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86-814: An apple is a round, edible fruit produced by an apple tree ( Malus spp. , among them the domestic or orchard apple ; Malus domestica ). Apple trees are cultivated worldwide and are the most widely grown species in the genus Malus . The tree originated in Central Asia , where its wild ancestor, Malus sieversii , is still found. Apples have been grown for thousands of years in Eurasia and were introduced to North America by European colonists . Apples have religious and mythological significance in many cultures, including Norse , Greek , and European Christian tradition. Apples grown from seed tend to be very different from those of their parents, and

172-423: A fruit is the seed -bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering (see Fruit anatomy ). Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propagated using the movements of humans and other animals in a symbiotic relationship that is the means for seed dispersal for the one group and nutrition for

258-564: A risk factor for cardiovascular diseases . Fruit consumption is under preliminary research for the potential to improve nutrition and affect chronic diseases. Regular consumption of fruit is generally associated with reduced risks of several diseases and functional declines associated with aging. For food safety , the CDC recommends proper fruit handling and preparation to reduce the risk of food contamination and foodborne illness . Fresh fruits and vegetables should be carefully selected; at

344-750: A 6-day overlap period. There are four to seven pollination groups in apples, depending on climate: One cultivar can be pollinated by a compatible cultivar from the same group or close (A with A, or A with B, but not A with C or D). Cultivars vary in their yield and the ultimate size of the tree, even when grown on the same rootstock. Some cultivars, if left unpruned, grow very large—letting them bear more fruit, but making harvesting more difficult. Depending on tree density (number of trees planted per unit surface area), mature trees typically bear 40–200 kg (90–440 lb) of apples each year, though productivity can be close to zero in poor years. Apples are harvested using three-point ladders that are designed to fit amongst

430-399: A bed of straw, chaff, or mats with windfalls kept separately. Though methods like this will extend the availabity of reasonably fresh apples, without refrigeration their lifespan is limited. Even sturdy winter apple varieties will only keep well until December in cool climates. For longer storage medieval Europeans strung up cored and peeled apples to dry, either whole or sliced into rings. Of

516-450: A blackberry is an aggregate-accessory fruit, and an apple is a simple-accessory fruit. Seedlessness is an important feature of some fruits of commerce. Commercial cultivars of bananas and pineapples are examples of seedless fruits . Some cultivars of citrus fruits (especially grapefruit , mandarin oranges , navel oranges , satsumas ), table grapes , and of watermelons are valued for their seedlessness. In some species, seedlessness

602-423: A certain time, harmful bacteria may grow on them and increase the risk of foodborne illness. Shoot (botany) In everyday speech, shoots are often synonymous with stems. Stems, which are an integral component of shoots, provide an axis for buds, fruits, and leaves. Young shoots are often eaten by animals because the fibers in the new growth have not yet completed secondary cell wall development, making

688-512: A generic term for all fruit, including nuts . This can be compared to the 14th-century Middle English expression appel of paradis , meaning a banana . The apple is a deciduous tree, generally standing 2 to 4.5 metres (6 to 15 feet) tall in cultivation and up to 15 m (49 ft) in the wild, though more typically 2 to 10 m (6.5 to 33 ft). When cultivated, the size, shape and branch density are determined by rootstock selection and trimming method. Apple trees may naturally have

774-522: A head, is produced first. After fertilization , each flower in the cluster develops into a drupe; as the drupes expand, they develop as a connate organ, merging into a multiple fleshy fruit called a syncarp . Progressive stages of multiple flowering and fruit development can be observed on a single branch of the Indian mulberry, or noni . During the sequence of development, a progression of second, third, and more inflorescences are initiated in turn at

860-417: A long period of time and permitted secondary introgression of genes from other species into the open-pollinated seeds. Significant exchange with Malus sylvestris , the crabapple, resulted in populations of apples being more related to crabapples than to the more morphologically similar progenitor Malus sieversii . In strains without recent admixture the contribution of the latter predominates. The apple

946-644: A new plant some distance away from the parent. Other fruits have evolved flattened and elongated wings or helicopter-like blades, e.g., elm , maple , and tuliptree . This mechanism increases dispersal distance away from the parent via wind. Other wind-dispersed fruit have tiny " parachutes ", e.g., dandelion , milkweed , salsify . Coconut fruits can float thousands of miles in the ocean, thereby spreading their seeds. Other fruits that can disperse via water are nipa palm and screw pine . Some fruits have evolved propulsive mechanisms that fling seeds substantial distances – perhaps up to 100 m (330 ft) in

