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98-628: The Annisquam River is a tidal, salt-water estuary in the Annisquam neighborhood of Gloucester, Massachusetts , connecting Annisquam Harbor on the north to Gloucester Harbor on the south. The segment between Gloucester Harbor and the Newburyport/Rockport Line bridge is also known as the Blynman Canal . The estuary is about 4.5 miles (7.2 km) long, navigable, and open to the ocean at both ends. Its surface area

196-409: A cell wall . Newly dead animals may be covered by an exoskeleton . Fragmentation processes, which break through these protective layers, accelerate the rate of microbial decomposition. Animals fragment detritus as they hunt for food, as does passage through the gut. Freeze-thaw cycles and cycles of wetting and drying also fragment dead material. The chemical alteration of the dead organic matter

294-495: A food chain . Real systems are much more complex than this—organisms will generally feed on more than one form of food, and may feed at more than one trophic level. Carnivores may capture some prey that is part of a plant-based trophic system and others that are part of a detritus-based trophic system (a bird that feeds both on herbivorous grasshoppers and earthworms, which consume detritus). Real systems, with all these complexities, form food webs rather than food chains which present

392-619: A habitat . Ecosystem ecology is the "study of the interactions between organisms and their environment as an integrated system". The size of ecosystems can range up to ten orders of magnitude , from the surface layers of rocks to the surface of the planet. The Hubbard Brook Ecosystem Study started in 1963 to study the White Mountains in New Hampshire . It was the first successful attempt to study an entire watershed as an ecosystem. The study used stream chemistry as

490-609: A central role over a wide range, for example, in the slow development of soil from bare rock and the faster recovery of a community from disturbance . Disturbance also plays an important role in ecological processes. F. Stuart Chapin and coauthors define disturbance as "a relatively discrete event in time that removes plant biomass". This can range from herbivore outbreaks, treefalls, fires, hurricanes, floods, glacial advances , to volcanic eruptions . Such disturbances can cause large changes in plant, animal and microbe populations, as well as soil organic matter content. Disturbance

588-507: A critical role in global nutrient cycling and ecosystem function. Phosphorus enters ecosystems through weathering . As ecosystems age this supply diminishes, making phosphorus-limitation more common in older landscapes (especially in the tropics). Calcium and sulfur are also produced by weathering, but acid deposition is an important source of sulfur in many ecosystems. Although magnesium and manganese are produced by weathering, exchanges between soil organic matter and living cells account for

686-517: A faster recovery. More severe and more frequent disturbance result in longer recovery times. From one year to another, ecosystems experience variation in their biotic and abiotic environments. A drought , a colder than usual winter, and a pest outbreak all are short-term variability in environmental conditions. Animal populations vary from year to year, building up during resource-rich periods and crashing as they overshoot their food supply. Longer-term changes also shape ecosystem processes. For example,

784-454: A form that can be readily used by plants and microbes. Ecosystems provide a variety of goods and services upon which people depend, and may be part of. Ecosystem goods include the "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like

882-427: A function-based typology has been proposed to leverage the strengths of these different approaches into a unified system. Human activities are important in almost all ecosystems. Although humans exist and operate within ecosystems, their cumulative effects are large enough to influence external factors like climate. Ecosystems provide a variety of goods and services upon which people depend. Ecosystem goods include

980-698: A general level, for example, tropical forests , temperate grasslands , and arctic tundra . There can be any degree of subcategories among ecosystem types that comprise a biome, e.g., needle-leafed boreal forests or wet tropical forests. Although ecosystems are most commonly categorized by their structure and geography, there are also other ways to categorize and classify ecosystems such as by their level of human impact (see anthropogenic biome ), or by their integration with social processes or technological processes or their novelty (e.g. novel ecosystem ). Each of these taxonomies of ecosystems tends to emphasize different structural or functional properties. None of these

1078-573: A harsh environment for organisms. Sediment often settles in intertidal mudflats which are extremely difficult to colonize. No points of attachment exist for algae , so vegetation based habitat is not established. Sediment can also clog feeding and respiratory structures of species, and special adaptations exist within mudflat species to cope with this problem. Lastly, dissolved oxygen variation can cause problems for life forms. Nutrient-rich sediment from human-made sources can promote primary production life cycles, perhaps leading to eventual decay removing

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1176-672: A means of monitoring ecosystem properties, and developed a detailed biogeochemical model of the ecosystem. Long-term research at the site led to the discovery of acid rain in North America in 1972. Researchers documented the depletion of soil cations (especially calcium) over the next several decades. Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Studies can be carried out at

