . The fourth ordinary meeting of

5720-562: The Parties uses expertise and support from several other bodies that are established by the Convention. In addition to committees or mechanisms established on an ad hoc basis, the main organs are: The CBD Secretariat, based in Montreal , Quebec, Canada, operates under UNEP, the United Nations Environment Programme . Its main functions are to organize meetings, draft documents, assist member governments in

5824-715: The SBI was held in October 2024 in Cali, Colombia. The Bureau of the Conference of the Parties serves as the Bureau of the SBI. The current chair of the SBI is Ms. Clarissa Souza Della Nina of Brazil . As of 2016, the Convention has 196 Parties, which includes 195 states and the European Union . All UN member states—with the exception of the United States—have ratified the treaty. Non-UN member states that have ratified are

5928-481: The absence of disturbance, net ecosystem production is equivalent to the net carbon accumulation in the ecosystem. Energy can also be released from an ecosystem through disturbances such as wildfire or transferred to other ecosystems (e.g., from a forest to a stream to a lake) by erosion . In aquatic systems , the proportion of plant biomass that gets consumed by herbivores is much higher than in terrestrial systems. In trophic systems, photosynthetic organisms are

6032-412: The acting executive secretary is Astrid Schomaker. The previous executive secretaries were: David Cooper (2023-2024), Elizabeth Maruma Mrema (2020-2023), pl:Cristiana Pașca Palmer (2017–2019), Braulio Ferreira de Souza Dias (2012–2017), Ahmed Djoghlaf (2006–2012), Hamdallah Zedan (1998–2005), Calestous Juma (1995–1998), and Angela Cropper (1993–1995). Some of the many issues dealt with under

6136-402: The amount of energy available to the ecosystem. Parent material determines the nature of the soil in an ecosystem, and influences the supply of mineral nutrients. Topography also controls ecosystem processes by affecting things like microclimate , soil development and the movement of water through a system. For example, ecosystems can be quite different if situated in a small depression on

6240-504: The amount of light available, the amount of leaf area a plant has to capture light (shading by other plants is a major limitation of photosynthesis), the rate at which carbon dioxide can be supplied to the chloroplasts to support photosynthesis, the availability of water, and the availability of suitable temperatures for carrying out photosynthesis. Energy and carbon enter ecosystems through photosynthesis, are incorporated into living tissue, transferred to other organisms that feed on

6344-401: The approval of genetically modified organisms (GMO) imports. Disappointing the United States, the panel nevertheless "decided not to decide" by not invalidating the stringent European biosafety regulations. Implementation by the Parties to the Convention is achieved using two means: National Biodiversity Strategies and Action Plans ( NBSAP ) are the principal instruments for implementing

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6448-408: The carbon makes up much of the organic matter in living and dead biomass, soil carbon and fossil fuels . It also drives the carbon cycle , which influences global climate via the greenhouse effect . Through the process of photosynthesis, plants capture energy from light and use it to combine carbon dioxide and water to produce carbohydrates and oxygen . The photosynthesis carried out by all

6552-432: The combustion of fossil fuels, ammonia gas which evaporates from agricultural fields which have had fertilizers applied to them, and dust. Anthropogenic nitrogen inputs account for about 80% of all nitrogen fluxes in ecosystems. When plant tissues are shed or are eaten, the nitrogen in those tissues becomes available to animals and microbes. Microbial decomposition releases nitrogen compounds from dead organic matter in

6656-418: The concept to draw attention to the importance of transfers of materials between organisms and their environment. He later refined the term, describing it as "The whole system, ... including not only the organism-complex, but also the whole complex of physical factors forming what we call the environment". Tansley regarded ecosystems not simply as natural units, but as "mental isolates". Tansley later defined

6760-512: The conservation and sustainable use of biological diversity, and it is often seen as the key document regarding sustainable development . The Convention was opened for signature at the Earth Summit in Rio de Janeiro on 5 June 1992 and entered into force on 29 December 1993. The United States is the only UN member state which has not ratified the Convention. It has two supplementary agreements,

6864-629: The contributions of biodiversity to achieving the SDGs. A new plan, known as the post-2020 Global Biodiversity Framework (GBF) was developed to guide action through 2030. A first draft of this framework was released in July 2021, and its final content was discussed and negotiated as part of the COP 15 meetings. Reducing agricultural pollution and sharing the benefits of digital sequence information arose as key points of contention among Parties during development of

6968-403: The dead organic matter would accumulate in an ecosystem, and nutrients and atmospheric carbon dioxide would be depleted. Decomposition processes can be separated into three categories— leaching , fragmentation and chemical alteration of dead material. As water moves through dead organic matter, it dissolves and carries with it the water-soluble components. These are then taken up by organisms in

