48-504: Galloisiana Grylloblatta Grylloblattella Grylloblattina Namkungia Grylloblattidae , commonly known as the icebugs or ice crawlers , is a family of extremophile ( psychrophile ) and wingless insects that live in the cold on top of mountains and the edges of glaciers. It is the only member of Grylloblattodea , which is generally considered an order . Alternatively, Grylloblattodea, along with Mantophasmatodea (rock crawlers), have been ranked as suborders of
96-480: A set that includes some organisms and excludes others, based on a detailed published description (for example a species description ) and on the provision of type material, which is usually available to scientists for examination in a major museum research collection, or similar institution. According to a precise set of rules laid down in the International Code of Zoological Nomenclature (ICZN) and
144-454: A type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes the defining features of that particular taxon . In older usage (pre-1900 in botany), a type was a taxon rather than a specimen. A taxon is a scientifically named grouping of organisms with other like organisms,
192-407: A clonotype. In zoological nomenclature , the type of a species or subspecies is a specimen or series of specimens. The type of a genus or subgenus is a species. The type of a suprageneric taxon (e.g., family, etc.) is a genus. Names higher than superfamily rank do not have types. A "name-bearing type" is a specimen or image that "provides the objective standard of reference whereby the application of
240-414: A diagnosis (typically, a discussion of similarities to and differences from closely related species), and an indication of where the type specimen or specimens are deposited for examination. The geographical location where a type specimen was originally found is known as its type locality . In the case of parasites, the term type host (or symbiotype) is used to indicate the host organism from which
288-410: A lot of small plants), dead and kept safe, "curated", in a herbarium (or the equivalent for fungi). Examples of where an illustration may serve as a type include: A type does not determine the circumscription of the taxon. For example, the common dandelion is a controversial taxon: some botanists consider it to consist of over a hundred species, and others regard it as a single species. The type of
336-485: A new generic name; the old generic name passes into synonymy and is abandoned unless there is a pressing need to make an exception (decided case-by-case, via petition to the International Commission on Zoological Nomenclature). A type genus is a genus from which the name of a family or subfamily is formed. As with type species, the type genus is not necessarily the most representative but is usually
384-546: A part of it that has been stolen, or improperly relocated. Type illustrations have also been used by zoologists, as in the case of the Réunion parakeet , which is known only from historical illustrations and descriptions. Recently, some species have been described where the type specimen was released alive back into the wild, such as the Bulo Burti boubou (a bushshrike ), described as Laniarius liberatus , in which
432-403: A passing remark on Linnaeus's contributions, "Linnaeus himself, must stand as the type of his Homo sapiens. He justified his choice by noting that the specimen that Linnaeus, who wrote his own autobiography five times, had most studied was probably himself. This sufficiently and correctly designated Linnaeus to be the lectotype for Homo sapiens . It has also been suggested that Edward Cope
480-405: A scientific name to a specific operational taxonomic unit . Type specimens are theoretically even allowed to be aberrant or deformed individuals or color variations, though this is rarely chosen to be the case, as it makes it hard to determine to which population the individual belonged. The usage of the term type is somewhat complicated by slightly different uses in botany and zoology . In
528-453: A single type specimen, a holotype, was often not designated. Also, types were not always carefully preserved, and intervening events such as wars and fires have resulted in the destruction of the original type material. The validity of a species name often rests upon the availability of original type specimens; or, if the type cannot be found, or one has never existed, upon the clarity of the description. The ICZN has existed only since 1961 when
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#1732800838951576-410: A type. Describing species and appointing type specimens is part of scientific nomenclature and alpha taxonomy . When identifying material, a scientist attempts to apply a taxon name to a specimen or group of specimens based on their understanding of the relevant taxa, based on (at least) having read the type description(s), preferably also based on an examination of all the type material of all of
624-671: Is a genus of insects in the family Grylloblattidae found in East Asia. It contains 12 species. These species belong to the genus Galloisiana : In Japan, Galloisiana species prefer rocky areas and cool, moist environments, where they are frequently found near streams (often under rocks) in well developed forests. Unlike the Grylloblatta species of North America, they are not known to forage in snowfields. They are most abundant in montane habitats in central Honshu . In central Honshu, Galloisiana specimens have been collected in