1274-528: A more important role in moving nutrients around. This can be especially important as the soil thaws in the spring, creating a pulse of nutrients that become available. Decomposition rates are low under very wet or very dry conditions. Decomposition rates are highest in wet, moist conditions with adequate levels of oxygen. Wet soils tend to become deficient in oxygen (this is especially true in wetlands ), which slows microbial growth. In dry soils, decomposition slows as well, but bacteria continue to grow (albeit at

1372-529: A number of coastal water bodies such as coastal lagoons and brackish seas. A more comprehensive definition of an estuary is "a semi-enclosed body of water connected to the sea as far as the tidal limit or the salt intrusion limit and receiving freshwater runoff; however the freshwater inflow may not be perennial, the connection to the sea may be closed for part of the year and tidal influence may be negligible". This broad definition also includes fjords , lagoons , river mouths , and tidal creeks . An estuary

1470-406: A number of common, non random properties in the topology of their network. The carbon and nutrients in dead organic matter are broken down by a group of processes known as decomposition. This releases nutrients that can then be re-used for plant and microbial production and returns carbon dioxide to the atmosphere (or water) where it can be used for photosynthesis. In the absence of decomposition,

1568-421: A process known as denitrification . Mycorrhizal fungi which are symbiotic with plant roots, use carbohydrates supplied by the plants and in return transfer phosphorus and nitrogen compounds back to the plant roots. This is an important pathway of organic nitrogen transfer from dead organic matter to plants. This mechanism may contribute to more than 70 Tg of annually assimilated plant nitrogen, thereby playing

1666-696: A significant portion of ecosystem fluxes. Potassium is primarily cycled between living cells and soil organic matter. Biodiversity plays an important role in ecosystem functioning. Ecosystem processes are driven by the species in an ecosystem, the nature of the individual species, and the relative abundance of organisms among these species. Ecosystem processes are the net effect of the actions of individual organisms as they interact with their environment. Ecological theory suggests that in order to coexist, species must have some level of limiting similarity —they must be different from one another in some fundamental way, otherwise, one species would competitively exclude

1764-408: A slower rate) even after soils become too dry to support plant growth. Ecosystems are dynamic entities. They are subject to periodic disturbances and are always in the process of recovering from past disturbances. When a perturbation occurs, an ecosystem responds by moving away from its initial state. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance,

1862-464: A small effect on ecosystem function. Ecologically distinct species, on the other hand, have a much larger effect. Similarly, dominant species have a large effect on ecosystem function, while rare species tend to have a small effect. Keystone species tend to have an effect on ecosystem function that is disproportionate to their abundance in an ecosystem. An ecosystem engineer is any organism that creates, significantly modifies, maintains or destroys

1960-526: A system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Biomes are general classes or categories of ecosystems. However, there

2058-408: A type of ecosystem in some estuaries that have been negatively impacted by eutrophication. Cordgrass vegetation dominates the salt marsh landscape. Excess nutrients allow the plants to grow at greater rates in above ground biomass, however less energy is allocated to the roots since nutrients is abundant. This leads to a lower biomass in the vegetation below ground which destabilizes the banks of

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2156-522: A variety of scales, ranging from whole-ecosystem studies to studying microcosms or mesocosms (simplified representations of ecosystems). American ecologist Stephen R. Carpenter has argued that microcosm experiments can be "irrelevant and diversionary" if they are not carried out in conjunction with field studies done at the ecosystem scale. In such cases, microcosm experiments may fail to accurately predict ecosystem-level dynamics. Biomes are general classes or categories of ecosystems. However, there

2254-563: A well-mixed water column and the disappearance of the vertical salinity gradient . The freshwater-seawater boundary is eliminated due to the intense turbulent mixing and eddy effects . The lower reaches of Delaware Bay and the Raritan River in New Jersey are examples of vertically homogeneous estuaries. Inverse estuaries occur in dry climates where evaporation greatly exceeds the inflow of freshwater. A salinity maximum zone

2352-436: A wholly marine embayment to any of the other estuary types. The most important variable characteristics of estuary water are the concentration of dissolved oxygen, salinity and sediment load. There is extreme spatial variability in salinity, with a range of near-zero at the tidal limit of tributary rivers to 3.4% at the estuary mouth. At any one point, the salinity will vary considerably over time and seasons, making it

2450-477: A wide effect on the surrounding water bodies.  In turn, this can decrease fishing industry sales in one area and across the country. Production in 2016 from recreational and commercial fishing contributes billions of dollars to the United States' gross domestic product (GDP). A decrease in production within this industry can affect any of the 1.7 million people the fishing industry employs yearly across

