116-401: Acacia pycnantha , most commonly known as the golden wattle , is a tree of the family Fabaceae . It grows to a height of 8 metres (26 feet) and has phyllodes (flattened leaf stalks) instead of true leaves. The profuse fragrant, golden flowers appear in late winter and spring, followed by long seed pods. Explorer Thomas Mitchell collected the type specimen , from which George Bentham wrote
232-480: A set that includes some organisms and excludes others, based on a detailed published description (for example a species description ) and on the provision of type material, which is usually available to scientists for examination in a major museum research collection, or similar institution. According to a precise set of rules laid down in the International Code of Zoological Nomenclature (ICZN) and
348-454: A type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes the defining features of that particular taxon . In older usage (pre-1900 in botany), a type was a taxon rather than a specimen. A taxon is a scientifically named grouping of organisms with other like organisms,
464-407: A clonotype. In zoological nomenclature , the type of a species or subspecies is a specimen or series of specimens. The type of a genus or subgenus is a species. The type of a suprageneric taxon (e.g., family, etc.) is a genus. Names higher than superfamily rank do not have types. A "name-bearing type" is a specimen or image that "provides the objective standard of reference whereby the application of
580-414: A diagnosis (typically, a discussion of similarities to and differences from closely related species), and an indication of where the type specimen or specimens are deposited for examination. The geographical location where a type specimen was originally found is known as its type locality . In the case of parasites, the term type host (or symbiotype) is used to indicate the host organism from which
696-428: A fuel wood. The scented flowers have been used for perfume making, and honey production in humid areas. However, the pollen is too dry to be collected by bees in dry climates. In southern Europe, it is one of several species grown for the cut-flower trade and sold as "mimosa". Like many other species of wattle, Acacia pycnantha exudes gum when stressed. Eaten by indigenous Australians , the gum has been investigated as
812-433: A golden wattle. Similarly, the green and gold colours used by Australian international sporting teams were inspired by the colours of wattles in general, rather than the golden wattle specifically. The species was depicted on a stamp captioned "wattle" as part of a 1959–60 Australian stamp set featuring Australian native flowers. In 1970, a 5c stamp labelled "Golden Wattle" was issued to complement an earlier set depicting
928-410: A lot of small plants), dead and kept safe, "curated", in a herbarium (or the equivalent for fungi). Examples of where an illustration may serve as a type include: A type does not determine the circumscription of the taxon. For example, the common dandelion is a controversial taxon: some botanists consider it to consist of over a hundred species, and others regard it as a single species. The type of
1044-485: A new generic name; the old generic name passes into synonymy and is abandoned unless there is a pressing need to make an exception (decided case-by-case, via petition to the International Commission on Zoological Nomenclature). A type genus is a genus from which the name of a family or subfamily is formed. As with type species, the type genus is not necessarily the most representative but is usually
1160-546: A part of it that has been stolen, or improperly relocated. Type illustrations have also been used by zoologists, as in the case of the Réunion parakeet , which is known only from historical illustrations and descriptions. Recently, some species have been described where the type specimen was released alive back into the wild, such as the Bulo Burti boubou (a bushshrike ), described as Laniarius liberatus , in which
1276-403: A passing remark on Linnaeus's contributions, "Linnaeus himself, must stand as the type of his Homo sapiens. He justified his choice by noting that the specimen that Linnaeus, who wrote his own autobiography five times, had most studied was probably himself. This sufficiently and correctly designated Linnaeus to be the lectotype for Homo sapiens . It has also been suggested that Edward Cope
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#17328023610791392-602: A plant grown at Hamburg Botanic Gardens from seed said to be from the Swan River Colony (Perth). Carl Meissner described A. falcinella from material from Port Lincoln in 1855. Bentham classified both as A. pycnantha in his 1864 Flora Australiensis , though he did categorise a possible subspecies angustifolia based on material from Spencer Gulf with narrower phyllodes and fewer inflorescences. However, no subspecies are currently recognised, though an informal classification distinguishes wetland and dryland forms,
1508-487: A possible alternative to gum arabic , commonly used in the food industry. Although wattles, and in particular the golden wattle, have been the informal floral emblem of Australia for many years (for instance, it represented Australia on the Coronation gown of Queen Elizabeth II in 1953), it was not until Australia's bicentenary in 1988 that the golden wattle was formally adopted as the floral emblem of Australia . This
1624-427: A reference to the dense cluster of flowers that make up the globular inflorescences. Queensland botanist Les Pedley reclassified the species as Racosperma pycnanthum in 2003, when he proposed placing almost all Australian members of the genus into the new genus Racosperma . However, this name is treated as a synonym of its original name. Johann Georg Christian Lehmann described Acacia petiolaris in 1851 from
1740-462: A reference, by one of the stereotyped Australian characters, to "the wattle" as being "the emblem of our land", with suggested methods of display, including "stick[ing] it in a bottle or hold[ing] it in your hand" — despite the wattle prop itself being a large, forked branch with sparse patches of leaves and generic yellow flowers. Fabaceae The Fabaceae ( / f ə ˈ b eɪ s i . iː , - ˌ aɪ / ) or Leguminosae , commonly known as
1856-837: A rhizobia species may often infect more than one host species. This means that one plant species may be infected by more than one species of bacteria. For example, nodules in Acacia senegal can contain seven species of rhizobia belonging to three different genera. The most distinctive characteristics that allow rhizobia to be distinguished apart are the rapidity of their growth and the type of root nodule that they form with their host. Root nodules can be classified as being either indeterminate, cylindrical and often branched, and determinate, spherical with prominent lenticels. Indeterminate nodules are characteristic of legumes from temperate climates, while determinate nodules are commonly found in species from tropical or subtropical climates. Nodule formation
