98-515: The Aphrophoridae or spittlebugs are a family of insects belonging to the order Hemiptera . There are at least 160 genera and 990 described species in Aphrophoridae. This Hemiptera article related to members of the insect suborder Auchenorrhyncha is a stub . You can help Misplaced Pages by expanding it . Hemiptera Hemiptera ( / h ɛ ˈ m ɪ p t ər ə / ; from Ancient Greek hemipterus 'half-winged')
196-408: A pupal stage and undergo complete metamorphosis . The majority of species are terrestrial, including a number of important agricultural pests, but some are found in freshwater habitats. These include the water boatmen , backswimmers , pond skaters , and giant water bugs . Hemiptera belong to the insect superorder Paraneoptera , which includes lice ( Psocodea ), thrips ( Thysanoptera ), and
294-582: A brilliant radiance by Athena, Achilles stands next to the Achaean wall and roars in rage. The Trojans are terrified by his appearance, and the Achaeans manage to bear Patroclus's body away. Polydamas again urges Hector to withdraw into the city; again, Hector refuses, and the Trojans camp on the plain at nightfall. Achilles mourns Patroclus, brokenhearted. Meanwhile, at Thetis's request, Hephaestus fashions
392-484: A conclusion determines the validity of evidence. Some scholars believe that the gods may have intervened in the mortal world because of quarrels they may have had with each other. Homer interprets the world at this time by using the passion and emotion of the gods to be determining factors of what happens on the human level. An example of one of these relationships in the Iliad occurs between Athena, Hera, and Aphrodite. In
490-673: A double layer of grains, coarser on the outside. The Amazon rain forest cicada Hemisciera maculipennis displays bright red deimatic flash coloration on its hindwings when threatened; the sudden contrast helps to startle predators, giving the cicada time to escape. The coloured patch on the hindwing is concealed at rest by an olive green patch of the same size on the forewing, enabling the insect to switch rapidly from cryptic to deimatic behaviour. Some hemipterans such as firebugs have bold aposematic warning coloration, often red and black, which appear to deter passerine birds. Many hemipterans including aphids, scale insects and especially
588-469: A dream to Agamemnon, urging him to attack Troy. Agamemnon heeds the dream but first decides to test the Achaean army's morale by telling them to go home. But nine years into the war, the soldiers' morale has worn thin. The plan backfires, and only the intervention of Odysseus, inspired by Athena , stops a rout . Odysseus confronts and beats Thersites , a common soldier who voices discontent about fighting Agamemnon's war. The Achaeans deploy in companies upon
686-443: A greater or lesser degree. The nymphs moult several times as they grow, and each instar resembles the adult more than the previous one. Wing buds grow in later stage nymphs; the final transformation involves little more than the development of functional wings (if they are present at all) and functioning sexual organs, with no intervening pupal stage as in holometabolous insects. Many aphids are parthenogenetic during part of
784-620: A herbaceous plant. Phloem sap, which has a higher concentration of sugars and nitrogen, is under positive pressure unlike the more dilute xylem sap. Most of the Sternorrhyncha and a number of Auchenorrhynchan groups feed on phloem. Phloem feeding is common in the Fulgoromorpha, most Cicadellidae and in the Heteroptera. The Typhlocybine Cicadellids specialize in feeding on non-vascular mesophyll tissue of leaves, which
882-404: A jam jar in and out. Stridulatory sounds are produced among the aquatic Corixidae and Notonectidae (backswimmers) using tibial combs rubbed across rostral ridges. Hemipterans are hemimetabolous , meaning that they do not undergo metamorphosis , the complete change of form between a larval phase and an adult phase . Instead, their young are called nymphs , and resemble the adults to
980-468: A new set of armor for Achilles, including a magnificently wrought shield . In the morning, Thetis brings Achilles his new set of armor, only to find him weeping over Patroclus's body. Achilles arms for battle and rallies the Achaean warriors. Agamemnon gives Achilles all the promised gifts, including Briseis , but Achilles is indifferent to them. The Achaeans take their meal; Achilles refuses to eat. His horse, Xanthos , prophesies Achilles's death; Achilles
1078-549: A priest of Apollo , offers the Achaeans wealth for the return of his daughter Chryseis , held captive by Agamemnon. Although most of the Achaean kings are in favor of the offer, Agamemnon refuses. Chryses prays for Apollo's help, and Apollo sends a plague to afflict the Achaean army. After nine days of plague, Achilles , the leader of the Myrmidon forces and aristos achaion ("best of the Greeks"), calls an assembly to deal with
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#17327903150201176-504: A result of this thinking, each god or goddess in polytheistic Greek religion is attributed to an aspect of the human world. For example, Poseidon is the god of the sea, Aphrodite is the goddess of beauty, Ares is the god of war, and so on and so forth for many other gods. This is how Greek culture was defined as many Athenians felt the presence of their gods through divine intervention in significant events in their lives. Oftentimes, they found these events to be mysterious and inexplicable. In