672-443: Is a specimen later selected to serve as the single type specimen for species originally described from a set of syntypes . In zoology, a lectotype is a kind of name-bearing type . When a species was originally described on the basis of a name-bearing type consisting of multiple specimens, one of those may be designated as the lectotype. Having a single name-bearing type reduces the potential for confusion, especially considering that it
720-504: Is a specimen of a kind of bird commonly known as the spotted harrier , which currently bears the scientific name Circus assimilis . This particular specimen is the holotype for that species; the name Circus assimilis refers, by definition, to the species of that particular specimen. That species was named and described by Jardine and Selby in 1828, and the holotype was placed in the museum collection so that other scientists might refer to it as necessary. At least for type specimens there
768-462: Is any additional specimen from among a set of syntypes after a lectotype has been designated from among them. These are not name-bearing types. A special case in Protistans where the type consists of two or more specimens of "directly related individuals" within a preparation medium such as a blood smear. The terms parahapantotype and lectohapantotype refer to type preparations additional to
816-425: Is clearly designated in the original description, this specimen is known as the holotype of that species. The holotype is typically placed in a major museum, or similar well-known public collection, so that it is freely available for later examination by other biologists. When the original description designated a holotype, there may be additional specimens that the author designates as additional representatives of
864-463: Is no requirement for a "typical" individual to be used. Genera and families , particularly those established by early taxonomists, tend to be named after species that are more "typical" for them, but here too this is not always the case and due to changes in systematics cannot be. Hence, the term name-bearing type or onomatophore is sometimes used, to denote the fact that biological types do not define "typical" individuals or taxa , but rather fix
912-454: Is not uncommon for a series of syntypes to contain specimens of more than one species. Formally, Carl Linnaeus is the lectotype for Homo sapiens , designated in 1959. He published the first book considered to be part of taxonomical nomenclature, the 10th edition of Systema Naturae, which included the first description of Homo sapiens and determined all valid syntypes for the species. Crucially, in 1959, Professor William Stearne wrote in
960-411: Is one of many species that are based on illustrations by Albertus Seba (1734). An ergatotype is a specimen selected to represent a worker member in hymenopterans which have polymorphic castes. A hypotype is a specimen whose details have previously been published that is used in a supplementary figure or description of the species. The term " kleptotype " informally refers to a type specimen or
1008-492: Is the lectotype for Homo sapiens , based on the 1994 reporting by Louie Psihoyos of an unpublished proposal by Bob Bakker to do so. However, this designation is invalid both because Edward Cope was not one of the specimens described in Systema Naturae 10th Ed., and therefore not being a syntype is not eligible, and because Stearne's designation in 1959 has seniority and invalidates future designations. A paralectotype
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#17328008389511056-529: The International Code of Nomenclature for algae, fungi, and plants (ICN), the scientific name of every taxon is almost always based on one particular specimen , or in some cases specimens. Types are of great significance to biologists, especially to taxonomists . Types are usually physical specimens that are kept in a museum or herbarium research collection, but failing that, an image of an individual of that taxon has sometimes been designated as
1104-692: The PhyloCode , type-based definitions are replaced by phylogenetic definitions . In some older taxonomic works the word "type" has sometimes been used differently. The meaning was similar in the first Laws of Botanical Nomenclature , but has a meaning closer to the term taxon in some other works: Ce seul caractère permet de distinguer ce type de toutes les autres espèces de la section. ... Après avoir étudié ces diverses formes, j'en arrivai à les considérer comme appartenant à un seul et même type spécifique. Translation: This single character permits [one to] distinguish this type from all other species of
1152-531: The Himalayas . They are omnivorous, but feed primarily on dead arthropods and carrion. When arthropod carcasses are scarce, they subsist on plant material. Grylloblattidae is generally thought to have emerged from within the " Grylloblattida ", a poorly defined group of extinct winged insects that first appeared in the Late Carboniferous , over 300 million years ago. The winged Aristovia from