2548-597: Is 1.9 square miles (4.9 km). The name "Annisquam" comes from an Algonquian term meaning "top of the rock, containing <wanashque> , "on top of", and <-ompsk> , "rock". The first European settlement in Annisquam was established in 1631. The river was dredged by the United States Army Corps of Engineers in 1965 to create a channel some 8 feet (2.4 m) deep, and 60 to 200 feet (18 to 61 m) wide. Annisquam Harbor Light stands on

2646-455: Is a dynamic ecosystem having a connection to the open sea through which the sea water enters with the rhythm of the tides . The effects of tides on estuaries can show nonlinear effects on the movement of water which can have important impacts on the ecosystem and waterflow. The seawater entering the estuary is diluted by the fresh water flowing from rivers and streams. The pattern of dilution varies between different estuaries and depends on

2744-458: Is an international synthesis by over 1000 of the world's leading biological scientists that analyzes the state of the Earth's ecosystems and provides summaries and guidelines for decision-makers. The report identified four major categories of ecosystem services: provisioning, regulating, cultural and supporting services. It concludes that human activity is having a significant and escalating impact on

2842-442: Is consumed by animals while still alive and enters the plant-based trophic system. After plants and animals die, the organic matter contained in them enters the detritus-based trophic system. Ecosystem respiration is the sum of respiration by all living organisms (plants, animals, and decomposers) in the ecosystem. Net ecosystem production is the difference between gross primary production (GPP) and ecosystem respiration. In

2940-423: Is controlled by internal factors. Therefore, internal factors not only control ecosystem processes but are also controlled by them. Ecosystems are dynamic entities—they are subject to periodic disturbances and are always in the process of recovering from some past disturbance. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance, is termed its resistance . The capacity of

3038-513: Is derived from the Latin word aestuarium meaning tidal inlet of the sea, which in itself is derived from the term aestus , meaning tide. There have been many definitions proposed to describe an estuary. The most widely accepted definition is: "a semi-enclosed coastal body of water, which has a free connection with the open sea, and within which seawater is measurably diluted with freshwater derived from land drainage". However, this definition excludes

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3136-569: Is followed by succession, a "directional change in ecosystem structure and functioning resulting from biotically driven changes in resource supply." The frequency and severity of disturbance determine the way it affects ecosystem function. A major disturbance like a volcanic eruption or glacial advance and retreat leave behind soils that lack plants, animals or organic matter. Ecosystems that experience such disturbances undergo primary succession . A less severe disturbance like forest fires, hurricanes or cultivation result in secondary succession and

3234-411: Is formed, and both riverine and oceanic water flow close to the surface towards this zone. This water is pushed downward and spreads along the bottom in both the seaward and landward direction. Examples of an inverse estuary are Spencer Gulf , South Australia, Saloum River and Casamance River , Senegal. Estuary type varies dramatically depending on freshwater input, and is capable of changing from

3332-554: Is governed by three sets of factors—the physical environment (temperature, moisture, and soil properties), the quantity and quality of the dead material available to decomposers, and the nature of the microbial community itself. Temperature controls the rate of microbial respiration; the higher the temperature, the faster the microbial decomposition occurs. Temperature also affects soil moisture, which affects decomposition. Freeze-thaw cycles also affect decomposition—freezing temperatures kill soil microorganisms, which allows leaching to play

3430-566: Is less restricted, and there is a slow but steady exchange of water between the estuary and the ocean. Fjord-type estuaries can be found along the coasts of Alaska , the Puget Sound region of western Washington state , British Columbia , eastern Canada, Greenland , Iceland , New Zealand, and Norway. These estuaries are formed by subsidence or land cut off from the ocean by land movement associated with faulting , volcanoes , and landslides . Inundation from eustatic sea-level rise during

3528-405: Is no clear distinction between biomes and ecosystems. Biomes are always defined at a very general level. Ecosystems can be described at levels that range from very general (in which case the names are sometimes the same as those of biomes) to very specific, such as "wet coastal needle-leafed forests". Biomes vary due to global variations in climate . Biomes are often defined by their structure: at

3626-516: Is no clear distinction between biomes and ecosystems. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of the definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Biotic factors of the ecosystem are living things; such as plants, animals, and bacteria, while abiotic are non-living components; such as water, soil and atmosphere. Plants allow energy to enter

3724-531: Is primarily achieved through bacterial and fungal action. Fungal hyphae produce enzymes that can break through the tough outer structures surrounding dead plant material. They also produce enzymes that break down lignin , which allows them access to both cell contents and the nitrogen in the lignin. Fungi can transfer carbon and nitrogen through their hyphal networks and thus, unlike bacteria, are not dependent solely on locally available resources. Decomposition rates vary among ecosystems. The rate of decomposition