1274-488: A single species. This kind of polymorphism tends to be helpful when habitats are temporary with more energy put into reproduction when food is available and into dispersal through flight when food becomes scarce. In aphids, both winged and wingless forms occur with winged forms produced in greater numbers when food resources are depleted. Aphids and whiteflies can sometimes be transported very long distances by atmospheric updrafts and high altitude winds. Wing-length polymorphism
1372-524: A stern admonition to come back to him and not to pursue the Trojans. Achilles says that after all has been made right, he and Patroclus will take Troy together. Patroclus leads the Myrmidons into battle and arrives as the Trojans set fire to the first ships. The Trojans are routed by the sudden onslaught, and Patroclus begins his assault by killing Zeus's son Sarpedon , a leading ally of the Trojans. Patroclus, ignoring Achilles's command, pursues and reaches
1470-601: A take-off velocity of up to 2.7 metres per second and an acceleration of up to 250 g. The instantaneous power output is much greater than that of normal muscle, implying that energy is stored and released to catapult the insect into the air. Cicadas, which are much larger, extend their hind legs for a jump in under a millisecond, again implying elastic storage of energy for sudden release. Instead of relying on any form of locomotion, most Sternorrhyncha females are sedentary or completely sessile, attached to their host plants by their thin feeding stylets which cannot be taken out of
1568-409: A variety of modes of locomotion including swimming, skating on a water surface and jumping, as well as walking and flying like other insects. Several families of Heteroptera are water bugs , adapted to an aquatic lifestyle , such as the water boatmen (Corixidae), water scorpions (Nepidae), and backswimmers (Notonectidae). They are mostly predatory, and have legs adapted as paddles to help
1666-536: A water surface, in the same way that a toy soap boat propels itself. Water bugs in the genus Microvelia ( Veliidae ) can travel at up to 17 cm/s, twice as fast as they can walk, by this means. Flight is well developed in the Hemiptera although mostly used for short distance movement and dispersal. Wing development is sometimes related to environmental conditions. In some groups of Hemiptera, there are variations of winged, short-winged, and wingless forms within
1764-451: A wide variety of habitats, generally terrestrial, though some are adapted to life in or on the surface of fresh water (e.g. pondskaters , water boatmen , giant water bugs ). Hemipterans are hemimetabolous , with young nymphs that somewhat resemble adults. Many aphids are capable of parthenogenesis , producing young from unfertilised eggs; this helps them to reproduce extremely rapidly in favourable conditions. Humans have interacted with
1862-405: Is an order of insects , commonly called true bugs , comprising over 80,000 species within groups such as the cicadas , aphids , planthoppers , leafhoppers , assassin bugs , bed bugs , and shield bugs . They range in size from 1 mm (0.04 in) to around 15 cm (6 in), and share a common arrangement of piercing-sucking mouthparts . The name "true bugs" is often limited to
1960-520: Is beaten back by Hephaestus's firestorm. The gods fight amongst themselves. The great gates of the city are opened to receive the fleeing Trojans, and Apollo leads Achilles away from the city by pretending to be a Trojan. When Apollo reveals himself to Achilles, the Trojans have retreated into the city, all except for Hector. Despite the counsel of Polydamas and the pleas of his parents, Priam and Hecuba , Hector resolves to face Achilles. When Achilles approaches, however, Hector's will fails him. He flees and
2058-412: Is chased by Achilles around the city. Finally, Athena tricks him into stopping, and he turns to face his opponent. After a brief duel, Achilles stabs Hector through the neck. Before dying, Hector reminds Achilles that he, too, is fated to die. Achilles strips Hector of his own armor, gloating over his death. Achilles then dishonors Hector's body by lashing it to the back of his chariot and dragging it around
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#17327903150202156-595: Is enraged by Poseidon's intervention. However, he reassures Hera that Troy is still fated to fall once Hector kills Patroclus. Poseidon is recalled from the battlefield, and Zeus sends Apollo to aid the Trojans. The Trojans once again breach the wall, and the battle reaches the ships. Patroclus cannot stand to watch any longer and goes to Achilles, weeping. He briefly admonishes him for his stubbornness and then asks him to allow him to fight in his place, wearing Achilles's armor so that he will be mistaken for him. Achilles relents and lends Patroclus his armor but sends him off with
2254-411: Is even lower in amino acids and contains monosaccharides rather than sucrose , as well as organic acids and minerals. No digestion is required (except for the hydrolysis of sucrose) and 90% of the nutrients in the xylem sap can be utilised. Some phloem sap feeders selectively mix phloem and xylem sap to control the osmotic potential of the liquid consumed. A striking adaptation to a very dilute diet
2352-502: Is found in many hemipterans: a filter chamber, a part of the gut looped back on itself as a countercurrent exchanger, which permits nutrients to be separated from excess water. The residue, mostly water with sugars and amino acids, is quickly excreted as sticky "honey dew", notably from aphids but also from other Auchenorrhycha and Sternorrhyncha. Some Sternorrhyncha including Psyllids and some aphids are gall formers. These sap-sucking hemipterans inject fluids containing plant hormones into