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1032-483: A number of organic and non-organic means. In 2010, the fruit's genome was sequenced as part of research on disease control and selective breeding in apple production. The word apple , whose Old English ancestor is æppel , is descended from the Proto-Germanic noun * aplaz , descended in turn from Proto-Indo-European * h₂ébōl . As late as the 17th century, the word also functioned as

1118-680: A number of different forms from a wide range of families, including carrot , parsnip , parsley , cumin . An aggregate fruit is also called an aggregation, or etaerio ; it develops from a single flower that presents numerous simple pistils . Each pistil contains one carpel ; together, they form a fruitlet. The ultimate (fruiting) development of the aggregation of pistils is called an aggregate fruit , etaerio fruit , or simply an etaerio . Different types of aggregate fruits can produce different etaerios, such as achenes, drupelets, follicles, and berries. Some other broadly recognized species and their etaerios (or aggregations) are: The pistils of

1204-532: A pantry and four to six weeks from the date of purchase in a refrigerator that maintains 4 to 0 °C (39 to 32 °F). Some varieties of apples (e.g. ' Granny Smith ' and ' Fuji ') have more than three times the storage life of others. Non-organic apples may be sprayed with a substance 1-methylcyclopropene blocking the apples' ethylene receptors, temporarily preventing them from ripening. Apple trees are susceptible to fungal and bacterial diseases, and to damage by insect pests. Many commercial orchards pursue

1290-447: A program of chemical sprays to maintain high fruit quality, tree health, and high yields. These prohibit the use of synthetic pesticides, though some older pesticides are allowed. Organic methods include, for instance, introducing its natural predator to reduce the population of a particular pest. A wide range of pests and diseases can affect the plant. Three of the more common diseases or pests are mildew, aphids, and apple scab. Among

1376-442: A prominent pointed terminal projection is said to be beaked . A fruit results from the fertilizing and maturing of one or more flowers. The gynoecium , which contains the stigma-style-ovary system, is centered in the flower-head, and it forms all or part of the fruit. Inside the ovary(ies) are one or more ovules . Here begins a complex sequence called double fertilization : a female gametophyte produces an egg cell for

1462-455: A rounded to erect crown with a dense canopy of leaves. The bark of the trunk is dark gray or gray-brown, but young branches are reddish or dark-brown with a smooth texture. Young twigs are covered in fine downy hairs; they become hairless when older. The buds are egg-shaped and dark red or purple in color; they range in size from 3 to 5 millimeters, but are usually less than 4 mm. The bud scales have very hairy edges. When emerging from

1548-404: A single flower with numerous pistils typically produces an aggregate fruit ; and the merging of several flowers, or a 'multiple' of flowers, results in a 'multiple' fruit. A simple fruit is further classified as either dry or fleshy. To distribute their seeds, dry fruits may split open and discharge their seeds to the winds, which is called dehiscence . Or the distribution process may rely upon

1634-408: A species has more than 100 alternative scientific names, or synonyms . In modern times, Malus pumila and Malus domestica are the two main names in use. M. pumila is the older name, but M. domestica has become much more commonly used starting in the 21st century, especially in the western world. Two proposals were made to make M. domestica a conserved name : the earlier proposal

1720-557: A strong minority following. Extremely sweet apples with barely any acid flavor are popular in Asia, especially the Indian subcontinent . Old cultivars are often oddly shaped, russeted, and grow in a variety of textures and colors. Some find them to have better flavor than modern cultivars, but they may have other problems that make them commercially unviable—low yield, disease susceptibility, poor tolerance for storage or transport, or just being

1806-974: Is a ripened ovary or carpel that contains seeds, e.g., an orange, pomegranate, tomato or a pumpkin. A nut is a type of fruit (and not a seed), and a seed is a ripened ovule . In culinary language, a fruit is the sweet- or not sweet- (even sour-) tasting produce of a specific plant (e.g., a peach, pear or lemon); nuts are hard, oily, non-sweet plant produce in shells ( hazelnut , acorn ). Vegetables , so-called, typically are savory or non-sweet produce ( zucchini , lettuce, broccoli, and tomato). but some may be sweet-tasting (sweet potato). Examples of botanically classified fruit that are typically called vegetables include cucumber , pumpkin , and squash (all are cucurbits ); beans , peanuts , and peas (all legumes ); and corn , eggplant , bell pepper (or sweet pepper), and tomato. Many spices are fruits, botanically speaking, including black pepper , chili pepper , cumin and allspice . In contrast, rhubarb

