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Apocrita

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Order ( Latin : ordo ) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy . It is classified between family and class . In biological classification , the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes . An immediately higher rank, superorder , is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families.

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19-413: Apocrita is a suborder of insects in the order Hymenoptera . It includes wasps , bees , and ants , and consists of many families. It contains the most advanced hymenopterans and is distinguished from Symphyta by the narrow "waist" ( petiole ) formed between the first two segments of the actual abdomen ; the first abdominal segment is fused to the thorax, and is called the propodeum . Therefore, it

38-461: A cohors (plural cohortes ). Some of the plant families still retain the names of Linnaean "natural orders" or even the names of pre-Linnaean natural groups recognized by Linnaeus as orders in his natural classification (e.g. Palmae or Labiatae ). Such names are known as descriptive family names. In the field of zoology , the Linnaean orders were used more consistently. That is,

57-509: A capital letter. For some groups of organisms, their orders may follow consistent naming schemes . Orders of plants , fungi , and algae use the suffix -ales (e.g. Dictyotales ). Orders of birds and fishes use the Latin suffix -iformes meaning 'having the form of' (e.g. Passeriformes ), but orders of mammals and invertebrates are not so consistent (e.g. Artiodactyla , Actiniaria , Primates ). For some clades covered by

76-570: A distinct rank of biological classification having its own distinctive name (and not just called a higher genus ( genus summum )) was first introduced by the German botanist Augustus Quirinus Rivinus in his classification of plants that appeared in a series of treatises in the 1690s. Carl Linnaeus was the first to apply it consistently to the division of all three kingdoms of nature (then minerals , plants , and animals ) in his Systema Naturae (1735, 1st. Ed.). For plants, Linnaeus' orders in

95-419: Is determined by a taxonomist , as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with

114-417: Is general practice, when discussing the body of an apocritan in a technical sense, to refer to the mesosoma and metasoma (or gaster ) rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the parasitoid lifestyle of the ancestral apocritan, allowing more maneuverability of the female's ovipositor . The ovipositor either extends freely or

133-522: Is nearly always killed. Many parasitic hymenopterans are used as biological control agents to control pests, such as caterpillars , true bugs and hoppers , flies , and weevils . Aculeata is a monophyletic group that includes those species in which the female's ovipositor is modified into a stinger to inject venom . Groups within Aculeata include the familiar ants, bees, and various types of parasitic and predatory wasps; it also includes all of

152-598: Is not fully resolved. Cladogram of Apocrita after Peters et al.(2017) Ceraphronoidea Ichneumonoidea Cynipoidea Platygastroidea Chalcidoidea Diaprioidea Proctotrupoidea Evanioidea Stephanoidea Trigonaloidea Chrysidoidea Vespoidea (potter, honey and social wasps) Pompiloidea (velvet ants, spider wasps and relatives) Thynnoidea Tiphioidea Scolioidea Formicoidea (ants) Ampulicidae Sphecidae " Crabronidae " (digger wasps) Anthophila (bees) Suborder (biology) What does and does not belong to each order

171-405: Is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host (plant or animal) or in a nest cell provisioned by their mothers. Apocrita has historically been split into two groups, Parasitica and Aculeata . Aculeata is a clade whose name is in standard use. "Parasitica" is not a clade, as it is paraphyletic :

190-528: The International Code of Zoological Nomenclature , several additional classifications are sometimes used, although not all of these are officially recognized. In their 1997 classification of mammals , McKenna and Bell used two extra levels between superorder and order: grandorder and mirorder . Michael Novacek (1986) inserted them at the same position. Michael Benton (2005) inserted them between superorder and magnorder instead. This position

209-556: The Parasitica (containing many families of parasitoid wasps ) have been found on molecular analysis to be paraphyletic . Parasitoidism evolved once, and it is found today across most Apocritan families, though it has been secondarily lost several times. The phylogenetic tree gives a condensed overview of the phylogeny, illustrated with major groups. The sawflies are paraphyletic as the Apocrita evolved inside that group. The tree

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228-815: The Systema Naturae and the Species Plantarum were strictly artificial, introduced to subdivide the artificial classes into more comprehensible smaller groups. When the word ordo was first consistently used for natural units of plants, in 19th-century works such as the Prodromus Systematis Naturalis Regni Vegetabilis of Augustin Pyramus de Candolle and the Genera Plantarum of Bentham & Hooker, it indicated taxa that are now given

247-416: The social hymenopterans. Among the nonparasitic and nonsocial Aculeata, larvae are fed with captured prey (typically alive and paralyzed) or may be fed pollen and nectar. The social Aculeata feed their young prey ( paper wasps and hornets ), or pollen and nectar (bees), or perhaps seeds, fungi , or nonviable eggs (ants). The Apocrita contains a large number of families. Some traditional taxa such as

266-635: The clade would contain the Aculeata. "Parasitica" is therefore a rankless grouping in many present classifications, if it appears at all. Parasitica comprises the majority of hymenopteran insects, its members living as parasitoids . Most species are small, with the ovipositor adapted for piercing. In some hosts, the parasitoids induce metamorphosis prematurely, and in others it is prolonged. There are even species that are hyperparasites , or parasitoids on other parasitoids. The Parasitica lay their eggs inside or on another insect (egg, larva or pupa) and their larvae grow and develop within or on that host. The host

285-708: The orders in the zoology part of the Systema Naturae refer to natural groups. Some of his ordinal names are still in use, e.g. Lepidoptera (moths and butterflies) and Diptera (flies, mosquitoes, midges, and gnats). In virology , the International Committee on Taxonomy of Viruses 's virus classification includes fifteen taxomomic ranks to be applied for viruses , viroids and satellite nucleic acids : realm , subrealm , kingdom , subkingdom, phylum , subphylum , class, subclass, order, suborder, family, subfamily , genus, subgenus , and species. There are currently fourteen viral orders, each ending in

304-564: The precursor of the currently used International Code of Nomenclature for algae, fungi, and plants . In the first international Rules of botanical nomenclature from the International Botanical Congress of 1905, the word family ( familia ) was assigned to the rank indicated by the French famille , while order ( ordo ) was reserved for a higher rank, for what in the 19th century had often been named

323-502: The rank of family (see ordo naturalis , ' natural order '). In French botanical publications, from Michel Adanson 's Familles naturelles des plantes (1763) and until the end of the 19th century, the word famille (plural: familles ) was used as a French equivalent for this Latin ordo . This equivalence was explicitly stated in the Alphonse Pyramus de Candolle 's Lois de la nomenclature botanique (1868),

342-555: The suffix -virales . Proctotrupoidea Proctotrupoidea is a hymenopteran superfamily containing seven extant families, though others have been recognized in the past, most of these having been removed to a recently erected superfamily Diaprioidea . Of the remaining families, only Proctotrupidae contains a substantial number of species, with over 400 described. The others are small, often relictual groups. See links for individual families for details of life history and diversity. This Apocrita -related article

361-418: Was adopted by Systema Naturae 2000 and others. In botany , the ranks of subclass and suborder are secondary ranks pre-defined as respectively above and below the rank of order. Any number of further ranks can be used as long as they are clearly defined. The superorder rank is commonly used, with the ending -anae that was initiated by Armen Takhtajan 's publications from 1966 onwards. The order as

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