1972-405: A scientific name to a specific operational taxonomic unit . Type specimens are theoretically even allowed to be aberrant or deformed individuals or color variations, though this is rarely chosen to be the case, as it makes it hard to determine to which population the individual belonged. The usage of the term type is somewhat complicated by slightly different uses in botany and zoology . In
2088-438: A severe fire, mature specimens are able to resprout . Seeds are able to persist in the soil for more than five years, germinating after fire. Like other wattles, Acacia pycnantha fixes nitrogen from the atmosphere. It hosts bacteria known as rhizobia that form root nodules, where they make nitrogen available in organic form and thus help the plant grow in poor soils. A field study across Australia and South Africa found that
2204-682: A short hypanthium , usually cup-shaped. There are normally ten stamens and one elongated superior ovary , with a curved style . They are usually arranged in indeterminate inflorescences . Fabaceae are typically entomophilous plants (i.e. they are pollinated by insects ), and the flowers are usually showy to attract pollinators . In the Caesalpinioideae , the flowers are often zygomorphic , as in Cercis , or nearly symmetrical with five equal petals, as in Bauhinia . The upper petal
2320-453: A single type specimen, a holotype, was often not designated. Also, types were not always carefully preserved, and intervening events such as wars and fires have resulted in the destruction of the original type material. The validity of a species name often rests upon the availability of original type specimens; or, if the type cannot be found, or one has never existed, upon the clarity of the description. The ICZN has existed only since 1961 when
2436-430: A staple, essential as a source of protein. Their ability to fix atmospheric nitrogen reduces fertilizer costs for farmers and gardeners who grow legumes, and means that legumes can be used in a crop rotation to replenish soil that has been depleted of nitrogen . Legume seeds and foliage have a comparatively higher protein content than non-legume materials, due to the additional nitrogen that legumes receive through
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#17328023610792552-410: A type. Describing species and appointing type specimens is part of scientific nomenclature and alpha taxonomy . When identifying material, a scientist attempts to apply a taxon name to a specimen or group of specimens based on their understanding of the relevant taxa, based on (at least) having read the type description(s), preferably also based on an examination of all the type material of all of
2668-409: A white bloom . The mature trees do not have true leaves but have phyllodes —flat and widened leaf stems—that hang down from the branches. Shiny and dark green, these are between 9 and 15 cm (3.5 and 5.9 in) long, 1–3.5 cm (0.4–1 in) wide and falcate ( sickle -shaped) to oblanceolate in shape. New growth has a bronze colouration. Field observations at Hale Conservation Park show
2784-472: A wide range of plant hosts, and Monochaetia lutea . Golden wattle is cultivated in Australia and was introduced to the northern hemisphere in the mid-1800s. Although it has a relatively short lifespan of 15 to 30 years, it is widely grown for its bright yellow, fragrant flowers. As well as being an ornamental plant, it has been used as a windbreak or in controlling erosion. Trees are sometimes planted with
2900-736: A wide variety of growth forms , including trees, shrubs, herbaceous plants, and even vines or lianas . The herbaceous plants can be annuals, biennials , or perennials, without basal or terminal leaf aggregations. Many Legumes have tendrils. They are upright plants, epiphytes , or vines. The latter support themselves by means of shoots that twist around a support or through cauline or foliar tendrils . Plants can be heliophytes , mesophytes , or xerophytes . The leaves are usually alternate and compound. Most often they are even- or odd- pinnately compound (e.g. Caragana and Robinia respectively), often trifoliate (e.g. Trifolium , Medicago ) and rarely palmately compound (e.g. Lupinus ), in
3016-513: A wide variety of taxa representing the main lineages in the Fabaceae have been found in the fossil record dating from the middle to the late Eocene , suggesting that the majority of the modern Fabaceae groups were already present and that a broad diversification occurred during this period. Therefore, the Fabaceae started their diversification approximately 60 million years ago and the most important clades separated 50 million years ago. The age of
3132-424: Is a ball-like structure covered by 40 to 100 small flowers that have five tiny petals ( pentamerous ) and long erect stamens , which give the flower head a fluffy appearance. Developing after flowering has finished, the seed pods are flattish, straight or slightly curved, 5–14 cm (2–6 in) long and 5–8 mm wide. They are initially bright green, maturing to dark brown and have slight constrictions between
3248-407: Is a host to rust fungus species in the genus Uromycladium that affect the phyllodes and branches. These include Uromycladium simplex that forms pustules and U. tepperianum that causes large swollen brown to black galls, which eventually lead to the death of the host plant. Two fungal species have been isolated from leaf spots on A. pycnantha : Seimatosporium arbuti , which is found on
3364-443: Is a specimen later selected to serve as the single type specimen for species originally described from a set of syntypes . In zoology, a lectotype is a kind of name-bearing type . When a species was originally described on the basis of a name-bearing type consisting of multiple specimens, one of those may be designated as the lectotype. Having a single name-bearing type reduces the potential for confusion, especially considering that it
3480-504: Is a specimen of a kind of bird commonly known as the spotted harrier , which currently bears the scientific name Circus assimilis . This particular specimen is the holotype for that species; the name Circus assimilis refers, by definition, to the species of that particular specimen. That species was named and described by Jardine and Selby in 1828, and the holotype was placed in the museum collection so that other scientists might refer to it as necessary. At least for type specimens there