2450-465: Is indifferent. Achilles goes into battle, with Automedon driving his chariot. Zeus lifts the ban on the gods' interference, and the gods freely help both sides. Achilles, burning with rage and grief, slays many. Achilles cuts off half the Trojans' number in the river and slaughters them, clogging the river with bodies. The river god, Scamander , confronts Achilles and commands him to stop killing Trojans, but Achilles refuses. They fight until Scamander
2548-405: Is lost in his grief and spends his days mourning Patroclus and dragging Hector's body behind his chariot. Dismayed by Achilles's continued abuse of Hector's body, Zeus decides that it must be returned to Priam. Led by Hermes , Priam takes a wagon filled with gifts out of Troy, across the plains, and into the Achaean camp unnoticed. He clasps Achilles by the knees and begs for his son's body. Achilles
2646-654: Is more nutritious than the leaf epidermis. Most Heteroptera also feed on mesophyll tissue where they are more likely to encounter defensive secondary plant metabolites which often leads to the evolution of host specificity. Obligate xylem feeding is a special habit that is found in the Auchenorrhyncha among Cicadoidea, Cercopoidea and in Cicadelline Cicadellids. Some phloem feeders may take to xylem sap facultatively, especially when facing dehydration. Xylem feeders tend to be polyphagous; to overcome
2744-437: Is moved to tears and finally relents in his anger. The two lament their losses in the war. Achilles agrees to give Hector's body back and to give the Trojans twelve days to properly mourn and bury him. Achilles apologizes to Patroclus, fearing he has dishonored him by returning Hector's body. After a meal, Priam carries Hector's body back into Troy. Hector is buried, and the city mourns. Ancient Greek religion had no founder and
2842-408: Is notably rare in tree-living Hemiptera. Many Auchenorrhyncha including representatives of the cicadas, leafhoppers, treehoppers, planthoppers, and froghoppers are adapted for jumping (saltation). Treehoppers, for example, jump by rapidly depressing their hind legs. Before jumping, the hind legs are raised and the femora are pressed tightly into curved indentations in the coxae. Treehoppers can attain
2940-522: Is one of the oldest extant works of literature still widely read by modern audiences. As with the Odyssey , the poem is divided into 24 books and was written in dactylic hexameter . It contains 15,693 lines in its most widely accepted version. Set towards the end of the Trojan War , a ten-year siege of the city of Troy by a coalition of Mycenaean Greek states, the poem depicts significant events in
3038-424: Is their "beak" in which the modified mandibles and maxillae form a "stylet" which is sheathed within a modified labium . The stylet is capable of piercing tissues and sucking liquids, while the labium supports it. The stylet contains a channel for the outward movement of saliva and another for the inward movement of liquid food. A salivary pump drives saliva into the prey; a cibarial pump extracts liquid from
Aphrophoridae - Misplaced Pages Continue
3136-418: Is true of many aquatic species which are predatory, either as nymphs or adults. The predatory shield bug for example stabs caterpillars with its beak and sucks out the body fluids. The saliva of predatory heteropterans contains digestive enzymes such as proteinase and phospholipase , and in some species also amylase . The mouthparts of these insects are adapted for predation. There are toothed stylets on
3234-562: The Asian citrus psyllid which transmits citrus greening disease . Iliad On the Greek side: On the Trojan side: The Iliad ( / ˈ ɪ l i ə d / ; Ancient Greek : Ἰλιάς , romanized : Iliás , [iː.li.ás] ; lit. ' [a poem] about Ilion (Troy) ' ) is one of two major ancient Greek epic poems attributed to Homer . It
3332-492: The Early Devonian period. Hemipterans can dramatically cut the mass of affected plants, especially in major outbreaks. They sometimes also change the mix of plants by predation on seeds or feeding on roots of certain species. Some sap-suckers move from one host to another at different times of year. Many aphids spend the winter as eggs on a woody host plant and the summer as parthenogenetically reproducing females on
3430-494: The Iliad as a major piece of evidence for his theory of the Bicameral Mind , which posits that until about the time described in the Iliad , humans had a far different mentality from present-day humans. He says that humans during that time were lacking what is today called consciousness. He suggests that humans heard and obeyed commands from what they identified as gods until the change in human mentality that incorporated
3528-431: The Iliad , Paris challenges any of the Achaeans to a single combat and Menelaus steps forward. Menelaus is dominating the battle and is on the verge of killing Paris. "Now he'd have hauled him off and won undying glory but Aphrodite, Zeus's daughter, was quick to the mark, snapped the rawhide strap." Aphrodite intervenes out of her own self-interest to save Paris from the wrath of Menelaus because Paris had helped her to win
3626-530: The Malpighian tubules in their proximal segment that produce mucopolysaccharides, which form the froth around spittlebugs, offering a measure of protection. Parental care is found in many species of Hemiptera especially in members of the Membracidae and numerous Heteroptera. In many species of shield bug, females stand guard over their egg clusters to protect them from egg parasitoids and predators. In