3822-485: Is termed its resistance . The capacity of a system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Resilience thinking also includes humanity as an integral part of the biosphere where we are dependent on ecosystem services for our survival and must build and maintain their natural capacities to withstand shocks and disturbances. Time plays

3920-633: Is the Colorado River Delta in Mexico, historically covered with marshlands and forests, but now essentially a salt flat. Ecosystem An ecosystem (or ecological system ) is a system that environments and their organisms form through their interaction. The biotic and abiotic components are linked together through nutrient cycles and energy flows. Ecosystems are controlled by external and internal factors . External factors such as climate , parent material which forms

4018-422: Is the "best" classification. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of the definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Different approaches to ecological classifications have been developed in terrestrial, freshwater and marine disciplines, and

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4116-646: Is the whitefish species from the European Alps . Eutrophication reduced the oxygen levels in their habitats so greatly that whitefish eggs could not survive, causing local extinctions. However, some animals, such as carnivorous fish, tend to do well in nutrient-enriched environments and can benefit from eutrophication. This can be seen in populations of bass or pikes. Eutrophication can affect many marine habitats which can lead to economic consequences. The commercial fishing industry relies upon estuaries for approximately 68 percent of their catch by value because of

4214-748: The Holocene Epoch has also contributed to the formation of these estuaries. There are only a small number of tectonically produced estuaries; one example is the San Francisco Bay , which was formed by the crustal movements of the San Andreas Fault system causing the inundation of the lower reaches of the Sacramento and San Joaquin rivers . In this type of estuary, river output greatly exceeds marine input and tidal effects have minor importance. Freshwater floats on top of

4312-473: The Holocene epoch with the flooding of river-eroded or glacially scoured valleys when the sea level began to rise about 10,000–12,000 years ago. Estuaries are typically classified according to their geomorphological features or to water-circulation patterns. They can have many different names, such as bays , harbors , lagoons , inlets , or sounds , although some of these water bodies do not strictly meet

4410-610: The Mandovi estuary in Goa during the monsoon period. As tidal forcing increases, river output becomes less than the marine input. Here, current induced turbulence causes mixing of the whole water column such that salinity varies more longitudinally rather than vertically, leading to a moderately stratified condition. Examples include the Chesapeake Bay and Narragansett Bay . Tidal mixing forces exceed river output, resulting in

4508-640: The Mid-Atlantic coast, and Galveston Bay and Tampa Bay along the Gulf Coast . Bar-built estuaries are found in a place where the deposition of sediment has kept pace with rising sea levels so that the estuaries are shallow and separated from the sea by sand spits or barrier islands. They are relatively common in tropical and subtropical locations. These estuaries are semi-isolated from ocean waters by barrier beaches ( barrier islands and barrier spits ). Formation of barrier beaches partially encloses

4606-734: The Severn Estuary in the United Kingdom and the Ems Dollard along the Dutch-German border. The width-to-depth ratio of these estuaries is typically large, appearing wedge-shaped (in cross-section) in the inner part and broadening and deepening seaward. Water depths rarely exceed 30 m (100 ft). Examples of this type of estuary in the U.S. are the Hudson River , Chesapeake Bay , and Delaware Bay along

4704-478: The black-tailed godwit , rely on estuaries. Two of the main challenges of estuarine life are the variability in salinity and sedimentation . Many species of fish and invertebrates have various methods to control or conform to the shifts in salt concentrations and are termed osmoconformers and osmoregulators . Many animals also burrow to avoid predation and to live in a more stable sedimental environment. However, large numbers of bacteria are found within

4802-555: The resource inputs are generally controlled by external processes like climate and parent material, the availability of these resources within the ecosystem is controlled by internal factors like decomposition, root competition or shading. Other factors like disturbance, succession or the types of species present are also internal factors. Primary production is the production of organic matter from inorganic carbon sources. This mainly occurs through photosynthesis . The energy incorporated through this process supports life on earth, while

4900-435: The "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . They also include less tangible items like tourism and recreation, and genes from wild plants and animals that can be used to improve domestic species. Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like

4998-400: The United States. Estuaries are incredibly dynamic systems, where temperature, salinity, turbidity, depth and flow all change daily in response to the tides. This dynamism makes estuaries highly productive habitats, but also make it difficult for many species to survive year-round. As a result, estuaries large and small experience strong seasonal variation in their fish communities. In winter,