3596-462: Is any additional specimen from among a set of syntypes after a lectotype has been designated from among them. These are not name-bearing types. A special case in Protistans where the type consists of two or more specimens of "directly related individuals" within a preparation medium such as a blood smear. The terms parahapantotype and lectohapantotype refer to type preparations additional to
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3712-425: Is clearly designated in the original description, this specimen is known as the holotype of that species. The holotype is typically placed in a major museum, or similar well-known public collection, so that it is freely available for later examination by other biologists. When the original description designated a holotype, there may be additional specimens that the author designates as additional representatives of
3828-511: Is closely related to human evolution . The family Fabaceae includes a number of plants that are common in agriculture including Glycine max ( soybean ), Phaseolus (beans), Pisum sativum ( pea ), Cicer arietinum ( chickpeas ), Vicia faba ( broad bean ), Medicago sativa ( alfalfa ), Arachis hypogaea ( peanut ), Ceratonia siliqua (carob), Trigonella foenum-graecum ( fenugreek ), and Glycyrrhiza glabra ( liquorice ). A number of species are also weedy pests in different parts of
3944-523: Is common throughout the Fabaceae. It is found in the majority of its members that only form an association with rhizobia, which in turn form an exclusive symbiosis with the Fabaceae (with the exception of Parasponia , the only genus of the 18 Ulmaceae genera that is capable of forming nodules). Nodule formation is present in all the Fabaceae sub-families, although it is less common in the Caesalpinioideae. All types of nodule formation are present in
4060-503: Is defined by isoxazolin-5-one derivatives. These compounds occur in particular together with 3-NPA and related derivatives at the same time in the same species, as found in Astragalus canadensis and Astragalus collinus . 3-NPA and isoxazlin-5-one derivatives also occur in many species of leaf beetles (see defense in insects ). Legumes are economically and culturally important plants due to their extraordinary diversity and abundance,
4176-992: Is found in the Darling Range and western wheatbelt as well as Esperance and Kalgoorlie . Outside Australia it has become naturalised in South Africa where it is considered an invasive alien plant and is uprooted to prevent water depletion and protect local flora, Tanzania, Italy, Portugal, Sardinia, India, Indonesia and New Zealand. It is present in California as a garden escapee, but is not considered to be naturalised there. In South Africa, where it had been introduced between 1858 and 1865 for dune stabilization and tannin production, it had spread along waterways into forest, mountain and lowland fynbos , and borderline areas between fynbos and karoo . The gall-forming wasp Trichilogaster signiventris has been introduced in South Africa for biological control and has reduced
4292-463: Is no requirement for a "typical" individual to be used. Genera and families , particularly those established by early taxonomists, tend to be named after species that are more "typical" for them, but here too this is not always the case and due to changes in systematics cannot be. Hence, the term name-bearing type or onomatophore is sometimes used, to denote the fact that biological types do not define "typical" individuals or taxa , but rather fix
4408-454: Is not uncommon for a series of syntypes to contain specimens of more than one species. Formally, Carl Linnaeus is the lectotype for Homo sapiens , designated in 1959. He published the first book considered to be part of taxonomical nomenclature, the 10th edition of Systema Naturae, which included the first description of Homo sapiens and determined all valid syntypes for the species. Crucially, in 1959, Professor William Stearne wrote in
4524-411: Is one of many species that are based on illustrations by Albertus Seba (1734). An ergatotype is a specimen selected to represent a worker member in hymenopterans which have polymorphic castes. A hypotype is a specimen whose details have previously been published that is used in a supplementary figure or description of the species. The term " kleptotype " informally refers to a type specimen or
4640-593: Is the innermost one, unlike in the Faboideae . Some species, like some in the genus Senna , have asymmetric flowers, with one of the lower petals larger than the opposing one, and the style bent to one side. The calyx, corolla, or stamens can be showy in this group. In the Mimosoideae , the flowers are actinomorphic and arranged in globose inflorescences. The petals are small and the stamens, which can be more than just 10, have long, coloured filaments, which are
4756-492: Is the lectotype for Homo sapiens , based on the 1994 reporting by Louie Psihoyos of an unpublished proposal by Bob Bakker to do so. However, this designation is invalid both because Edward Cope was not one of the specimens described in Systema Naturae 10th Ed., and therefore not being a syntype is not eligible, and because Stearne's designation in 1959 has seniority and invalidates future designations. A paralectotype
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4872-545: Is the most common family found in tropical rainforests and dry forests of the Americas and Africa. Recent molecular and morphological evidence supports the fact that the Fabaceae is a single monophyletic family. This conclusion has been supported not only by the degree of interrelation shown by different groups within the family compared with that found among the Leguminosae and their closest relations, but also by all
4988-448: Is vulnerable to gall attack in cultivation. Propagation is from seed which has been pre-soaked in hot water to soften the hard seed coating. Golden wattle has been grown in temperate regions around the world for the tannin in its bark, as it provides the highest yield of all wattles . Trees can be harvested for tannin from seven to ten years of age. Commercial use of its timber is limited by the small size of trees, but it has high value as
5104-529: The International Code of Nomenclature for algae, fungi, and plants (ICN), the scientific name of every taxon is almost always based on one particular specimen , or in some cases specimens. Types are of great significance to biologists, especially to taxonomists . Types are usually physical specimens that are kept in a museum or herbarium research collection, but failing that, an image of an individual of that taxon has sometimes been designated as