3724-591: The Psylloidea and Aleyrodoidea . Aphids and Coccoids appear in the Triassic . The Coleorrhyncha extend back to the Lower Jurassic . The Heteroptera first appeared in the Triassic . The present members of the order Hemiptera (sometimes referred to as Rhynchota) were historically placed into two orders, the so-called Homoptera and Heteroptera/Hemiptera, based on differences in wing structure and
3822-530: The Sternorrhyncha and Auchenorrhyncha , or partially hardened, as in most Heteroptera . The name "Hemiptera" is from the Greek ἡμι- ( hemi ; "half") and πτερόν ( pteron ; "wing"), referring to the forewings of many heteropterans which are hardened near the base, but membranous at the ends. Wings modified in this manner are termed hemelytra (singular: hemelytron ), by analogy with
3920-482: The honeydew that these hemipterans secrete. The relationship is mutualistic , as both ant and aphid benefit. Ants such as the yellow anthill ant, Lasius flavus , breed aphids of at least four species, Geoica utricularia , Tetraneura ulmi , Forda marginata and Forda formicaria , taking eggs with them when they found a new colony; in return, these aphids are obligately associated with the ant, breeding mainly or wholly asexually inside anthills. Ants may also protect
4018-599: The lovebug is a fly and the Maybug and ladybug are beetles . The term is occasionally extended to colloquial names for freshwater or marine crustaceans (e.g. Balmain bug , Moreton Bay bug , mudbug ) and used by physicians and bacteriologists for disease-causing germs (e.g. superbugs ). Most hemipterans feed on plants, using their sucking and piercing mouthparts to extract plant sap . Some are bloodsucking, or hematophagous , while others are predators that feed on other insects or small invertebrates . They live in
Aphrophoridae - Misplaced Pages Continue
4116-615: The Gods, allows it. This motif recurs when he considers sparing Hector, whom he loves and respects. This time, it is Athena who challenges him: Father of the shining bolt, dark misted, what is this you said? Do you wish to bring back a man who is mortal, one long since doomed by his destiny, from ill-sounding death and release him? Do it, then; but not all the rest of us gods shall approve you. Again, Zeus appears capable of altering fate, but does not, deciding instead to abide by set outcomes; similarly, fate spares Aeneas after Apollo convinces
4214-452: The Hemiptera for millennia. Some species, including many aphids, are significant agricultural pests , damaging crops by sucking the sap. Others harm humans more directly as vectors of serious viral diseases. The bed bug is a persistent parasite of humans, and some kissing bugs can transmit Chagas disease . Some species have been used for biological control of insect pests or of invasive plants. A few hemipterans, have been cultivated for
4312-477: The Trojan Dolon , and wreak havoc in the camps of some Thracian allies of Troy. In the morning, the fighting is fierce, and Agamemnon, Diomedes, and Odysseus are all wounded. Achilles sends Patroclus from his camp to inquire about the Achaean casualties, and while there, Patroclus is moved to pity by a speech by Nestor . Nestor asks Patroclus to beg Achilles to rejoin the fighting, or if he will not, to lead
4410-530: The Trojan archer Pandarus to shoot Menelaus. Menelaus is wounded, and the truce is broken. Fighting breaks out, and many minor Trojans are killed. In the fighting, Diomedes kills many Trojans, including Pandarus, and defeats Aeneas . Aphrodite rescues him before he can be killed, but Diomedes attacks her and wounds the goddess's wrist. Apollo faces Diomedes and warns him against warring with gods, which Diomedes ignores. Apollo sends Ares to defeat Diomedes. Many heroes and commanders join in, including Hector, and
4508-411: The Trojan plain. When news of the Achaean deployment reaches King Priam , the Trojans respond in a sortie upon the plain. The armies approach each other, but before they meet, Paris offers to end the war by fighting a duel with Menelaus , urged by Hector , his brother and hero of Troy . Here, the initial cause of the entire war is explained: Helen , wife of Menelaus, and the most beautiful woman in
4606-433: The Trojans back. Poseidon's nephew Amphimachus is killed in the battle; Poseidon imbues Idomeneus with godly power. Many fall on both sides. The Trojan seer Polydamas urges Hector to fall back because of a bad omen but is ignored. Hera seduces Zeus and lulls him to sleep, allowing Poseidon to help the Greeks. The Trojans are driven back onto the plain. Ajax wounds Hector, who is then carried back to Troy. Zeus awakes and
4704-414: The animal move through the water. The pondskaters or water striders (Gerridae) are also associated with water, but use the surface tension of standing water to keep them above the surface; they include the sea skaters in the genus Halobates , the only truly marine group of insects. Marangoni effect propulsion exploits the change in surface tension when a soap-like surfactant is released on to
4802-413: The aquatic Belostomatidae, females lay their eggs on the back of the male which guards the eggs. Protection provided by ants is common in the Auchenorrhyncha. Although many species of Hemiptera are significant pests of crops and garden plants, including many species of aphid and scale insects , other species are harmless. The damage done is often not so much the deprivation of the plant of its sap, but
4900-423: The army wearing Achilles's armor. The Trojans attack the Achaean wall on foot. Hector leads the terrible fighting, despite an omen that their charge will fail. The Achaeans are overwhelmed and routed, the wall's gate is broken, and Hector charges in. The Achaeans fall back to their ships. Poseidon pities the Achaeans and decides to disobey Zeus and help them. He rallies the Achaeans' spirits, and they begin to push