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5096-513: The above definition of an estuary and could be fully saline. Many estuaries suffer degeneration from a variety of factors including soil erosion , deforestation , overgrazing , overfishing and the filling of wetlands. Eutrophication may lead to excessive nutrients from sewage and animal wastes; pollutants including heavy metals , polychlorinated biphenyls , radionuclides and hydrocarbons from sewage inputs; and diking or damming for flood control or water diversion. The word "estuary"

5194-481: The absence of disturbance, net ecosystem production is equivalent to the net carbon accumulation in the ecosystem. Energy can also be released from an ecosystem through disturbances such as wildfire or transferred to other ecosystems (e.g., from a forest to a stream to a lake) by erosion . In aquatic systems , the proportion of plant biomass that gets consumed by herbivores is much higher than in terrestrial systems. In trophic systems, photosynthetic organisms are

5292-402: The amount of energy available to the ecosystem. Parent material determines the nature of the soil in an ecosystem, and influences the supply of mineral nutrients. Topography also controls ecosystem processes by affecting things like microclimate , soil development and the movement of water through a system. For example, ecosystems can be quite different if situated in a small depression on

5390-504: The amount of light available, the amount of leaf area a plant has to capture light (shading by other plants is a major limitation of photosynthesis), the rate at which carbon dioxide can be supplied to the chloroplasts to support photosynthesis, the availability of water, and the availability of suitable temperatures for carrying out photosynthesis. Energy and carbon enter ecosystems through photosynthesis, are incorporated into living tissue, transferred to other organisms that feed on

5488-452: The bottom where they are harmless. Historically the oysters filtered the estuary's entire water volume of excess nutrients every three or four days. Today that process takes almost a year, and sediment, nutrients, and algae can cause problems in local waters. Some major rivers that run through deserts historically had vast, expansive estuaries that have been reduced to a fraction of their former size, because of dams and diversions. One example

5586-408: The carbon makes up much of the organic matter in living and dead biomass, soil carbon and fossil fuels . It also drives the carbon cycle , which influences global climate via the greenhouse effect . Through the process of photosynthesis, plants capture energy from light and use it to combine carbon dioxide and water to produce carbohydrates and oxygen . The photosynthesis carried out by all

5684-432: The combustion of fossil fuels, ammonia gas which evaporates from agricultural fields which have had fertilizers applied to them, and dust. Anthropogenic nitrogen inputs account for about 80% of all nitrogen fluxes in ecosystems. When plant tissues are shed or are eaten, the nitrogen in those tissues becomes available to animals and microbes. Microbial decomposition releases nitrogen compounds from dead organic matter in

5782-418: The concept to draw attention to the importance of transfers of materials between organisms and their environment. He later refined the term, describing it as "The whole system, ... including not only the organism-complex, but also the whole complex of physical factors forming what we call the environment". Tansley regarded ecosystems not simply as natural units, but as "mental isolates". Tansley later defined

5880-403: The dead organic matter would accumulate in an ecosystem, and nutrients and atmospheric carbon dioxide would be depleted. Decomposition processes can be separated into three categories— leaching , fragmentation and chemical alteration of dead material. As water moves through dead organic matter, it dissolves and carries with it the water-soluble components. These are then taken up by organisms in

5978-409: The dissolved oxygen from the water; thus hypoxic or anoxic zones can develop. Nitrogen is often the lead cause of eutrophication in estuaries in temperate zones. During a eutrophication event, biogeochemical feedback decreases the amount of available silica . These feedbacks also increase the supply of nitrogen and phosphorus, creating conditions where harmful algal blooms can persist. Given

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6076-688: The east, Cape Ann, side of the north entrance to the river. There are only three fixed crossings of the river: Massachusetts Route 128 , the Rockport Branch of the MBTA Newburyport/Rockport Line, and Western Avenue ( Massachusetts Route 127 ). There are strong tidal currents in the river, and unusually the current flows in opposite directions at the two ends. 42°39′18″N 70°41′8″W  /  42.65500°N 70.68556°W  / 42.65500; -70.68556 Estuary Most existing estuaries formed during

6174-655: The ecosystem or to gradual disruption of biotic processes and degradation of abiotic conditions of the ecosystem. Once the original ecosystem has lost its defining features, it is considered "collapsed ". Ecosystem restoration can contribute to achieving the Sustainable Development Goals . An ecosystem (or ecological system) consists of all the organisms and the abiotic pools (or physical environment) with which they interact. The biotic and abiotic components are linked together through nutrient cycles and energy flows. "Ecosystem processes" are