5220-692: The PhyloCode , type-based definitions are replaced by phylogenetic definitions . In some older taxonomic works the word "type" has sometimes been used differently. The meaning was similar in the first Laws of Botanical Nomenclature , but has a meaning closer to the term taxon in some other works: Ce seul caractère permet de distinguer ce type de toutes les autres espèces de la section. ... Après avoir étudié ces diverses formes, j'en arrivai à les considérer comme appartenant à un seul et même type spécifique. Translation: This single character permits [one to] distinguish this type from all other species of
5336-555: The Australian Capital Territory . It is found in the understorey of open eucalypt forests on dry, shallow soils. The species has become naturalised beyond its original range in Australia. In New South Wales it is especially prevalent around Sydney and the Central Coast region . In Tasmania it has spread in the east of the state and become weedy in bushland near Hobart . In Western Australia, it
5452-528: The Orchidaceae and Asteraceae , with about 765 genera and nearly 20,000 known species. The five largest genera of the family are Astragalus (over 3,000 species), Acacia (over 1,000 species), Indigofera (around 700 species), Crotalaria (around 700 species), and Mimosa (around 400 species), which constitute about a quarter of all legume species. The c. 19,000 known legume species amount to about 7% of flowering plant species. Fabaceae
5568-526: The Paleogene to become a ubiquitous part of the modern earth's biota , along with many other families belonging to the flowering plants. The Fabaceae have an abundant and diverse fossil record, especially for the Tertiary period. Fossils of flowers, fruit, leaves, wood and pollen from this period have been found in numerous locations. The earliest fossils that can be definitively assigned to
5684-613: The Rosidae clade (as established by the gene molecular phylogeny of rbcL , a gene coding for part of the RuBisCO enzyme in the chloroplast ). This grouping indicates that the predisposition for forming nodules probably only arose once in flowering plants and that it can be considered as an ancestral characteristic that has been conserved or lost in certain lineages. However, such a wide distribution of families and genera within this lineage indicates that nodulation had multiple origins. Of
5800-580: The Tethys seaway during the Palaeogene Period. However, others contend that Africa (or even the Americas ) cannot yet be ruled out as the origin of the family. The current hypothesis about the evolution of the genes needed for nodulation is that they were recruited from other pathways after a polyploidy event. Several different pathways have been implicated as donating duplicated genes to
5916-726: The Whipstick forest near Bendigo in Victoria, putative hybrids with Whirrakee wattle ( Acacia williamsonii ) have been identified; these resemble hakea wattle ( Acacia hakeoides ). Garden hybrids with Queensland silver wattle ( Acacia podalyriifolia ) raised in Europe have been given the names Acacia x siebertiana and Acacia x deneufvillei . Golden wattle occurs in south-eastern Australia from South Australia 's southern Eyre Peninsula and Flinders Ranges across Victoria and northwards into inland areas of southern New South Wales and
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#17328023610796032-466: The flowers of one plant to others thereby ensuring pollination. Many Fabaceae species are important sources of pollen and nectar for bees, including for honey production in the beekeeping industry. Example Fabaceae such as alfalfa , and various clovers including white clover and sweet clover , are important sources of nectar and honey for the Western honey bee . Type specimen In biology ,
6148-458: The fruit of these plants, which are called legumes . Fabaceae range in habit from giant trees (like Koompassia excelsa ) to small annual herbs , with the majority being herbaceous perennials. Plants have indeterminate inflorescences, which are sometimes reduced to a single flower. The flowers have a short hypanthium and a single carpel with a short gynophore , and after fertilization produce fruits that are legumes. The Fabaceae have
6264-406: The jewel beetle species Agrilus assimilis , A. australasiae and A. hypoleucus . The larvae of a number of butterfly species feed on the foliage including the fiery jewel , icilius blue , lithocroa blue and wattle blue . Trichilogaster wasps form galls in the flowerheads, disrupting seed set and Acizzia acaciaepycnanthae , a psyllid , sucks sap from the leaves. Acacia pycnantha
6380-399: The legume , pea , or bean family , are a large and agriculturally important family of flowering plants . It includes trees , shrubs , and perennial or annual herbaceous plants , which are easily recognized by their fruit ( legume ) and their compound, stipulate leaves. The family is widely distributed, and is the third-largest land plant family in number of species, behind only
6496-566: The nomenclature code applied to the organism in question, a type can be a specimen, a culture, an illustration , or (under the bacteriological code) a description. Some codes consider a subordinate taxon to be the type, but under the botanical code, the type is always a specimen or illustration. For example, in the research collection of the Natural History Museum in London, there is a bird specimen numbered 1886.6.24.20. This
6612-697: The species description in 1842. The species is native to southeastern Australia as an understorey plant in eucalyptus forest. Plants are cross-pollinated by several species of honeyeater and thornbill , which visit nectaries on the phyllodes and brush against flowers, transferring pollen between them. A. pycnantha has become a weed in areas of Australia, as well as in Africa and Eurasia. Its bark produces more tannin than any other wattle species, resulting in its commercial cultivation for production of this compound. It has been widely grown as an ornamental garden plant and for cut flower production. A. pycnantha
6728-503: The 10 families within the Rosidae, 8 have nodules formed by actinomyces ( Betulaceae , Casuarinaceae , Coriariaceae , Datiscaceae , Elaeagnaceae , Myricaceae , Rhamnaceae and Rosaceae ), and the two remaining families, Ulmaceae and Fabaceae have nodules formed by rhizobia. The rhizobia and their hosts must be able to recognize each other for nodule formation to commence. Rhizobia are specific to particular host species although
6844-462: The Fabaceae appeared in the early Palaeocene (approximately 65 million years ago). Representatives of the 3 sub-families traditionally recognised as being members of the Fabaceae – Cesalpinioideae, Papilionoideae and Mimosoideae – as well as members of the large clades within these sub-families – such as the genistoides – have been found in periods later, starting between 55 and 50 million years ago. In fact,