4998-432: The beauty pageant. The partisanship of Aphrodite towards Paris induces constant intervention by all of the gods, especially to give motivational speeches to their respective protégés, while often appearing in the shape of a human being they are familiar with. This connection of emotions to actions is just one example out of many that occur throughout the poem. Fate ( κήρ , kēr , 'fated death') propels most of
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#17327903150205096-428: The body. When Achilles hears of Patroclus's death, he screams so loudly in his grief that his mother, Thetis, hears him from the bottom of the ocean. Thetis grieves too, knowing that Achilles is fated to die young if he kills Hector. Though he knows it will seal his own fate, Achilles vows to kill Hector in order to avenge Patroclus. Achilles is urged to help retrieve Patroclus's body but has no armor to wear. Bathed in
5194-459: The city walls. He then rejoins the battle. Hector duels with Ajax , but nightfall interrupts the fight, and both sides retire. The Trojans quarrel about returning Helen. Paris offers to return the treasure he took and give further wealth as compensation, but not Helen, and the offer is refused. Both sides agree to a day's truce to burn the dead. The Achaeans also build a wall and trench to protect their camp and ships. The next morning, Zeus prohibits
5292-406: The city. The Trojans grieve. The ghost of Patroclus comes to Achilles in a dream, urging him to carry out the burial rites so that his spirit can move on to the underworld. Patroclus asks Achilles to arrange for their bones to be entombed together in a single urn; Achilles agrees, and Patroclus's body is cremated. The Achaeans hold a day of funeral games, and Achilles gives out the prizes. Achilles
5390-513: The completely hardened elytra of beetles , and occur only in the suborder Heteroptera . In all suborders, the hindwings – if present at all – are entirely membranous and usually shorter than the forewings. The forewings may be held "roofwise" over the body (typical of Sternorrhyncha and Auchenorrhyncha ), or held flat on the back, with the ends overlapping (typical of Heteroptera ). The antennae in Hemiptera typically consist of four or five segments, although they can still be quite long, and
5488-496: The dangerous Chagas disease . The first known hemipteran to feed in this way on vertebrates was the extinct assassin bug Triatoma dominicana found fossilized in amber and dating back about twenty million years. Faecal pellets fossilised beside it show that it transmitted a disease-causing Trypanosoma and the amber included hairs of the likely host, a bat . Some species of ant protect and farm aphids (Sternorrhyncha) and other sap-sucking hemipterans, gathering and eating
5586-824: The disparity in the analyses is suggested to be the long branch attraction effect in phylogenetic analysis, due to rapidly evolving DNA regions. The cladogram shows Hemiptera's placement within Paraneoptera , as well as how Hemiptera's four suborders are related. English names are given in parentheses where possible. Other insects [REDACTED] (barklice, booklice, lice) [REDACTED] Thysanoptera (thrips) [REDACTED] (aphids, scale insects, etc) [REDACTED] (shield bugs, assassin bugs, etc) [REDACTED] (moss bugs) [REDACTED] Fulgoromorpha (planthoppers) [REDACTED] Cicadomorpha (cicadas, leafhoppers, spittlebugs, etc) [REDACTED] The defining feature of hemipterans
5684-466: The events of the Iliad . Once set, gods and men abide it, neither truly able nor willing to contest it. How fate is set is unknown, but it is told by the Fates and by Zeus through sending omens to seers such as Calchas . Men and their gods continually speak of heroic acceptance and cowardly avoidance of one's slated fate. Fate does not determine every action, incident, and occurrence, but it does determine
5782-620: The extraction of dyestuffs such as cochineal and carmine , and for shellac . Cicadas have been used as food, and have appeared in literature since the Iliad in Ancient Greece . Hemiptera is the largest order of hemimetabolous insects (not undergoing complete metamorphosis; though some examples such as male scale insects do undergo a form of complete metamorphosis ), containing over 95,000 named species. Other insect orders with more species are all holometabolous , meaning they have
5880-429: The fact that they transmit serious viral diseases between plants. They often produce copious amounts of honeydew which encourages the growth of sooty mould . Significant pests include the cottony cushion scale , a pest of citrus fruit trees, the green peach aphid and other aphids which attack crops worldwide and transmit diseases, and jumping plant lice which can be plant-specific and transmit diseases, as with
5978-399: The fighting. Achilles and his companion Patroclus receive the embassy well. However, considering the slight to his honor too great, Achilles angrily refuses Agamemnon's offer and declares that he will only return to battle if the Trojans reach his ships and threaten them with fire. The embassy returns empty-handed. Later that night, Odysseus and Diomedes venture out to the Trojan lines, kill
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#17327903150206076-470: The final book of the poem, Homer writes, "He offended Athena and Hera—both goddesses." Athena and Hera are envious of Aphrodite because of a beauty pageant on Mount Olympus in which Paris chose Aphrodite to be the most beautiful goddess over both Hera and Athena. Wolfgang Kullmann further goes on to say, "Hera's and Athena's disappointment over the victory of Aphrodite in the Judgement of Paris determines