1200-566: The nomenclature code applied to the organism in question, a type can be a specimen, a culture, an illustration , or (under the bacteriological code) a description. Some codes consider a subordinate taxon to be the type, but under the botanical code, the type is always a specimen or illustration. For example, in the research collection of the Natural History Museum in London, there is a bird specimen numbered 1886.6.24.20. This
1248-415: The edges of ice sheets. Their optimal living temperature is between 1–4 °C (34–39 °F). They can be killed at colder temperatures due to ice formation in the body, so when the temperature drops below their optimal range they survive by living under snow pack near the soil. They have a very narrow range of temperatures that they prefer and cannot withstand high temperatures; many species are killed when
1296-471: The essential characteristics of the genus to which it belongs, but this is subjective and, ultimately, technically irrelevant, as it is not a requirement of the Code. If the type species proves, upon closer examination, to belong to a pre-existing genus (a common occurrence), then all of the constituent species must be either moved into the pre-existing genus or disassociated from the original type species and given
1344-432: The first edition of the Code was published. The ICZN does not always demand a type specimen for the historical validity of a species, and many "type-less" species do exist. The current edition of the Code, Article 75.3, prohibits the designation of a neotype unless there is "an exceptional need" for "clarifying the taxonomic status" of a species (Article 75.2). There are many other permutations and variations on terms using
1392-420: The first species named was Grylloblatta campodeiformis , which means " cricket - cockroach shaped like a Campodea " (a kind of two-pronged bristletail). Most are nocturnal and appear to feed on detritus . They have long antennae (23–45 segments) and long cerci (5–8 segments), but no wings. Their eyes are either missing or reduced and they have no ocelli (simple eyes). Their closest living relatives are
1440-529: The following mountains and mountain ranges. Specimens have also been collected in Shikoku , which are genetically close to specimens from the Suzuka Mountains . Galloisiana species are generalist scavengers that feed on both plant and animal materials. This Notoptera related article is a stub . You can help Misplaced Pages by expanding it . Type locality (biology) In biology ,
1488-474: The hapantotype and designated by the describing author. As with other type designations the use of the prefix "Neo-", such as Neohapantotype , is employed when a replacement for the original hapantotype is designated, or when an original description did not include a designated type specimen. An illustration on which a new species or subspecies was based. For instance, the Burmese python, Python bivittatus ,
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1536-440: The mid-Cretaceous Burmese amber of Myanmar, around 100 million years ago is thought to be closely related to modern Grylloblattidae due to its very similar mouthparts. List of Grylloblattodea genera and species along with their type localities : In total, there are 33 extant species and 5 extant genera described as of 2015. Galloisiana 12 species Galloisia Caudell & King 1924 (preoccupied) Galloisiana
1584-408: The name Taraxacum officinale is the same whether the circumscription of the species includes all those small species ( Taraxacum officinale is a "big" species) or whether the circumscription is limited to only one small species among the other hundred ( Taraxacum officinale is a "small" species). The name Taraxacum officinale is the same and the type of the name is the same, but the extent to which
1632-525: The name actually applies varies greatly. Setting the circumscription of a taxon is done by a taxonomist in a publication. Miscellaneous notes: The ICN provides a listing of the various kinds of types (article 9 and the Glossary), the most important of which is the holotype. These are The word "type" appears in botanical literature as a part of some older terms that have no status under the ICN : for example
1680-482: The name of a nominal taxon can be determined." Although in reality biologists may examine many specimens (when available) of a new taxon before writing an official published species description, nonetheless, under the formal rules for naming species (the International Code of Zoological Nomenclature), a single type must be designated, as part of the published description. A type description must include
1728-462: The order Notoptera . Grylloblattids are wingless insects mostly less than 3 cm long, with a head resembling that of a cockroach , with long antennae and having elongated cerci arising from the tip of their abdomen. They cannot tolerate warmth (most species will die at 10 °C) and many species have small distribution ranges. Grylloblattids, ice crawlers or icebugs puzzled the scientists who discovered them in 1914, E.M. Walker and T.B. Kurata;
1776-535: The original author never cited a specimen. A syntype is any one of two or more specimens that is listed in a species description where no holotype was designated; historically, syntypes were often explicitly designated as such, and under the present ICZN this is a requirement, but modern attempts to publish species description based on syntypes are generally frowned upon by practicing taxonomists, and most are gradually being replaced by lectotypes. Those that still exist are still considered name-bearing types. A lectotype