6960-536: The Fabaceae. Forisome proteins are found in the sieve tubes of Fabaceae; uniquely they are not dependent on ADT . The order Fabales contains around 7.3% of eudicot species and the greatest part of this diversity is contained in just one of the four families that the order contains: Fabaceae. This clade also includes the families Polygalaceae , Surianaceae and Quillajaceae and its origins date back 94 to 89 million years, although it started its diversification 79 to 74 million years ago. The Fabaceae diversified during
7076-725: The Faboideae, even though diversification within each genus was relatively recent. For instance, Astragalus separated from the Oxytropis 16 to 12 million years ago. In addition, the separation of the aneuploid species of Neoastragalus started 4 million years ago. Inga , another genus of the Papilionoideae with approximately 350 species, seems to have diverged in the last 2 million years. It has been suggested, based on fossil and phylogenetic evidence, that legumes originally evolved in arid and/or semi-arid regions along
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#17328023610797192-577: The Mimosoideae and the Caesalpinioideae commonly bipinnate (e.g. Acacia , Mimosa ). They always have stipules , which can be leaf-like (e.g. Pisum ), thorn-like (e.g. Robinia ) or be rather inconspicuous. Leaf margins are entire or, occasionally, serrate . Both the leaves and the leaflets often have wrinkled pulvini to permit nastic movements . In some species, leaflets have evolved into tendrils (e.g. Vicia ). Many species have leaves with structures that attract ants which protect
7308-583: The ability to take nitrogen gas (N 2 ) out of the air and convert it to a form of nitrogen that is usable to the host plant ( NO 3 or NH 3 ). This process is called nitrogen fixation . The legume, acting as a host, and rhizobia , acting as a provider of usable nitrate, form a symbiotic relationship. Members of the Phaseoleae genus Apios form tubers, which can be edible. The flowers often have five generally fused sepals and five free petals . They are generally hermaphroditic and have
7424-500: The apex (remaining free at the base), forming a boat-like structure called the keel. The stamens are always ten in number, and their filaments can be fused in various configurations, often in a group of nine stamens plus one separate stamen. Various genes in the CYCLOIDEA (CYC)/DICHOTOMA (DICH) family are expressed in the upper (also called dorsal or adaxial) petal; in some species, such as Cadia , these genes are expressed throughout
7540-429: The basic legume fruit. The Fabaceae are rarely cyanogenic . Where they are, the cyanogenic compounds are derived from tyrosine , phenylalanine or leucine . They frequently contain alkaloids . Proanthocyanidins can be present either as cyanidin or delphinidine or both at the same time. Flavonoids such as kaempferol , quercitin and myricetin are often present. Ellagic acid has never been found in any of
7656-591: The bulk of new growth occurs over spring and summer from October to January. Floral buds are produced year-round on the tips of new growth, but only those initiated between November and May go on to flower several months later. Flowering usually takes place from July to November (late winter to early summer) in the golden wattle's native range, and because the later buds develop faster, flowering peaks over July and August. The bright yellow inflorescences occur in groups of 40 to 80 on 2.5–9 cm (1–4 in)-long racemes that arise from axillary buds . Each inflorescence
7772-527: The capacity of trees to reproduce throughout their range. The eggs are laid by adult wasps into buds of flower heads in the summer, before hatching in May and June when the larvae induce the formation of the grape-like galls and prevent flower development. The galls can be so heavy that branches break under their weight. In addition, the introduction in 2001 of the acacia seed weevil Melanterius compactus has also proved effective. Though plants are usually killed by
7888-436: The chloroplast genes rbcL and matK , or the ribosomal spacers ITS ) and cladistic analysis in order to investigate the relationships between the family's different lineages. Fabaceae is consistently recovered as monophyletic . The studies further confirmed that the traditional subfamilies Mimosoideae and Papilionoideae were each monophyletic but both were nested within the paraphyletic subfamily Caesalpinioideae. All
8004-560: The compound 3-nitropropanoic acid (3-NPA, beta-nitropropionic acid ). The free acid 3-NPA is an irreversible inhibitor of mitochondrial respiration , and thus the compound inhibits the tricarboxylic acid cycle . This inhibition caused by 3-NPA is especially toxic to nerve cells and represents a very general toxic mechanism suggesting a profound ecological importance due to the big number of species producing this compound and its derivatives. A second and closely related class of secondary metabolites that occur in many species of leguminous plants
8120-548: The different approaches yielded similar results regarding the relationships between the family's main clades. Following extensive discussion in the legume phylogenetics community, the Legume Phylogeny Working Group reclassified Fabaceae into six subfamilies, which necessitated the segregation of four new subfamilies from Caesalpinioideae and merging Caesapinioideae sensu stricto with the former subfamily Mimosoideae. The exact branching order of
8236-557: The different subfamilies is still unresolved. Polygalaceae ( outgroup ) Surianaceae (outgroup) Quillajaceae (outgroup) Cercidoideae Detarioideae Duparquetioideae Dialioideae Caesalpinioideae Faboideae The Fabaceae are placed in the order Fabales according to most taxonomic systems, including the APG III system . The family now includes six subfamilies: The Fabaceae have an essentially worldwide distribution, being found everywhere except Antarctica and
8352-471: The essential characteristics of the genus to which it belongs, but this is subjective and, ultimately, technically irrelevant, as it is not a requirement of the Code. If the type species proves, upon closer examination, to belong to a pre-existing genus (a common occurrence), then all of the constituent species must be either moved into the pre-existing genus or disassociated from the original type species and given
8468-432: The first edition of the Code was published. The ICZN does not always demand a type specimen for the historical validity of a species, and many "type-less" species do exist. The current edition of the Code, Article 75.3, prohibits the designation of a neotype unless there is "an exceptional need" for "clarifying the taxonomic status" of a species (Article 75.2). There are many other permutations and variations on terms using