6272-401: The effects of modifying the estuarine circulation. Fjord -type estuaries are formed in deeply eroded valleys formed by glaciers . These U-shaped estuaries typically have steep sides, rock bottoms, and underwater sills contoured by glacial movement. The estuary is shallowest at its mouth, where terminal glacial moraines or rock bars form sills that restrict water flow. In the upper reaches of

6370-601: The estuary impacted by human activities, and over time may shift the basic composition of the ecosystem, and the reversible or irreversible changes in the abiotic and biotic parts of the systems from the bottom up. For example, Chinese and Russian industrial pollution, such as phenols and heavy metals, has devastated fish stocks in the Amur River and damaged its estuary soil. Estuaries tend to be naturally eutrophic because land runoff discharges nutrients into estuaries. With human activities, land run-off also now includes

6468-424: The estuary, the depth can exceed 300 m (1,000 ft). The width-to-depth ratio is generally small. In estuaries with very shallow sills, tidal oscillations only affect the water down to the depth of the sill, and the waters deeper than that may remain stagnant for a very long time, so there is only an occasional exchange of the deep water of the estuary with the ocean. If the sill depth is deep, water circulation

6566-776: The estuary, with only narrow inlets allowing contact with the ocean waters. Bar-built estuaries typically develop on gently sloping plains located along tectonically stable edges of continents and marginal sea coasts. They are extensive along the Atlantic and Gulf coasts of the U.S. in areas with active coastal deposition of sediments and where tidal ranges are less than 4 m (13 ft). The barrier beaches that enclose bar-built estuaries have been developed in several ways: Fjords were formed where Pleistocene glaciers deepened and widened existing river valleys so that they become U-shaped in cross-sections. At their mouths there are typically rocks, bars or sills of glacial deposits , which have

6664-609: The fish community is dominated by hardy marine residents, and in summer a variety of marine and anadromous fishes move into and out of estuaries, capitalizing on their high productivity. Estuaries provide a critical habitat to a variety of species that rely on estuaries for life-cycle completion. Pacific Herring ( Clupea pallasii ) are known to lay their eggs in estuaries and bays, surfperch give birth in estuaries, juvenile flatfish and rockfish migrate to estuaries to rear, and anadromous salmonids and lampreys use estuaries as migration corridors. Also, migratory bird populations, such as

6762-421: The flow of energy through a lake was the primary driver of the ecosystem. Hutchinson's students, brothers Howard T. Odum and Eugene P. Odum , further developed a "systems approach" to the study of ecosystems. This allowed them to study the flow of energy and material through ecological systems. Ecosystems are controlled by both external and internal factors. External factors, also called state factors, control

6860-458: The forests of eastern North America still show legacies of cultivation which ceased in 1850 when large areas were reverted to forests. Another example is the methane production in eastern Siberian lakes that is controlled by organic matter which accumulated during the Pleistocene . Ecosystems continually exchange energy and carbon with the wider environment . Mineral nutrients, on

6958-400: The great biodiversity of this ecosystem. During an algal bloom , fishermen have noticed a significant increase in the quantity of fish. A sudden increase in primary productivity causes spikes in fish populations which leads to more oxygen being utilized. It is the continued deoxygenation of the water that then causes a decline in fish populations. These effects can begin in estuaries and have

7056-420: The impacts do not end there. Plant death alters the entire food web structure which can result in the death of animals within the afflicted biome . Estuaries are hotspots for biodiversity , containing a majority of commercial fish catch, making the impacts of eutrophication that much greater within estuaries. Some specific estuarine animals feel the effects of eutrophication more strongly than others. One example

7154-691: The landscape, versus one present on an adjacent steep hillside. Other external factors that play an important role in ecosystem functioning include time and potential biota , the organisms that are present in a region and could potentially occupy a particular site. Ecosystems in similar environments that are located in different parts of the world can end up doing things very differently simply because they have different pools of species present. The introduction of non-native species can cause substantial shifts in ecosystem function. Unlike external factors, internal factors in ecosystems not only control ecosystem processes but are also controlled by them. While

7252-423: The living and dead plant matter, and eventually released through respiration. The carbon and energy incorporated into plant tissues (net primary production) is either consumed by animals while the plant is alive, or it remains uneaten when the plant tissue dies and becomes detritus . In terrestrial ecosystems , the vast majority of the net primary production ends up being broken down by decomposers . The remainder

7350-471: The maintenance of hydrological cycles , cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. Many ecosystems become degraded through human impacts, such as soil loss , air and water pollution , habitat fragmentation , water diversion , fire suppression , and introduced species and invasive species . These threats can lead to abrupt transformation of