7072-447: The definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Biotic factors of the ecosystem are living things; such as plants, animals, and bacteria, while abiotic are non-living components; such as water, soil and atmosphere. Plants allow energy to enter the system through photosynthesis , building up plant tissue. Animals play an important role in

7176-566: The description of Ecologically or Biologically Significant Marine Areas (EBSAs) and describing new areas. These have focused on the North-East, North-West and South-Eastern Atlantic Ocean , Baltic Sea , Caspian Sea , Black Sea , Seas of East Asia, North-West Indian Ocean and Adjacent Gulf Areas, Southern and North-East Indian Ocean, Mediterranean Sea , North and South Pacific, Eastern Tropical and Temperate Pacific, Wider Caribbean and Western Mid-Atlantic. The workshop meetings have followed

7280-655: The ecosystem or to gradual disruption of biotic processes and degradation of abiotic conditions of the ecosystem. Once the original ecosystem has lost its defining features, it is considered "collapsed ". Ecosystem restoration can contribute to achieving the Sustainable Development Goals . An ecosystem (or ecological system) consists of all the organisms and the abiotic pools (or physical environment) with which they interact. The biotic and abiotic components are linked together through nutrient cycles and energy flows. "Ecosystem processes" are

7384-528: The ecosystem. Internal factors are controlled, for example, by decomposition , root competition, shading, disturbance, succession, and the types of species present. While the resource inputs are generally controlled by external processes, the availability of these resources within the ecosystem is controlled by internal factors. Therefore, internal factors not only control ecosystem processes but are also controlled by them. Ecosystems are dynamic entities—they are subject to periodic disturbances and are always in

7488-421: The effective implementation of one of the three objectives of the CBD: the fair and equitable sharing of benefits arising out of the utilization of genetic resources. It thereby contributes to the conservation and sustainable use of biodiversity. Also at the tenth meeting of the Conference of the Parties , held from 18 to 29 October 2010 in Nagoya, a revised and updated "Strategic Plan for Biodiversity, 2011-2020"

7592-463: The finite status of natural resources and sets out a philosophy of sustainable use . While past conservation efforts were aimed at protecting particular species and habitats, the Convention recognizes that ecosystems, species and genes must be used for the benefit of humans. However, this should be done in a way and at a rate that does not lead to the long-term decline of biological diversity. The Convention also offers decision-makers guidance based on

7696-422: The first time in international law that the conservation of biodiversity is "a common concern of humankind" and is an integral part of the development process. The agreement covers all ecosystems , species , and genetic resources. It links traditional conservation efforts to the economic goal of using biological resources sustainably. It sets principles for the fair and equitable sharing of the benefits arising from

7800-421: The flow of energy through a lake was the primary driver of the ecosystem. Hutchinson's students, brothers Howard T. Odum and Eugene P. Odum , further developed a "systems approach" to the study of ecosystems. This allowed them to study the flow of energy and material through ecological systems. Ecosystems are controlled by both external and internal factors. External factors, also called state factors, control

7904-458: The forests of eastern North America still show legacies of cultivation which ceased in 1850 when large areas were reverted to forests. Another example is the methane production in eastern Siberian lakes that is controlled by organic matter which accumulated during the Pleistocene . Ecosystems continually exchange energy and carbon with the wider environment . Mineral nutrients, on

8008-438: The framework. A final version was adopted by the Convention on 19 December 2022. The framework includes a number of ambitious goals, including a commitment to designate at least 30 percent of global land and sea as protected areas (known as the " 30 by 30 " initiative). The CBD has a significant focus on marine and coastal biodiversity . A series of expert workshops have been held (2018–2022) to identify options for modifying

8112-488: The implementation of the programme of work, coordinate with other international organizations, and collect and disseminate information. The SBSTTA is a committee composed of experts from member governments competent in relevant fields. It plays a key role in making recommendations to the COP on scientific and technical issues. It provides assessments of the status of biological diversity and of various measures taken in accordance with Convention, and also gives recommendations to

8216-691: The landscape, versus one present on an adjacent steep hillside. Other external factors that play an important role in ecosystem functioning include time and potential biota , the organisms that are present in a region and could potentially occupy a particular site. Ecosystems in similar environments that are located in different parts of the world can end up doing things very differently simply because they have different pools of species present. The introduction of non-native species can cause substantial shifts in ecosystem function. Unlike external factors, internal factors in ecosystems not only control ecosystem processes but are also controlled by them. While