8584-506: The floral emblems of Australia. To mark Australia Day in 1990, a 41c stamp labelled "Acacia pycnantha" was issued. Another stamp labelled "Golden Wattle", with a value of 70c, was issued in 2014. Clare Waight Keller included golden wattles to represent Australia in Meghan Markle's wedding veil, which included the distinctive flora of each Commonwealth country. The 1970 Monty Python's Flying Circus Bruces sketch includes
8700-427: The flower heads and dislodge pollen and often visit multiple trees. Several species of honeyeater , including the white-naped , yellow-faced , New Holland , and occasionally white-plumed and crescent honeyeaters , and eastern spinebills have been observed foraging. Other bird species include the silvereye , striated , buff-rumped and brown thornbills . As well as eating nectar, birds often pick off insects on
8816-432: The flower, producing a radially symmetrical flower. The ovary most typically develops into a legume . A legume is a simple dry fruit that usually dehisces (opens along a seam) on two sides. A common name for this type of fruit is a "pod", although that can also be applied to a few other fruit types. A few species have evolved samarae , loments , follicles , indehiscent legumes, achenes , drupes , and berries from
8932-435: The foliage. Honeybees, native bees, ants and flies also visit nectaries, but generally do not come into contact with the flowers during this activity. The presence of A. pycnantha is positively correlated with numbers of swift parrots overwintering in box–ironbark forest in central Victoria, though it is not clear whether the parrots are feeding on them or some other factor is at play. The wood serves as food for larvae of
9048-429: The genera or species analysed. Sugars are transported within the plants in the form of sucrose . C3 photosynthesis has been found in a wide variety of genera. The family has also evolved a unique chemistry. Many legumes contain toxic and indigestible substances, antinutrients , which may be removed through various processing methods. Pterocarpans are a class of molecules (derivatives of isoflavonoids ) found only in
9164-474: The hapantotype and designated by the describing author. As with other type designations the use of the prefix "Neo-", such as Neohapantotype , is employed when a replacement for the original hapantotype is designated, or when an original description did not include a designated type specimen. An illustration on which a new species or subspecies was based. For instance, the Burmese python, Python bivittatus ,
9280-457: The high Arctic. The trees are often found in tropical regions, while the herbaceous plants and shrubs are predominant outside the tropics. Biological nitrogen fixation (BNF, performed by the organisms called diazotrophs ) is a very old process that probably originated in the Archean eon when the primitive atmosphere lacked oxygen . It is only carried out by Euryarchaeota and just 6 of
9396-428: The infection thread development in that infection threads grow in a polar manner that is similar to a pollen tubes polar growth towards the ovules. Both pathways include the same type of enzymes, pectin-degrading cell wall enzymes. The enzymes needed to reduce nitrogen, nitrogenases, require a substantial input of ATP but at the same time are sensitive to free oxygen. To meet the requirements of this paradoxical situation,
9512-505: The latter with narrower phyllodes. In 1921 Joseph Maiden described Acacia westonii from the northern and western slopes of Mount Jerrabomberra near Queanbeyan in New South Wales . He felt it was similar to, but distinct from, A. pycnantha and was uncertain whether it warranted species rank. His colleague Richard Hind Cambage grew seedlings and reported they had much longer internodes than those of A. pycnantha , and that
9628-468: The main Cesalpinioideae clades have been estimated as between 56 and 34 million years and the basal group of the Mimosoideae as 44 ± 2.6 million years. The division between Mimosoideae and Faboideae is dated as occurring between 59 and 34 million years ago and the basal group of the Faboideae as 58.6 ± 0.2 million years ago. It has been possible to date the divergence of some of the groups within
9744-761: The microbes are genetically diverse, belonging to various strains of the species Bradyrhizobium japonicum and genus Burkholderia in both countries. It is unclear whether the golden wattle was accompanied by the bacteria to the African continent or encountered new populations there. Self-incompatible , A. pycnantha cannot fertilise itself and requires cross- pollination between plants to set seed. Birds facilitate this and field experiments keeping birds away from flowers greatly reduces seed production. Nectaries are located on phyllodes; those near open flowers become active, producing nectar that birds feed upon just before or during flowering. While feeding, birds brush against
9860-585: The more than 50 phyla of bacteria . Some of these lineages co-evolved together with the flowering plants establishing the molecular basis of a mutually beneficial symbiotic relationship. BNF is carried out in nodules that are mainly located in the root cortex, although they are occasionally located in the stem as in Sesbania rostrata . The spermatophytes that co-evolved with actinorhizal diazotrophs ( Frankia ) or with rhizobia to establish their symbiotic relationship belong to 11 families contained within
9976-408: The name Taraxacum officinale is the same whether the circumscription of the species includes all those small species ( Taraxacum officinale is a "big" species) or whether the circumscription is limited to only one small species among the other hundred ( Taraxacum officinale is a "small" species). The name Taraxacum officinale is the same and the type of the name is the same, but the extent to which
10092-525: The name actually applies varies greatly. Setting the circumscription of a taxon is done by a taxonomist in a publication. Miscellaneous notes: The ICN provides a listing of the various kinds of types (article 9 and the Glossary), the most important of which is the holotype. These are The word "type" appears in botanical literature as a part of some older terms that have no status under the ICN : for example
10208-482: The name of a nominal taxon can be determined." Although in reality biologists may examine many specimens (when available) of a new taxon before writing an official published species description, nonetheless, under the formal rules for naming species (the International Code of Zoological Nomenclature), a single type must be designated, as part of the published description. A type description must include