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1892-479: Is generally considered the center of origin for apples due to the genetic variability in specimens there. The wild ancestor of Malus domestica was Malus sieversii , found growing wild in the mountains of Central Asia in southern Kazakhstan , Kyrgyzstan , Tajikistan , and northwestern China . Cultivation of the species, most likely beginning on the forested flanks of the Tian Shan mountains, progressed over

1978-559: Is generally pale yellowish-white, though it can be pink, yellow or green. Important volatile compounds in apples that contribute to their scent and flavour include acetaldehyde , ethyl acetate , 1-butanal , ethanol , 2-methylbutanal, 3-methylbutanal , ethyl propionate , ethyl 2-methylpropionate, ethyl butyrate , ethyl 2-methyl butyrate, hexanal , 1-butanol , 3-methylbutyl acetate , 2-methylbutyl acetate, 1-propyl butyrate, ethyl pentanoate , amyl acetate , 2-methyl-1-butanol , trans-2-hexenal, ethyl hexanoate , hexanol . The apple as

2064-570: Is hard to distinguish in the archeological record between foraged wild apples and apple plantations. There is indirect evidence of apple cultivation in the third millennium BCE in the Middle East . There is direct evidence, apple cores, dated to the 10th century BCE from a Judean site between the Sinai and Negev. There was substantial apple production in European classical antiquity, and grafting

2150-417: Is often called a fruit when used in making pies , but the edible produce of rhubarb is actually the leaf stalk or petiole of the plant. Edible gymnosperm seeds are often given fruit names, e.g., ginkgo nuts and pine nuts . Botanically, a cereal grain, such as corn , rice , or wheat is a kind of fruit (termed a caryopsis ). However, the fruit wall is thin and fused to the seed coat, so almost all

2236-813: Is pressed for olive oil and similar processing is applied to other oil-bearing fruits and vegetables. Some fruits are available all year round, while others (such as blackberries and apricots in the UK) are subject to seasonal availability. Fruits are also used for socializing and gift-giving in the form of fruit baskets and fruit bouquets . Typically, many botanical fruits – "vegetables" in culinary parlance – (including tomato, green beans, leaf greens, bell pepper, cucumber, eggplant, okra, pumpkin, squash, zucchini) are bought and sold daily in fresh produce markets and greengroceries and carried back to kitchens, at home or restaurant, for preparation of meals. All fruits benefit from proper post-harvest care, and in many fruits,

2322-485: Is quite variable and may be nearly round, elongated, conical, or short and wide. The groundcolor of ripe apples is yellow, green, yellow-green or whitish yellow. The overcolor of ripe apples can be orange-red, pink-red, red, purple-red or brown-red. The overcolor amount can be 0–100%. The skin may be wholly or partly russeted , making it rough and brown. The skin is covered in a protective layer of epicuticular wax . The skin may also be marked with scattered dots. The flesh

2408-701: Is the responsibility of the Department of Environment, Food, and Rural Affairs, includes a collection of over 2,000 cultivars of apple tree in Kent. The University of Reading , which is responsible for developing the UK national collection database, provides access to search the national collection. The University of Reading's work is part of the European Cooperative Programme for Plant Genetic Resources of which there are 38 countries participating in

2494-522: Is the result of parthenocarpy , where fruits set without fertilization. Parthenocarpic fruit-set may (or may not) require pollination, but most seedless citrus fruits require a stimulus from pollination to produce fruit. Seedless bananas and grapes are triploids , and seedlessness results from the abortion of the embryonic plant that is produced by fertilization, a phenomenon known as stenospermocarpy , which requires normal pollination and fertilization. Variations in fruit structures largely depend on

2580-597: Is thought to have been domesticated 4,000–10,000 years ago in the Tian Shan mountains, and then to have travelled along the Silk Road to Europe, with hybridization and introgression of wild crabapples from Siberia ( M. baccata ), the Caucasus ( M. orientalis ), and Europe ( M. sylvestris ). Only the M. sieversii trees growing on the western side of the Tian Shan mountains contributed genetically to

2666-505: The pericarp (fruit wall) is fleshy at maturity are termed fleshy simple fruits . Types of fleshy simple fruits, (with examples) include: Berries are a type of simple fleshy fruit that issue from a single ovary. (The ovary itself may be compound, with several carpels.) The botanical term true berry includes grapes, currants, cucumbers, eggplants (aubergines), tomatoes, chili peppers, and bananas, but excludes certain fruits that are called "-berry" by culinary custom or by common usage of