7448-408: The maintenance of hydrological cycles, cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. While material from the ecosystem had traditionally been recognized as being the basis for things of economic value, ecosystem services tend to be taken for granted. The Millennium Ecosystem Assessment

7546-525: The many chemicals used as fertilizers in agriculture as well as waste from livestock and humans. Excess oxygen-depleting chemicals in the water can lead to hypoxia and the creation of dead zones . This can result in reductions in water quality, fish, and other animal populations. Overfishing also occurs. Chesapeake Bay once had a flourishing oyster population that has been almost wiped out by overfishing. Oysters filter these pollutants, and either eat them or shape them into small packets that are deposited on

7644-422: The marine environment, such as plastics , pesticides , furans , dioxins , phenols and heavy metals . Such toxins can accumulate in the tissues of many species of aquatic life in a process called bioaccumulation . They also accumulate in benthic environments, such as estuaries and bay muds : a geological record of human activities of the last century. The elemental composition of biofilm reflect areas of

7742-575: The marsh causing increased rates of erosion . A similar phenomenon occurs in mangrove swamps , which are another potential ecosystem in estuaries. An increase in nitrogen causes an increase in shoot growth and a decrease in root growth. Weaker root systems cause a mangrove tree to be less resilient in seasons of drought, which can lead to the death of the mangrove. This shift in above ground and below ground biomass caused by eutrophication could hindered plant success in these ecosystems. Across all biomes, eutrophication often results in plant death but

7840-440: The now off-balance nitrogen cycle , estuaries can be driven to phosphorus limitation instead of nitrogen limitation. Estuaries can be severely impacted by an unbalanced phosphorus cycle, as phosphorus interacts with nitrogen and silica availability. With an abundance of nutrients in the ecosystem, plants and algae overgrow and eventually decompose, which produce a significant amount of carbon dioxide. While releasing CO 2 into

7938-537: The other hand, are mostly cycled back and forth between plants, animals, microbes and the soil. Most nitrogen enters ecosystems through biological nitrogen fixation , is deposited through precipitation, dust, gases or is applied as fertilizer . Most terrestrial ecosystems are nitrogen-limited in the short term making nitrogen cycling an important control on ecosystem production. Over the long term, phosphorus availability can also be critical. Macronutrients which are required by all plants in large quantities include

8036-509: The other. Despite this, the cumulative effect of additional species in an ecosystem is not linear: additional species may enhance nitrogen retention, for example. However, beyond some level of species richness, additional species may have little additive effect unless they differ substantially from species already present. This is the case for example for exotic species . The addition (or loss) of species that are ecologically similar to those already present in an ecosystem tends to only have

8134-403: The overall structure of an ecosystem and the way things work within it, but are not themselves influenced by the ecosystem. On broad geographic scales, climate is the factor that "most strongly determines ecosystem processes and structure". Climate determines the biome in which the ecosystem is embedded. Rainfall patterns and seasonal temperatures influence photosynthesis and thereby determine

8232-405: The plants in an ecosystem is called the gross primary production (GPP). About half of the gross GPP is respired by plants in order to provide the energy that supports their growth and maintenance. The remainder, that portion of GPP that is not used up by respiration, is known as the net primary production (NPP). Total photosynthesis is limited by a range of environmental factors. These include

8330-585: The primary nutrients (which are most limiting as they are used in largest amounts): Nitrogen, phosphorus, potassium. Secondary major nutrients (less often limiting) include: Calcium, magnesium, sulfur. Micronutrients required by all plants in small quantities include boron, chloride, copper, iron, manganese, molybdenum, zinc. Finally, there are also beneficial nutrients which may be required by certain plants or by plants under specific environmental conditions: aluminum, cobalt, iodine, nickel, selenium, silicon, sodium, vanadium. Until modern times, nitrogen fixation

8428-413: The primary producers. The organisms that consume their tissues are called primary consumers or secondary producers — herbivores . Organisms which feed on microbes ( bacteria and fungi ) are termed microbivores . Animals that feed on primary consumers— carnivores —are secondary consumers. Each of these constitutes a trophic level. The sequence of consumption—from plant to herbivore, to carnivore—forms

8526-427: The seawater in a layer that gradually thins as it moves seaward. The denser seawater moves landward along the bottom of the estuary, forming a wedge-shaped layer that is thinner as it approaches land. As a velocity difference develops between the two layers, shear forces generate internal waves at the interface, mixing the seawater upward with the freshwater. An examples of a salt wedge estuary is Mississippi River and