8320-535: The limited nature of a plurilateral treaty, the full cooperation of the parties to the treaty is required in order for the object of the treaty to be met. As a result, reservations to plurilateral treaties are not allowed without the consent of all other parties to the treaty. This principle is codified in international law by article 20(2) of the Vienna Convention on the Law of Treaties : When it appears from

8424-514: The limited number of the negotiating states and the object and purpose of a treaty that the application of the treaty in its entirety between all the parties is an essential condition of the consent of each one to be bound by the treaty, a reservation requires acceptance by all the parties. An example of a plurilateral treaty is the Antarctic Treaty , signed on 1 December 1959. Ecosystem An ecosystem (or ecological system )

8528-423: The living and dead plant matter, and eventually released through respiration. The carbon and energy incorporated into plant tissues (net primary production) is either consumed by animals while the plant is alive, or it remains uneaten when the plant tissue dies and becomes detritus . In terrestrial ecosystems , the vast majority of the net primary production ends up being broken down by decomposers . The remainder

8632-408: The maintenance of hydrological cycles, cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. While material from the ecosystem had traditionally been recognized as being the basis for things of economic value, ecosystem services tend to be taken for granted. The Millennium Ecosystem Assessment

8736-405: The maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. Many ecosystems become degraded through human impacts, such as soil loss , air and water pollution , habitat fragmentation , water diversion , fire suppression , and introduced species and invasive species . These threats can lead to abrupt transformation of

8840-433: The movement of matter and energy through the system, by feeding on plants and on one another. They also influence the quantity of plant and microbial biomass present. By breaking down dead organic matter , decomposers release carbon back to the atmosphere and facilitate nutrient cycling by converting nutrients stored in dead biomass back to a form that can be readily used by plants and microbes. Ecosystems provide

8944-537: The other hand, are mostly cycled back and forth between plants, animals, microbes and the soil. Most nitrogen enters ecosystems through biological nitrogen fixation , is deposited through precipitation, dust, gases or is applied as fertilizer . Most terrestrial ecosystems are nitrogen-limited in the short term making nitrogen cycling an important control on ecosystem production. Over the long term, phosphorus availability can also be critical. Macronutrients which are required by all plants in large quantities include

9048-509: The other. Despite this, the cumulative effect of additional species in an ecosystem is not linear: additional species may enhance nitrogen retention, for example. However, beyond some level of species richness, additional species may have little additive effect unless they differ substantially from species already present. This is the case for example for exotic species . The addition (or loss) of species that are ecologically similar to those already present in an ecosystem tends to only have

9152-403: The overall structure of an ecosystem and the way things work within it, but are not themselves influenced by the ecosystem. On broad geographic scales, climate is the factor that "most strongly determines ecosystem processes and structure". Climate determines the biome in which the ecosystem is embedded. Rainfall patterns and seasonal temperatures influence photosynthesis and thereby determine

9256-405: The plants in an ecosystem is called the gross primary production (GPP). About half of the gross GPP is respired by plants in order to provide the energy that supports their growth and maintenance. The remainder, that portion of GPP that is not used up by respiration, is known as the net primary production (NPP). Total photosynthesis is limited by a range of environmental factors. These include

9360-585: The primary nutrients (which are most limiting as they are used in largest amounts): Nitrogen, phosphorus, potassium. Secondary major nutrients (less often limiting) include: Calcium, magnesium, sulfur. Micronutrients required by all plants in small quantities include boron, chloride, copper, iron, manganese, molybdenum, zinc. Finally, there are also beneficial nutrients which may be required by certain plants or by plants under specific environmental conditions: aluminum, cobalt, iodine, nickel, selenium, silicon, sodium, vanadium. Until modern times, nitrogen fixation

9464-413: The primary producers. The organisms that consume their tissues are called primary consumers or secondary producers — herbivores . Organisms which feed on microbes ( bacteria and fungi ) are termed microbivores . Animals that feed on primary consumers— carnivores —are secondary consumers. Each of these constitutes a trophic level. The sequence of consumption—from plant to herbivore, to carnivore—forms

9568-419: The process of recovering from some past disturbance. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance, is termed its resistance . The capacity of a system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Ecosystems can be studied through

9672-406: The protection of weaker states. A plurilateral treaty is a special type of multilateral treaty. A plurilateral treaty is a treaty between a limited number of states with a particular interest in the subject of the treaty. The primary difference between a plurilateral treaty and other multilateral treaties is that the availability of reservations is more limited under a plurilateral treaty. Due to