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2752-457: The apple clearwing moth (red-belted clearwing) burrow through the bark and into the phloem of apple trees, potentially causing significant damage. There are more than 7,500 known cultivars (cultivated varieties) of apples. Cultivars vary in their yield and the ultimate size of the tree, even when grown on the same rootstock . Different cultivars are available for temperate and subtropical climates. The UK's National Fruit Collection, which

2838-603: The modes of dispersal applied to their seeds. Dispersal is achieved by wind or water, by explosive dehiscence , and by interactions with animals. Some fruits present their outer skins or shells coated with spikes or hooked burrs; these evolved either to deter would-be foragers from feeding on them or to serve to attach themselves to the hair, feathers, legs, or clothing of animals, thereby using them as dispersal agents. These plants are termed zoochorous ; common examples include cocklebur , unicorn plant , and beggarticks (or Spanish needle) . By developments of mutual evolution,

2924-484: The petals are rose-pink and fade to white or light pink when fully open with each flower 3-to-4-centimeter (1-to- 1 + 1 ⁄ 2 -inch) in diameter. The five-petaled flowers are group in an inflorescence consisting of a cyme with 3–7 flowers. The central flower of the inflorescence is called the "king bloom"; it opens first and can develop a larger fruit. Open apple blossoms are damaged by even brief exposures to temperatures −2 °C (28 °F) or less, although

3010-434: The raspberry are called drupelets because each pistil is like a small drupe attached to the receptacle. In some bramble fruits, such as blackberry , the receptacle, an accessory part, elongates and then develops as part of the fruit, making the blackberry an aggregate-accessory fruit. The strawberry is also an aggregate-accessory fruit, of which the seeds are contained in the achenes . Notably in all these examples,

3096-547: The "best" cultivars, showing the proliferation of new North American cultivars by the early 19th century. In the 20th century, irrigation projects in Eastern Washington began and allowed the development of the multibillion-dollar fruit industry, of which the apple is the leading product. Until the 20th century, farmers stored apples in frostproof cellars during the winter for their own use or for sale. Improved transportation of fresh apples by train and road replaced

3182-714: The "wrong" size. A few old cultivars are still produced on a large scale, but many have been preserved by home gardeners and farmers that sell directly to local markets. Many unusual and locally important cultivars with their own unique taste and appearance exist; apple conservation campaigns have sprung up around the world to preserve such local cultivars from extinction. In the United Kingdom, old cultivars such as ' Cox's Orange Pippin ' and ' Egremont Russet ' are still commercially important even though by modern standards they are low yielding and susceptible to disease. 2022, millions of tonnes Fruit In botany ,

3268-605: The ' Cosmic Crisp ' cultivar in 2017. The third most grown apple cultivar in the United States is the ' Honeycrisp ', released by the University of Minnesota program in 1991. Unusually for a popular cultivar, the 'Honeycrisp' is not directly related to another popular apple cultivar but instead to two unsuccessful cultivars. In Europe there are also many breeding programs such as the Julius Kühn-Institut ,

3354-412: The (deposited) pollen through the stigma down the style into the ovary to the ovule. Two sperm are transferred from the pollen to a megagametophyte. Within the megagametophyte, one sperm unites with the egg, forming a zygote , while the second sperm enters the central cell forming the endosperm mother cell, which completes the double fertilization process. Later, the zygote will give rise to the embryo of

3440-812: The 15th century and later went through several cycles of popularity and decline throughout the world. The majority of the rootstocks used to control size in apples were developed in England in the early 1900s. The East Malling Research Station conducted extensive research into rootstocks, and their rootstocks are given an "M" prefix to designate their origin. Rootstocks marked with an "MM" prefix are Malling-series cultivars later crossed with trees of ' Northern Spy ' in Merton, England . Most new apple cultivars originate as seedlings, which either arise by chance or are bred by deliberately crossing cultivars with promising characteristics. The words "seedling", "pippin", and "kernel" in

3526-588: The German federal research center for cultivated plants. Commercially popular apple cultivars are soft but crisp. Other desirable qualities in modern commercial apple breeding are a colorful skin, absence of russeting , ease of shipping, lengthy storage ability, high yields, disease resistance, common apple shape, and developed flavor. Modern apples are generally sweeter than older cultivars, as popular tastes in apples have varied over time. Most North Americans and Europeans favor sweet, subacid apples, but tart apples have

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3612-538: The Malus/Pyrus work group. The UK's national fruit collection database contains much information on the characteristics and origin of many apples, including alternative names for what is essentially the same "genetic" apple cultivar. Most of these cultivars are bred for eating fresh (dessert apples), though some are cultivated specifically for cooking ( cooking apples ) or producing cider . Cider apples are typically too tart and astringent to eat fresh, but they give