8624-411: The sediment which has a very high oxygen demand. This reduces the levels of oxygen within the sediment often resulting in partially anoxic conditions, which can be further exacerbated by limited water flow. Phytoplankton are key primary producers in estuaries. They move with the water bodies and can be flushed in and out with the tides . Their productivity is largely dependent upon the turbidity of

8722-409: The soil and topography , control the overall structure of an ecosystem but are not themselves influenced by the ecosystem. Internal factors are controlled, for example, by decomposition , root competition, shading, disturbance, succession, and the types of species present. While the resource inputs are generally controlled by external processes, the availability of these resources within the ecosystem

8820-611: The soil, react with mineral soil, or are transported beyond the confines of the ecosystem (and are considered lost to it). Newly shed leaves and newly dead animals have high concentrations of water-soluble components and include sugars , amino acids and mineral nutrients. Leaching is more important in wet environments and less important in dry ones. Fragmentation processes break organic material into smaller pieces, exposing new surfaces for colonization by microbes. Freshly shed leaf litter may be inaccessible due to an outer layer of cuticle or bark , and cell contents are protected by

8918-511: The soil, where plants, fungi, and bacteria compete for it. Some soil bacteria use organic nitrogen-containing compounds as a source of carbon, and release ammonium ions into the soil. This process is known as nitrogen mineralization . Others convert ammonium to nitrite and nitrate ions, a process known as nitrification . Nitric oxide and nitrous oxide are also produced during nitrification. Under nitrogen-rich and oxygen-poor conditions, nitrates and nitrites are converted to nitrogen gas ,

9016-483: The spatial extent of ecosystems using the term " ecotope ". G. Evelyn Hutchinson , a limnologist who was a contemporary of Tansley's, combined Charles Elton 's ideas about trophic ecology with those of Russian geochemist Vladimir Vernadsky . As a result, he suggested that mineral nutrient availability in a lake limited algal production . This would, in turn, limit the abundance of animals that feed on algae. Raymond Lindeman took these ideas further to suggest that

9114-454: The system through photosynthesis , building up plant tissue. Animals play an important role in the movement of matter and energy through the system, by feeding on plants and on one another. They also influence the quantity of plant and microbial biomass present. By breaking down dead organic matter , decomposers release carbon back to the atmosphere and facilitate nutrient cycling by converting nutrients stored in dead biomass back to

9212-421: The transfers of energy and materials from one pool to another. Ecosystem processes are known to "take place at a wide range of scales". Therefore, the correct scale of study depends on the question asked. The term "ecosystem" was first used in 1935 in a publication by British ecologist Arthur Tansley . The term was coined by Arthur Roy Clapham , who came up with the word at Tansley's request. Tansley devised

9310-456: The volume of freshwater, the tidal range, and the extent of evaporation of the water in the estuary. Drowned river valleys are also known as coastal plain estuaries. In places where the sea level is rising relative to the land, sea water progressively penetrates into river valleys and the topography of the estuary remains similar to that of a river valley. This is the most common type of estuary in temperate climates. Well-studied estuaries include

9408-453: The water and atmosphere, these organisms are also intaking all or nearly all of the available oxygen creating a hypoxic environment and unbalanced oxygen cycle . The excess carbon in the form of CO 2 can lead to low pH levels and ocean acidification , which is more harmful for vulnerable coastal regions like estuaries. Eutrophication has been seen to negatively impact many plant communities in estuarine ecosystems . Salt marshes are

9506-835: The water. The main phytoplankton present are diatoms and dinoflagellates which are abundant in the sediment. A primary source of food for many organisms on estuaries, including bacteria , is detritus from the settlement of the sedimentation. Of the thirty-two largest cities in the world in the early 1990s, twenty-two were located on estuaries. As ecosystems, estuaries are under threat from human activities such as pollution and overfishing . They are also threatened by sewage, coastal settlement, land clearance and much more. Estuaries are affected by events far upstream, and concentrate materials such as pollutants and sediments. Land run-off and industrial, agricultural, and domestic waste enter rivers and are discharged into estuaries. Contaminants can be introduced which do not disintegrate rapidly in

9604-679: Was the major source of nitrogen for ecosystems. Nitrogen-fixing bacteria either live symbiotically with plants or live freely in the soil. The energetic cost is high for plants that support nitrogen-fixing symbionts—as much as 25% of gross primary production when measured in controlled conditions. Many members of the legume plant family support nitrogen-fixing symbionts. Some cyanobacteria are also capable of nitrogen fixation. These are phototrophs , which carry out photosynthesis. Like other nitrogen-fixing bacteria, they can either be free-living or have symbiotic relationships with plants. Other sources of nitrogen include acid deposition produced through

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