6174-408: The first writers to name and describe the gods' appearance and character. Mary Lefkowitz discusses the relevance of divine action in the Iliad , attempting to answer the question of whether divine intervention is a discrete occurrence (for its own sake) or if such godly behaviors are mere human character metaphors. The intellectual interest of Classic-era authors, such as Thucydides and Plato ,
6272-428: The gates of Troy, where Apollo himself stops him. Patroclus kills Hector's brother Cebriones , is set upon by Apollo and Euphorbos , and is finally killed by Hector. Hector takes Achilles's armor from the fallen Patroclus. The Achaeans fight to retrieve Patroclus's body from the Trojans, who attempt to carry it back to Troy at Hector's command. Antilochus is sent to tell Achilles the news and asks him to help retrieve
6370-532: The gods from interfering, and fighting begins anew. The Trojans prevail and force the Achaeans back to their wall. Hera and Athena are forbidden to help. Night falls before the Trojans can assail the Achaean wall. They camp in the field to attack at first light, and their watchfires light the plain like stars. Meanwhile, the Achaeans are desperate. Agamemnon admits his error and sends an embassy composed of Odysseus, Ajax, Phoenix , and two heralds to offer Briseis and extensive gifts to Achilles, if only he will return to
6468-535: The gods supporting each side try to influence the battle. Emboldened by Athena, Diomedes wounds Ares and puts him out of action. Hector rallies the Trojans and prevents a rout. Diomedes and the Trojan Glaucus find common ground after a duel and exchange unequal gifts, while Glaucus tells Diomedes the story of Bellerophon . Hector enters the city, urges prayers and sacrifices, incites Paris to battle, and bids his wife Andromache and son Astyanax farewell on
6566-491: The hands of Aiakos' great son, Achilleus. Here, Patroclus alludes to his fated death by Hector's hand and to Hector's fated death by Achilles's hand. Each accepts the outcome of his life, yet no one knows if the gods can alter fate. The first instance of this doubt occurs in Book 16. Seeing Patroclus about to kill Sarpedon, his mortal son, Zeus says: Ah me, that it is destined that the dearest of men, Sarpedon, must go down under
6664-399: The hands of Menoitios' son Patroclus. About his dilemma, Hera asks Zeus: Majesty, son of Kronos, what sort of thing have you spoken? Do you wish to bring back a man who is mortal, one long since doomed by his destiny, from ill-sounding death and release him? Do it, then; but not all the rest of us gods shall approve you. In deciding between losing a son or abiding fate, Zeus, King of
6762-457: The heteropteran Oncopeltus fasciatus when it consumes milkweeds , while the coreid stinkbug Amorbus rubiginosus acquires 2- hexenal from its food plant, Eucalyptus . Some long-legged bugs mimic twigs, rocking to and fro to simulate the motion of a plant part in the wind. The nymph of the Masked hunter bug camouflages itself with sand grains, using its hind legs and tarsal fan to form
6860-562: The late 8th or early 7th century BC. Homer's authorship was infrequently questioned in antiquity , but contemporary scholarship predominantly assumes that the Iliad and the Odyssey were composed independently and that the stories formed as part of a long oral tradition . The poem was performed by professional reciters of Homer known as rhapsodes . Critical themes in the poem include kleos (glory), pride, fate and wrath. Despite being predominantly known for its tragic and serious themes,
6958-426: The life cycle, such that females can produce unfertilized eggs, which are clones of their mother. All such young are females ( thelytoky ), so 100% of the population at these times can produce more offspring. Many species of aphid are also viviparous : the young are born live rather than laid as eggs. These adaptations enable aphids to reproduce extremely rapidly when conditions are suitable. Hemipterans make use of
7056-566: The literary Trojan War of the Iliad , the Olympian gods, goddesses, and minor deities fight among themselves and participate in human warfare, often by interfering with humans to counter other gods. Unlike their portrayals in Greek religion, Homer's portrayal of gods suits his narrative purpose. The gods in traditional thought of 4th-century Athenians were not spoken of in terms familiar to the works of Homer. The Classical-era historian Herodotus says that Homer and Hesiod , his contemporary, were
7154-438: The mandibles able to cut into and abrade tissues of their prey. There are further stylets on the maxillae, adapted as tubular canals to inject saliva and to extract the pre-digested and liquified contents of the prey. A few hemipterans are haematophagic ectoparasites ), feeding on the blood of larger animals. These include bedbugs and the triatomine kissing bugs of the assassin bug family Reduviidae , which can transmit
7252-412: The motivating force into the conscious self. He points out that almost every action in the Iliad is directed, caused, or influenced by a god and that earlier translations show an astonishing lack of words suggesting thought, planning, or introspection. Those that do appear, he argues, are misinterpretations made by translators imposing a modern mentality on the characters, a form of reverse logic by which