3698-408: The attachment of other floral parts – there are parts (including petals, sepals, and stamens) that fuse with the ovary and ripen with it. For such a case, when floral parts other than the ovary form a significant part of the fruit that develops, it is called an accessory fruit . Examples of accessory fruits include apple, rose hip, strawberry, and pineapple. Because several parts of the flower besides

3784-427: The base of the flower. The seeds are egg- to pear-shaped and may be colored from light brown or tan to a very dark brown, often with red shades or even purplish-black. They may have a blunt or sharp point. The five sepals remain attached and stand out from the surface of the apple. The size of the fruit varies widely between cultivars, but generally has a diameter between 2.5 and 12 cm (1 and 5 in). The shape

3870-478: The beverage a rich flavor that dessert apples cannot. In the United States there are many apple breeding programs associated with universities. Cornell University has had a program operating since 1880 in Geneva, New York . Among their recent well known apples is the 'SnapDragon' cultivar released in 2013. In the west Washington State University started a program to support their apple industry in 1994 and released

3956-691: The branches. Trees grafted on dwarfing rootstocks bear about 10–80 kg (20–180 lb) of fruit per year. Some farms with apple orchards open them to the public so consumers can pick their own apples. Crops ripen at different times of the year according to the cultivar. Cultivar that yield their crop in the summer include ' Sweet Bough ' and 'Duchess'; fall producers include 'Blenheim'; winter producers include 'King', ' Swayzie ', and 'Tolman Sweet'. Commercially, apples can be stored for months in controlled atmosphere chambers. Apples are commonly stored in chambers with lowered concentrations of oxygen to reduce respiration and slow softening and other changes if

4042-418: The buds, the leaves are convolute , meaning that their edges overlap each other. Leaves can be simple ovals ( elliptic ), medium or wide in width, somewhat egg-shaped with the wider portion toward their base ( ovate ), or even with sides that are more parallel to each other instead of curved ( oblong ) with a narrow pointed end. The edges have broadly-angled teeth, but do not have lobes. The top surface of

4128-890: The case of the sandbox tree – via explosive dehiscence or other such mechanisms (see impatiens and squirting cucumber ). A cornucopia of fruits – fleshy (simple) fruits from apples to berries to watermelon; dry (simple) fruits including beans and rice and coconuts; aggregate fruits including strawberries, raspberries, blackberries, pawpaw; and multiple fruits such as pineapple, fig, mulberries – are commercially valuable as human food. They are eaten both fresh and as jams, marmalade and other fruit preserves . They are used extensively in manufactured and processed foods (cakes, cookies, baked goods, flavorings, ice cream, yogurt, canned vegetables, frozen vegetables and meals) and beverages such as fruit juices and alcoholic beverages (brandy, fruit beer , wine). Spices like vanilla, black pepper, paprika, and allspice are derived from berries. Olive fruit

4214-452: The case when a triploid plant can produce a seed (apples are an example), it occurs infrequently, and seedlings rarely survive. Because apples are not true breeders when planted as seeds, propagation usually involves grafting of cuttings. The rootstock used for the bottom of the graft can be selected to produce trees of a large variety of sizes, as well as changing the winter hardiness, insect and disease resistance, and soil preference of

4300-409: The characteristics of the parent. This is because seedling apples are " extreme heterozygotes ". Rather than resembling their parents, seedlings are all different from each other and from their parents. Triploid cultivars have an additional reproductive barrier in that three sets of chromosomes cannot be divided evenly during meiosis, yielding unequal segregation of the chromosomes (aneuploids). Even in

4386-490: The decay and degradation of the fruit to expose the seeds; or it may rely upon the eating of fruit and excreting of seeds by frugivores – both are called indehiscence . Fleshy fruits do not split open, but they also are indehiscent and they may also rely on frugivores for distribution of their seeds. Typically, the entire outer layer of the ovary wall ripens into a potentially edible pericarp . Types of dry simple fruits, (with examples) include: Fruits in which part or all of

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4472-580: The domesticated apple, not the isolated population on the eastern side. Chinese soft apples, such as M. asiatica and M. prunifolia , have been cultivated as dessert apples for more than 2,000 years in China. These are thought to be hybrids between M. baccata and M. sieversii in Kazakhstan. Among the traits selected for by human growers are size, fruit acidity, color, firmness, and soluble sugar. Unusually for domesticated fruits,