10324-535: The original author never cited a specimen. A syntype is any one of two or more specimens that is listed in a species description where no holotype was designated; historically, syntypes were often explicitly designated as such, and under the present ICZN this is a requirement, but modern attempts to publish species description based on syntypes are generally frowned upon by practicing taxonomists, and most are gradually being replaced by lectotypes. Those that still exist are still considered name-bearing types. A lectotype
10440-409: The pathways need for nodulation. The main donors to the pathway were the genes associated with the arbuscular mycorrhiza symbiosis genes, the pollen tube formation genes and the haemoglobin genes. One of the main genes shown to be shared between the arbuscular mycorrhiza pathway and the nodulation pathway is SYMRK and it is involved in the plant-bacterial recognition. The pollen tube growth is similar to
10556-549: The phyllodes appeared to have three nectaries rather than the single one of the latter species. It is now regarded as a synonym of A. pycnantha . Common names recorded include golden wattle, green wattle, black wattle, and broad-leaved wattle. At Ebenezer Mission in the Wergaia country of north-western Victoria the Aboriginal people referred to it as witch . Hybrids of the species are known in nature and cultivation. In
10672-431: The plant from herbivore insects (a form of mutualism ). Extrafloral nectaries are common among the Mimosoideae and the Caesalpinioideae, and are also found in some Faboideae (e.g. Vicia sativa ). In some Acacia , the modified hollow stipules are inhabited by ants and are known as domatia . Many Fabaceae host bacteria in their roots within structures called root nodules . These bacteria, known as rhizobia , have
10788-436: The plants express a type of haemoglobin called leghaemoglobin that is believed to be recruited after a duplication event. These three genetic pathways are believed to be part of a gene duplication event then recruited to work in nodulation. The phylogeny of the legumes has been the object of many studies by research groups from around the world. These studies have used morphology, DNA data (the chloroplast intron trnL ,
10904-858: The process. Legumes are commonly used as natural fertilizers. Some legume species perform hydraulic lift , which makes them ideal for intercropping . Farmed legumes can belong to numerous classes, including forage , grain , blooms, pharmaceutical/industrial, fallow/green manure and timber species, with most commercially farmed species filling two or more roles simultaneously. There are of two broad types of forage legumes. Some, like alfalfa , clover , vetch , and Arachis , are sown in pasture and grazed by livestock. Other forage legumes such as Leucaena or Albizia are woody shrub or tree species that are either broken down by livestock or regularly cut by humans to provide fodder . Grain legumes are cultivated for their seeds , and are also called pulses . The seeds are used for human and animal consumption or for
11020-656: The production of indigo , Acacia , for gum arabic , and Derris , for the insecticide action of rotenone , a compound it produces. Fallow or green manure legume species are cultivated to be tilled back into the soil to exploit the high nitrogen levels found in most legumes. Numerous legumes are farmed for this purpose, including Leucaena , Cyamopsis and Sesbania . Various legume species are farmed for timber production worldwide, including numerous Acacia species, Dalbergia species, and Castanospermum australe . Melliferous plants offer nectar to bees and other insects to encourage them to carry pollen from
11136-660: The production of oils for industrial uses. Grain legumes include both herbaceous plants like beans , lentils , lupins , peas and peanuts , and trees such as carob , mesquite and tamarind . Lathyrus tuberosus , once extensively cultivated in Europe, forms tubers used for human consumption. Bloom legume species include species such as lupin , which are farmed commercially for their blooms, and thus are popular in gardens worldwide. Laburnum , Robinia , Gleditsia (honey locust), Acacia , Mimosa , and Delonix are ornamental trees and shrubs . Industrial farmed legumes include Indigofera , cultivated for
11252-404: The recent phylogenetic studies based on DNA sequences. These studies confirm that the Fabaceae are a monophyletic group that is closely related to the families Polygalaceae , Surianaceae and Quillajaceae and that they belong to the order Fabales . Along with the cereals , some fruits and tropical roots, a number of Leguminosae have been a staple human food for millennia and their use
11368-409: The relevant taxa. If there is more than one named type that all appear to be the same taxon, then the oldest name takes precedence and is considered to be the correct name of the material in hand. If on the other hand, the taxon appears never to have been named at all, then the scientist or another qualified expert picks a type specimen and publishes a new name and an official description. Depending on
11484-508: The same species, termed paratypes. These are not name-bearing types . An allotype is a specimen of the opposite sex to the holotype, designated from among paratypes. The word was also formerly used for a specimen that shows features not seen in the holotype of a fossil. The term is not regulated by the ICZN . A neotype is a specimen later selected to serve as the single type specimen when an original holotype has been lost or destroyed or where
11600-406: The section ... After studying the diverse forms, I came to consider them as belonging to the one and the same specific type. In botanical nomenclature , a type ( typus , nomenclatural type ), "is that element to which the name of a taxon is permanently attached." (article 7.2) In botany, a type is either a specimen or an illustration. A specimen is a real plant (or one or more parts of a plant or
11716-711: The seeds, which are arranged in a line in the pod. The oblong seeds themselves are 5.5 to 6 mm long, black and shiny, with a clavate (club-shaped) aril . They are released in December and January, when the pods are fully ripe. Species similar in appearance include mountain hickory wattle ( A. obliquinervia ), coast golden wattle ( A. leiophylla ) and golden wreath wattle ( A. saligna ). Acacia obliquinervia has grey-green phyllodes, fewer flowers in its flower heads, and broader (1.25–2.5 cm (0.5–1 in)-wide) seed pods. A. leiophylla has paler phyllodes. A. saligna has longer, narrower phyllodes. Acacia pycnantha
11832-425: The showiest part of the flower. All of the flowers in an inflorescence open at once. In the Faboideae , the flowers are zygomorphic, and have a specialized structure . The upper petal, called the banner or standard, is large and envelops the rest of the petals in bud, often reflexing when the flower blooms. The two adjacent petals, the wings, surround the two bottom petals. The two bottom petals are fused together at