1824-758: The recently discovered Mantophasmatodea . Most species are less than 3 cm long, the largest being Namkungia magnus . The family has its own order , Grylloblattodea (sometimes considered a suborder of Notoptera). It contains 5 genera and about 34 extant species. Most species have restricted distributions and small populations and with increased warming their habitats are threatened, making them endangered. In North America some species like Grylloblatta barberi and G. oregonensis are known from single sites. Grylloblattodea are nocturnal extremophiles typically found in leaf litter and under stones in extremely cold environments, usually at higher elevations. They are known to inhabit cold temperate forests to glaciers and
1872-409: The relevant taxa. If there is more than one named type that all appear to be the same taxon, then the oldest name takes precedence and is considered to be the correct name of the material in hand. If on the other hand, the taxon appears never to have been named at all, then the scientist or another qualified expert picks a type specimen and publishes a new name and an official description. Depending on
1920-508: The same species, termed paratypes. These are not name-bearing types . An allotype is a specimen of the opposite sex to the holotype, designated from among paratypes. The word was also formerly used for a specimen that shows features not seen in the holotype of a fossil. The term is not regulated by the ICZN . A neotype is a specimen later selected to serve as the single type specimen when an original holotype has been lost or destroyed or where
1968-406: The section ... After studying the diverse forms, I came to consider them as belonging to the one and the same specific type. In botanical nomenclature , a type ( typus , nomenclatural type ), "is that element to which the name of a taxon is permanently attached." (article 7.2) In botany, a type is either a specimen or an illustration. A specimen is a real plant (or one or more parts of a plant or
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2016-405: The species description included DNA sequences from blood and feather samples. Assuming there is no future question as to the status of such a species, the absence of a type specimen does not invalidate the name, but it may be necessary for the future to designate a neotype for such a taxon, should any questions arise. However, in the case of the bushshrike, ornithologists have argued that the specimen
2064-414: The suffix "-type" (e.g., allotype , cotype, topotype , generitype , isotype , isoneotype, isolectotype, etc.) but these are not formally regulated by the Code, and a great many are obsolete and/or idiosyncratic. However, some of these categories can potentially apply to genuine type specimens, such as a neotype; e.g., isotypic/topotypic specimens are preferred to other specimens, when they are available at
2112-548: The temperature rises about 5 °C above their optimal temperature. They move in response to the seasons so as to maintain an optimal temperature in their foraging habitat. Grylloblattidae are patchily distributed in glaciers, caves, montane environments, and occasionally also lower-elevation forests in western North America, East Asia (Korea and Japan), Central Asia (Siberia, China, and Kazakhstan). They are predicted to occur in several other mountain chains in Asia, including parts of
2160-415: The time a neotype is chosen (because they are from the same time and/or place as the original type). A topotype is a specimen that was obtained from the same location that the original type specimen came from. The term fixation is used by the Code for the declaration of a name-bearing type, whether by original or subsequent designation. Each genus must have a designated type species (the term "genotype"
2208-406: The type specimen was obtained. Zoological collections are maintained by universities and museums. Ensuring that types are kept in good condition and made available for examination by taxonomists are two important functions of such collections. And, while there is only one holotype designated, there can be other "type" specimens, the following of which are formally defined: When a single specimen
2256-493: Was a rare and hitherto unknown color morph of a long-known species, using only the available blood and feather samples. While there is still some debate on the need to deposit actual killed individuals as type specimens, it can be observed that given proper vouchering and storage, tissue samples can be just as valuable should dispute about the validity of a species arise. The various types listed above are necessary because many species were described one or two centuries ago, when
2304-420: Was once used for this but has been abandoned because the word has become much better known as the term for a different concept in genetics ). The description of a genus is usually based primarily on its type species, modified and expanded by the features of other included species. The generic name is permanently associated with the name-bearing type of its type species. Ideally, a type species best exemplifies
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