4558-401: The edible grain-fruit is actually a seed. The outer layer, often edible, of most fruits is called the pericarp . Typically formed from the ovary, it surrounds the seeds; in some species, however, other structural tissues contribute to or form the edible portion. The pericarp may be described in three layers from outer to inner, i.e., the epicarp , mesocarp and endocarp . Fruit that bears

4644-477: The fleshy produce of fruits typically appeals to hungry animals, such that the seeds contained within are taken in, carried away, and later deposited (i.e., defecated ) at a distance from the parent plant. Likewise, the nutritious, oily kernels of nuts typically motivate birds and squirrels to hoard them, burying them in soil to retrieve later during the winter of scarcity; thereby, uneaten seeds are sown effectively under natural conditions to germinate and grow

4730-481: The flowering each season, apple growers often utilize pollinators to carry pollen. Honey bees are most commonly used. Orchard mason bees are also used as supplemental pollinators in commercial orchards. Bumblebee queens are sometimes present in orchards, but not usually in sufficient number to be significant pollinators. Cultivars are sometimes classified by the day of peak bloom in the average 30-day blossom period, with pollinizers selected from cultivars within

4816-429: The fruit develops from a single flower, with numerous pistils. A multiple fruit is formed from a cluster of flowers, (a 'multiple' of flowers) – also called an inflorescence . Each ('smallish') flower produces a single fruitlet, which, as all develop, all merge into one mass of fruit. Examples include pineapple , fig , mulberry , Osage orange , and breadfruit . An inflorescence (a cluster) of white flowers, called

4902-482: The fruit is already fully ripe. The gas ethylene is used by plants as a hormone which promotes ripening, decreasing the time an apple can be stored. For storage longer than about six months the apples are picked earlier, before full ripeness, when ethylene production by the fruit is low. However, in many varieties this increases their sensitivity to carbon dioxide , which also must be controlled. For home storage, most culitvars of apple can be stored for three weeks in

4988-416: The fruits develop, but they are not evolutionarily relevant as diverse plant taxa may be in the same group. While the section of a fungus that produces spores is called a fruiting body, fungi are members of the fungi kingdom and not of the plant kingdom . Simple fruits are the result of the ripening-to-fruit of a simple or compound ovary in a single flower with a single pistil . In contrast,

5074-419: The head of the branch or stem. Fruits may incorporate tissues derived from other floral parts besides the ovary, including the receptacle, hypanthium, petals, or sepals. Accessory fruits occur in all three classes of fruit development – simple, aggregate, and multiple. Accessory fruits are frequently designated by the hyphenated term showing both characters. For example, a pineapple is a multiple-accessory fruit,

5160-496: The latter term meaning the fleshy part is derived not from the plant's ovaries but from the receptacle that holds the ovaries . Numerous dry achenes are attached to the outside of the fruit-flesh; they appear to be seeds but each is actually an ovary of a flower, with a seed inside. Schizocarps are dry fruits, though some appear to be fleshy. They originate from syncarpous ovaries but do not actually dehisce ; rather, they split into segments with one or more seeds. They include

5246-413: The leaves are glabrescent , almost hairless, while the undersides are densely covered in fine hairs. The leaves are attached alternately by short leaf stems 1-to-3.5 cm ( 1 ⁄ 2 -to- 1 + 1 ⁄ 2  in) long. Blossoms are produced in spring simultaneously with the budding of the leaves and are produced on spurs and some long shoots . When the flower buds first begin to open

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5332-484: The majority of flowers and fruit. A similar pattern occurs in some conifers and in Ginkgo , although the "short shoots" of some genera such as Picea are so small that they can be mistaken for part of the leaf that they have produced. A related phenomenon is seasonal heterophylly , which involves visibly different leaves from spring growth and later lammas growth . Whereas spring growth mostly comes from buds formed

5418-685: The many Old World plants that the Spanish introduced to Chiloé Archipelago in the 16th century, apple trees became particularly well adapted. Apples were introduced to North America by colonists in the 17th century, and the first named apple cultivar was introduced in Boston by Reverend William Blaxton in 1640. The only apples native to North America are crab apples . Apple cultivars brought as seed from Europe were spread along Native American trade routes, as well as being cultivated on colonial farms. An 1845 United States apples nursery catalogue sold 350 of

5504-428: The most serious disease problems is a bacterial disease called fireblight , and three fungal diseases: Gymnosporangium rust, black spot , and bitter rot . Codling moths , and the apple maggots of fruit flies, cause serious damage to apple fruits, making them unsaleable. Young apple trees are also prone to mammal pests like mice and deer, which feed on the soft bark of the trees, especially in winter. The larvae of