11948-405: The species description included DNA sequences from blood and feather samples. Assuming there is no future question as to the status of such a species, the absence of a type specimen does not invalidate the name, but it may be necessary for the future to designate a neotype for such a taxon, should any questions arise. However, in the case of the bushshrike, ornithologists have argued that the specimen
12064-577: The subfamily Papilionoideae: indeterminate (with the meristem retained), determinate (without meristem) and the type included in Aeschynomene . The latter two are thought to be the most modern and specialised type of nodule as they are only present in some lines of the subfamily Papilionoideae. Even though nodule formation is common in the two monophyletic subfamilies Papilionoideae and Mimosoideae they also contain species that do not form nodules. The presence or absence of nodule-forming species within
12180-414: The suffix "-type" (e.g., allotype , cotype, topotype , generitype , isotype , isoneotype, isolectotype, etc.) but these are not formally regulated by the Code, and a great many are obsolete and/or idiosyncratic. However, some of these categories can potentially apply to genuine type specimens, such as a neotype; e.g., isotypic/topotypic specimens are preferred to other specimens, when they are available at
12296-626: The taller sugar gum ( Eucalyptus cladocalyx ) to make a two-layered windbreak. One form widely cultivated was originally collected on Mount Arapiles in western Victoria. It is floriferous , with fragrant flowers appearing from April to July. The species has a degree of frost tolerance and is adaptable to a wide range of soil conditions, but it prefers good drainage. It tolerates heavy soils in dry climates, as well as mild soil salinity . It can suffer yellowing ( chlorosis ) in limestone-based (alkaline) soils. Highly drought-tolerant, it needs 370–550 mm (10–20 in) winter rainfall for cultivation. It
12412-654: The three sub-families indicates that nodule formation has arisen several times during the evolution of the Fabaceae and that this ability has been lost in some lineages. For example, within the genus Acacia , a member of the Mimosoideae, A. pentagona does not form nodules, while other species of the same genus readily form nodules, as is the case for Acacia senegal , which forms both rapidly and slow growing rhizobial nodules. A large number of species within many genera of leguminous plants, e.g. Astragalus , Coronilla , Hippocrepis , Indigofera , Lotus , Securigera and Scorpiurus , produce chemicals that derive from
12528-415: The time a neotype is chosen (because they are from the same time and/or place as the original type). A topotype is a specimen that was obtained from the same location that the original type specimen came from. The term fixation is used by the Code for the declaration of a name-bearing type, whether by original or subsequent designation. Each genus must have a designated type species (the term "genotype"
12644-406: The type specimen was obtained. Zoological collections are maintained by universities and museums. Ensuring that types are kept in good condition and made available for examination by taxonomists are two important functions of such collections. And, while there is only one holotype designated, there can be other "type" specimens, the following of which are formally defined: When a single specimen
12760-545: The wide variety of edible vegetables they represent and due to the variety of uses they can be put to: in horticulture and agriculture, as a food, for the compounds they contain that have medicinal uses and for the oil and fats they contain that have a variety of uses. The history of legumes is tied in closely with that of human civilization, appearing early in Asia , the Americas (the common bean , several varieties) and Europe (broad beans) by 6,000 BCE , where they became
12876-461: The world, including Cytisus scoparius (broom), Robinia pseudoacacia (black locust) , Ulex europaeus (gorse), Pueraria montana (kudzu), and a number of Lupinus species. The name 'Fabaceae' comes from the defunct genus Faba , now included in Vicia . The term "faba" comes from Latin, and appears to simply mean "bean". Leguminosae is an older name still considered valid, and refers to
12992-493: Was a rare and hitherto unknown color morph of a long-known species, using only the available blood and feather samples. While there is still some debate on the need to deposit actual killed individuals as type specimens, it can be observed that given proper vouchering and storage, tissue samples can be just as valuable should dispute about the validity of a species arise. The various types listed above are necessary because many species were described one or two centuries ago, when
13108-633: Was first formally described by botanist George Bentham in the London Journal of Botany in 1842. The type specimen was collected by the explorer Thomas Mitchell in present-day northern Victoria between Pyramid Hill and the Loddon River . Bentham thought it was related to A. leiophylla , which he described in the same paper. The specific epithet pycnantha is derived from the Greek words pyknos (dense) and anthos (flower),
13224-526: Was made the official floral emblem of Australia in 1988, and has been featured on the country's postal stamps. Acacia pycnantha generally grows as a small tree to between 3 and 8 metres (10 and 30 feet) in height, though trees of up to 12 m (40 ft) high have been reported in Morocco. The bark is generally dark brown to grey—smooth in younger plants though it can be furrowed and rough in older plants. Branchlets may be bare and smooth or covered with
13340-420: Was once used for this but has been abandoned because the word has become much better known as the term for a different concept in genetics ). The description of a genus is usually based primarily on its type species, modified and expanded by the features of other included species. The generic name is permanently associated with the name-bearing type of its type species. Ideally, a type species best exemplifies
13456-668: Was proclaimed by Governor-General Sir Ninian Stephen in the Government gazette published on 1 September. The day was marked by a ceremony at the Australian National Botanic Gardens , which included the planting of a golden wattle by Hazel Hawke , the Prime Minister's wife. In 1992, 1 September was formally declared " National Wattle Day ". The Australian Coat of Arms includes a wreath of wattle; this does not, however, accurately represent
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