9776-585: The protocol to make it illegal to publicly share genetic information , e.g. via GenBank . William Yancey Brown , when with the Brookings Institution , suggested that the Convention on Biological Diversity should include the preservation of intact genomes and viable cells for every known species and for new species as they are discovered. A Conference of the Parties (COP) was held annually for three years after 1994, and thence biennially on even-numbered years. The first ordinary meeting of

9880-669: The rest as deficient, seriously deficient or totally deficient. Scientists working with biodiversity and medical research are expressing fears that the Nagoya Protocol is counterproductive, and will hamper disease prevention and conservation efforts, and that the threat of imprisonment of scientists will have a chilling effect on research. Non-commercial researchers and institutions such as natural history museums fear maintaining biological reference collections and exchanging material between institutions will become difficult, and medical researchers have expressed alarm at plans to expand

9984-611: The soil, react with mineral soil, or are transported beyond the confines of the ecosystem (and are considered lost to it). Newly shed leaves and newly dead animals have high concentrations of water-soluble components and include sugars , amino acids and mineral nutrients. Leaching is more important in wet environments and less important in dry ones. Fragmentation processes break organic material into smaller pieces, exposing new surfaces for colonization by microbes. Freshly shed leaf litter may be inaccessible due to an outer layer of cuticle or bark , and cell contents are protected by

10088-511: The soil, where plants, fungi, and bacteria compete for it. Some soil bacteria use organic nitrogen-containing compounds as a source of carbon, and release ammonium ions into the soil. This process is known as nitrogen mineralization . Others convert ammonium to nitrite and nitrate ions, a process known as nitrification . Nitric oxide and nitrous oxide are also produced during nitrification. Under nitrogen-rich and oxygen-poor conditions, nitrates and nitrites are converted to nitrogen gas ,

10192-483: The spatial extent of ecosystems using the term " ecotope ". G. Evelyn Hutchinson , a limnologist who was a contemporary of Tansley's, combined Charles Elton 's ideas about trophic ecology with those of Russian geochemist Vladimir Vernadsky . As a result, he suggested that mineral nutrient availability in a lake limited algal production . This would, in turn, limit the abundance of animals that feed on algae. Raymond Lindeman took these ideas further to suggest that

10296-421: The transfers of energy and materials from one pool to another. Ecosystem processes are known to "take place at a wide range of scales". Therefore, the correct scale of study depends on the question asked. The term "ecosystem" was first used in 1935 in a publication by British ecologist Arthur Tansley . The term was coined by Arthur Roy Clapham , who came up with the word at Tansley's request. Tansley devised

10400-435: The use of genetic resources, notably those destined for commercial use. It also covers the rapidly expanding field of biotechnology through its Cartagena Protocol on Biosafety , addressing technology development and transfer, benefit-sharing and biosafety issues. Importantly, the Convention is legally binding; countries that join it ('Parties') are obliged to implement its provisions. The Convention reminds decision-makers of

10504-797: Was adopted on 29 October 2010 in Nagoya, Japan , and entered into force on 12 October 2014. 2010 was also the International Year of Biodiversity , and the Secretariat of the CBD was its focal point. Following a recommendation of CBD signatories at Nagoya, the UN declared 2011 to 2020 as the United Nations Decade on Biodiversity in December 2010. The Convention's Strategic Plan for Biodiversity 2011-2020 , created in 2010, include

10608-446: Was agreed and published. This document included the "Aichi Biodiversity Targets", comprising 20 targets that address each of five strategic goals defined in the plan. The strategic plan includes the following strategic goals: Upon the launch of Agenda 2030 , CBD released a technical note mapping and identifying synergies between the 17 Sustainable Development Goals (SDGs) and the 20 Aichi Biodiversity Targets. This helps to understand

10712-582: Was reached in May 2003. In accordance with the provisions of its Article 37, the Protocol entered into force on 11 September 2003. In April 2002, the Parties of the UN CBD adopted the recommendations of the Gran Canaria Declaration Calling for a Global Plant Conservation Strategy, and adopted a 16-point plan aiming to slow the rate of plant extinctions around the world by 2010. The Nagoya Protocol on Access to Genetic Resources and

10816-679: Was the major source of nitrogen for ecosystems. Nitrogen-fixing bacteria either live symbiotically with plants or live freely in the soil. The energetic cost is high for plants that support nitrogen-fixing symbionts—as much as 25% of gross primary production when measured in controlled conditions. Many members of the legume plant family support nitrogen-fixing symbionts. Some cyanobacteria are also capable of nitrogen fixation. These are phototrophs , which carry out photosynthesis. Like other nitrogen-fixing bacteria, they can either be free-living or have symbiotic relationships with plants. Other sources of nitrogen include acid deposition produced through

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