5590-623: The name of an apple cultivar suggest that it originated as a seedling. Apples can also form bud sports (mutations on a single branch). Some bud sports turn out to be improved strains of the parent cultivar. Some differ sufficiently from the parent tree to be considered new cultivars. Apples have been acclimatized in Ecuador at very high altitudes, where they can often, with the needed factors, provide crops twice per year because of constant temperate conditions year-round. Apples are self-incompatible; they must cross-pollinate to develop fruit. During

5676-455: The necessity for storage. Controlled atmosphere facilities are used to keep apples fresh year-round. Controlled atmosphere facilities use high humidity, low oxygen, and controlled carbon dioxide levels to maintain fruit freshness. They were first researched at Cambridge University in the 1920s and first used in the United States in the 1950s. Many apples grow readily from seeds. However, apples must be propagated asexually to obtain cuttings with

5762-445: The number of fertilized ovules. The pericarp typically is differentiated into two or three distinct layers; these are called the exocarp (outer layer, also called epicarp), mesocarp (middle layer), and endocarp (inner layer). In some fruits, the sepals , petals , stamens or the style of the flower fall away as the fleshy fruit ripens. However, for simple fruits derived from an inferior ovary – i.e., one that lies below

5848-473: The other; humans, and many other animals, have become dependent on fruits as a source of food. Consequently, fruits account for a substantial fraction of the world's agricultural output, and some (such as the apple and the pomegranate ) have acquired extensive cultural and symbolic meanings. In common language and culinary usage, fruit normally means the seed-associated fleshy structures (or produce) of plants that typically are sweet (or sour) and edible in

5934-437: The ovary may contribute to the structure of a fruit, it is important to understand how a particular fruit forms. There are three general modes of fruit development: Consistent with the three modes of fruit development, plant scientists have classified fruits into three main groups: simple fruits, aggregate fruits, and multiple (or composite) fruits. The groupings reflect how the ovary and other flower organs are arranged and how

6020-412: The overwintering wood and buds are hardy down to −40 °C (−40 °F). The fruit is a pome that matures in late summer or autumn . The true fruits or carpels are the harder interior chambers inside the apple's core. There are usually five carpels inside an apple, but there may be as few as three. Each of the chambers contains one or two seeds. The edible flesh is formed from the receptacle at

6106-486: The pears, apples, and quinces were combined into one genus that he named Pyrus and he named the apple as Pyrus malus . This was widely accepted, however the botanist Philip Miller published an alternate classification in The Gardeners Dictionary with the apple species separated from Pyrus in 1754. He did not clearly indicate that by Malus pumila he meant the domesticated apple. Nonetheless, it

6192-614: The plant hormone ethylene causes ripening . Therefore, maintaining most fruits in an efficient cold chain is optimal for post-harvest storage, with the aim of extending and ensuring shelf life. Various culinary fruits provide significant amounts of fiber and water, and many are generally high in vitamin C . An overview of numerous studies showed that fruits (e.g., whole apples or whole oranges) are satisfying (filling) by simply eating and chewing them. The dietary fiber consumed in eating fruit promotes satiety , and may help to control body weight and aid reduction of blood cholesterol ,

6278-401: The purpose of fertilization. (A female gametophyte is called a megagametophyte , and also called the embryo sac .) After double fertilization, the ovules will become seeds. Ovules are fertilized in a process that starts with pollination , which is the movement of pollen from the stamens to the stigma-style-ovary system within the flower-head. After pollination, a pollen tube grows from

6364-403: The raw state, such as apples, bananas, grapes, lemons, oranges, and strawberries. In botanical usage, the term fruit also includes many structures that are not commonly called as such in everyday language, such as nuts, bean pods, corn kernels, tomatoes, and wheat grains. Many common language terms used for fruit and seeds differ from botanical classifications. For example, in botany, a fruit

6450-685: The resultant fruit frequently lacks desired characteristics. For commercial purposes, including botanical evaluation, apple cultivars are propagated by clonal grafting onto rootstocks . Apple trees grown without rootstocks tend to be larger and much slower to fruit after planting. Rootstocks are used to control the speed of growth and the size of the resulting tree, allowing for easier harvesting. There are more than 7,500 cultivars of apples . Different cultivars are bred for various tastes and uses, including cooking , eating raw, and cider or apple juice production. Trees and fruit are prone to fungal , bacterial, and pest problems, which can be controlled by