7350-582: The mouthparts of Siphonaptera , some Diptera and Thysanoptera superficially resemble the rostrum of the Hemiptera, but on closer inspection the differences are considerable. Aside from the mouthparts, various other insects can be confused with Hemiptera, but they all have biting mandibles and maxillae instead of the rostrum. Examples include cockroaches and psocids , both of which have longer, many-segmented antennae, and some beetles , but these have fully hardened forewings which do not overlap. The forewings of Hemiptera are either entirely membranous, as in
7448-719: The mysterious origin of fate is a power beyond the gods. Fate implies the primeval, tripartite division of the world that Zeus, Poseidon, and Hades effected in deposing their father, Cronus , for its dominion. Zeus took the Air and the Sky, Poseidon the Waters, and Hades the Underworld , the land of the dead—yet they share dominion of the Earth. Despite the earthly powers of the Olympic gods, only
7546-471: The negative pressure of xylem requires a special cibarial pump. Phloem feeding hemiptera typically have symbiotic micro-organisms in their gut that help to convert amino acids . Phloem feeders produce honeydew from their anus. A variety of organisms that feed on honeydew form symbiotic associations with phloem-feeders. Phloem sap is a sugary liquid low in amino acids, so insects have to process large quantities to meet their nutritional requirements. Xylem sap
7644-474: The outcome of life—before killing him, Hector calls Patroclus a fool for cowardly avoidance of his fate, by attempting his defeat; Patroclus retorts: No, deadly destiny, with the son of Leto, has killed me, and of men it was Euphorbos; you are only my third slayer. And put away in your heart this other thing that I tell you. You yourself are not one who shall live long, but now already death and powerful destiny are standing beside you, to go down under
7742-555: The overmatched Trojan to fight Achilles. Poseidon cautiously speaks: But come, let us ourselves get him away from death, for fear the son of Kronos may be angered if now Achilleus kills this man. It is destined that he shall be the survivor, that the generation of Dardanos shall not die… Divinely aided, Aeneas escapes the wrath of Achilles and survives the Trojan War. Whether or not the gods can alter fate, they do abide by it, despite its countering their human allegiances; thus,
7840-444: The plague. In the meantime, Agamemnon's messengers take Briseis away. Achilles becomes very upset and prays to his mother, Thetis , a minor goddess and sea nymph. Achilles asks his mother to ask Zeus to allow the Achaeans to be beaten back by the Trojans until their ships are at risk of burning. Only then will Agamemnon realize how much the Achaeans need Achilles and restore his honor. Thetis does so, and Zeus agrees. Zeus then sends
7938-947: The plant bugs from their natural enemies, removing the eggs of predatory beetles and preventing access by parasitic wasps. Some leafhoppers (Auchenorrhyncha) are similarly "milked" by ants. In the Corcovado rain forest of Costa Rica, wasps compete with ants to protect and milk leafhoppers; the leafhoppers preferentially give more honeydew, more often, to the wasps, which are larger and may offer better protection. Hemiptera form prey to predators including vertebrates, such as birds, and other invertebrates such as ladybirds . In response, hemipterans have evolved antipredator adaptations . Ranatra may feign death ( thanatosis ). Others such as Carpocoris purpureipennis secrete toxic fluids to ward off arthropod predators; some Pentatomidae such as Dolycoris are able to direct these fluids at an attacker. Toxic cardenolide compounds are accumulated by
8036-603: The plant quickly. Most hemipterans are phytophagous, using their sucking and piercing mouthparts to feed on plant sap. These include cicadas, leafhoppers, treehoppers, planthoppers, froghoppers, aphids, whiteflies, scale insects , and some other groups. Some are monophages , being host specific and only found on one plant taxon, others are oligophages , feeding on a few plant groups, while others again are less discriminating polyphages and feed on many species of plant. The relationship between hemipterans and plants appears to be ancient, with piercing and sucking of plants evident in
8134-454: The plant tissues inducing the production of tissue that covers to protects the insect and also act as sinks for nutrition that they feed on. The hackleberry gall psyllid for example, causes a woody gall on the leaf petioles of the hackleberry tree it infests, and the nymph of another psyllid produces a protective lerp out of hardened honeydew. Most other hemipterans are predatory , feeding on other insects, or even small vertebrates. This
8232-459: The planthoppers secrete wax to protect themselves from threats such as fungi, parasitoidal insects and predators, as well as abiotic factors like desiccation. Hard waxy coverings are especially important in the sedentary Sternorrhyncha such as scale insects, which have no means of escaping from predators; other Sternorrhyncha evade detection and attack by creating and living inside plant galls. Nymphal Cicadoidea and Cercopoidea have glands attached to
8330-441: The poem also contains instances of comedy and laughter. The poem is frequently described as a masculine or heroic epic, especially compared with the Odyssey . It contains detailed descriptions of ancient war instruments and battle tactics, and fewer female characters. The Olympian gods also play a major role in the poem, aiding their favoured warriors on the battlefield and intervening in personal disputes. Their characterisation in