6536-520: The resulting tree. Dwarf rootstocks can be used to produce very small trees (less than 3.0 m or 10 ft high at maturity), which bear fruit many years earlier in their life cycle than full size trees, and are easier to harvest. Dwarf rootstocks for apple trees can be traced as far back as 300 BCE, to the area of Persia and Asia Minor . Alexander the Great sent samples of dwarf apple trees to Aristotle 's Lyceum . Dwarf rootstocks became common by

6622-401: The seed, and the endosperm mother cell will give rise to endosperm , a nutritive tissue used by the embryo. As the ovules develop into seeds, the ovary begins to ripen and the ovary wall, the pericarp , may become fleshy (as in berries or drupes ), or it may form a hard outer covering (as in nuts). In some multi-seeded fruits, the extent to which a fleshy structure develops is proportional to

6708-730: The store, they should not be damaged or bruised; and precut pieces should be refrigerated or surrounded by ice. All fruits and vegetables should be rinsed before eating. This recommendation also applies to produce with rinds or skins that are not eaten. It should be done just before preparing or eating to avoid premature spoilage. Fruits and vegetables should be kept separate from raw foods like meat, poultry, and seafood, as well as from utensils that have come in contact with raw foods. Fruits and vegetables that are not going to be cooked should be thrown away if they have touched raw meat, poultry, seafood, or eggs. All cut, peeled, or cooked fruits and vegetables should be refrigerated within two hours. After

6794-417: The term – such as strawberries and raspberries. Berries may be formed from one or more carpels (i.e., from the simple or compound ovary) from the same, single flower. Seeds typically are embedded in the fleshy interior of the ovary. Examples include: The strawberry, regardless of its appearance, is classified as a dry, not a fleshy fruit. Botanically, it is not a berry ; it is an aggregate-accessory fruit,

6880-592: The wild M. sieversii origin is only slightly smaller than the modern domesticated apple. At the Sammardenchia-Cueis site near Udine in Northeastern Italy, seeds from some form of apples have been found in material carbon dated to between 6570 and 5684 BCE. Genetic analysis has not yet been successfully used to determine whether such ancient apples were wild Malus sylvestris or Malus domesticus containing Malus sieversii ancestry. It

6966-415: The young shoots softer and easier to chew and digest. As shoots grow and age, the cells develop secondary cell walls that have a hard and tough structure. Some plants (e.g. bracken ) produce toxins that make their shoots inedible or less palatable. Many woody plants have distinct short shoots and long shoots . In some angiosperms , the short shoots, also called spur shoots or fruit spurs , produce

7052-429: Was based on the diploid cultivar ' Golden Delicious '. However, this first whole genome sequence contained several errors, in part owing to the high degree of heterozygosity in diploid apples which, in combination with an ancient genome duplication, complicated the assembly. Recently, double- and trihaploid individuals have been sequenced, yielding whole genome sequences of higher quality. The first whole genome assembly

7138-453: Was certainly known then. Grafting is an essential part of modern domesticated apple production, to be able to propagate the best cultivars; it is unclear when apple tree grafting was invented. The Roman writer Pliny the Elder describes a method of storage for apples from his time in the 1st century. He says they should be placed in a room with good air circulation from a north facing window on

7224-465: Was estimated to contain around 57,000 genes, though the more recent genome sequences support estimates between 42,000 and 44,700 protein-coding genes. The availability of whole genome sequences has provided evidence that the wild ancestor of the cultivated apple most likely is Malus sieversii . Re-sequencing of multiple accessions has supported this, while also suggesting extensive introgression from Malus sylvestris following domestication. Central Asia

7310-637: Was used as such by many botanists. When Moritz Balthasar Borkhausen published his scientific description of the apple in 1803 it may have been a new combination of P. malus var. domestica , but this was not directly referenced by Borkhausen. The earliest use of var. domestica for the apple was by Georg Adolf Suckow in 1786. Apples are diploid , with two sets of chromosomes per cell (though triploid cultivars, with three sets, are not uncommon), have 17 chromosomes and an estimated genome size of approximately 650 Mb. Several whole genome sequences have been completed and made available. The first one in 2010

7396-743: Was voted down by the Committee for Vascular Plants of the IAPT in 2014, but in April 2017 the Committee decided, with a narrow majority, that the newly popular name should be conserved. The General Committee of the IAPT decided in June 2017 to approve this change, officially conserving M. domestica . Nevertheless, some works published after 2017 still use M. pumila as the correct name , under an alternate taxonomy. When first classified by Linnaeus in 1753,

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