8428-665: The poem humanised them for Ancient Greek audiences, giving a concrete sense of their cultural and religious tradition. In terms of formal style, the poem's repetitions and use of similes and epithets are often explored by scholars. The story begins with an invocation to the Muse . The events begin in medias res towards the end of the Trojan War, fought between the Trojans and the besieging Achaeans . The Achaean forces consist of armies from many different Greek kingdoms, led by their respective kings or princes. Agamemnon , king of Mycenae , acts as commander for these united armies. Chryses ,
8526-440: The position of the rostrum . "Homoptera" was established as paraphyletic group and an obsolete name. The order is now divided into four suborders, Heteroptera , Sternorrhyncha , Auchenorrhyncha , and Coleorrhyncha . The earlier work was based on nuclear DNA , but later phylogenetic analysis using mitochondrial DNA suggests that Homoptera may be monophyletic after all, a sister group of Heteroptera . The cause of
8624-455: The prey. Both pumps are powered by substantial dilator muscles in the head. The beak is usually folded under the body when not in use. The diet is typically plant sap , but some hemipterans such as assassin bugs are predators . Both herbivorous and predatory hemipterans inject enzymes to begin digestion extra-orally (before the food is taken into the body). These enzymes include amylase to hydrolyse starch , polygalacturonase to weaken
8722-422: The problem. Under pressure, Agamemnon agrees to return Chryseis to her father but decides to take Achilles's slave, Briseis , as compensation. Because war prizes were correlated with honor, Agamemnon's decision dishonors Achilles in front of the assembled Achaean forces. Achilles furiously declares that he and his men will no longer fight for Agamemnon. Odysseus returns Chryseis to her father, causing Apollo to end
8820-528: The siege's final weeks. In particular, it depicts a fierce quarrel between King Agamemnon and a celebrated warrior, Achilles . It is a central part of the Epic Cycle . The Iliad is often regarded as the first substantial piece of European literature . The Iliad and the Odyssey were likely written down in Homeric Greek , a literary mixture of Ionic Greek and other dialects, probably around
8918-576: The suborder Heteroptera . Entomologists reserve the term bug for Hemiptera or Heteroptera, which does not include other arthropods or insects of other orders such as ants , bees , beetles , or butterflies . In some varieties of English, all terrestrial arthropods (including non-insect arachnids , and myriapods ) also fall under the colloquial understanding of bug . Many insects with "bug" in their common name , especially in American English , belong to other orders; for example,
9016-401: The tarsi of the legs have two or three segments. Many hemipterans can produce sound for communication. The "song" of male cicadas, the loudest of any insect, is produced by tymbal organs on the underside of the abdomen, and is used to attract mates. The tymbals are drumlike disks of cuticle, which are clicked in and out repeatedly, making a sound in the same way as popping the metal lid of
9114-406: The tough cell walls of plants, and proteinases to break down proteins. Although the Hemiptera vary widely in their overall form, their mouthparts form a distinctive "rostrum". Other insect orders with mouthparts modified into anything like the rostrum and stylets of the Hemiptera include some Phthiraptera , but for other reasons they generally are easy to recognize as non-hemipteran. Similarly,
9212-800: The true bugs of Hemiptera. Within Paraneoptera, Hemiptera is most closely related to the sister clade Thysanoptera. The fossil record of hemipterans goes back to the Carboniferous ( Moscovian ). The oldest fossils are of the Archescytinidae from the Lower Permian and are thought to be basal to the Auchenorrhyncha . Fulgoromorpha and Cicadomorpha appear in the Upper Permian , as do Sternorrhyncha of
9310-517: The whole conduct of both goddesses in The Iliad and is the cause of their hatred for Paris, the Judge, and his town Troy." Hera and Athena then continue to support the Achaean forces throughout the poem because Paris is part of the Trojans, while Aphrodite aids Paris and the Trojans. The emotions between the goddesses often translate to actions they take in the mortal world. For example, in Book 3 of
9408-476: The world, is either through seduction or by force, taken by Paris from Menelaus's home in Sparta . Menelaus and Paris agree to duel; Helen will marry the victor. However, when Paris is beaten, Aphrodite rescues him and leads him to bed with Helen before Menelaus can kill him. The gods deliberate over whether the war should end here, but Hera convinces Zeus to wait for the utter destruction of Troy. Athena prompts
9506-436: Was limited to their utility as "a way of talking about human life rather than a description or a truth", because, if the gods remain religious figures, rather than human metaphors, their "existence"—without the foundation of either dogma or a bible of faiths—then allowed Greek culture the intellectual breadth and freedom to conjure gods fitting any religious function they required as a people. Psychologist Julian Jaynes uses
9604-536: Was not the creation of an inspired teacher. Rather, the religion arose out of the diverse beliefs of the Greek people. These beliefs coincide to the thoughts about the gods in polytheistic Greek religion. Adkins and Pollard agree with this by saying, "The early Greeks personalized every aspect of their world, natural and cultural, and their experiences in it. The earth, the sea, the mountains, the rivers, custom-law (themis), and one's share in society and its goods were all seen in personal as well as naturalistic terms." As
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