The Granitz is a wooded ridge in the southeast of Germany's largest island, Rügen , between the Baltic Sea resorts of Binz and Sellin . The woods cover an area of 982 hectares and are designated as a nature reserve . Since 1991 they have been part of the Southeast Rügen Biosphere Reserve .
91-664: Characteristic of the Granitz are its rich stands of beech and sessile oak and its rolling landscape of push end moraines , which in some ways resemble a mountain landscape. To the north and east the Granitz is bounded by a cliffed coast on the Baltic Sea. In the south it adjoins the Mönchgut region and in the north, the narrow bar of the Schmale Heide . The 23-acre (9.3 ha) Black Lake ( Schwarzer See ) lies in
182-588: A bitter taste (though not nearly as bitter as acorns ) and a high tannin content. The most recent classification system of the genus recognizes 14 species in two distinct subgenera, subgenus Englerianae and Fagus . Beech species can be diagnosed by phenotypical and/or genotypical traits. Species of subgenus Engleriana are found only in East Asia, and are notably distinct from species of subgenus Fagus in that these beeches are low-branching trees, often made up of several major trunks with yellowish bark and
273-651: A bitter, astringent, or in some cases, mild and nut-like taste. According to the Roman statesman Pliny the Elder in his work Natural History , beechnut was eaten by the people of Chios when the town was besieged, writing of the fruit: "that of the beech is the sweetest of all; so much so, that, according to Cornelius Alexander, the people of the city of Chios, when besieged, supported themselves wholly on mast". They can also be roasted and pulverized into an adequate coffee substitute . The leaves can be steeped in liquor to give
364-429: A decorative wood. The timber can be used to build chalets, houses, and log cabins. Beech wood is used for the stocks of military rifles when traditionally preferred woods such as walnut are scarce or unavailable or as a lower-cost alternative. The edible fruit of the beech tree, known as beechnuts or mast, is found in small burrs that drop from the tree in autumn. They are small, roughly triangular, and edible, with
455-408: A decussate pattern, in which each node rotates by 1/4 (90°) as in the herb basil . The leaves of tricussate plants such as Nerium oleander form a triple helix. The leaves of some plants do not form helices. In some plants, the divergence angle changes as the plant grows. In orixate phyllotaxis, named after Orixa japonica , the divergence angle is not constant. Instead, it is periodic and follows
546-399: A letter of the alphabet) is буква ( bukva ), while Serbo-Croatian and Slovene use " bukva " to refer to the tree. The pigment bistre was made from beech wood soot . Beech litter raking as a replacement for straw in animal husbandry was an old non-timber practice in forest management that once occurred in parts of Switzerland in the 17th century. Beech has been listed as one of
637-459: A light green/yellow liqueur. In antiquity, the bark of the beech tree was used by Indo-European people for writing-related purposes, especially in a religious context. Beech wood tablets were a common writing material in Germanic societies before the development of paper . The Old English bōc has the primary sense of "beech" but also a secondary sense of "book", and it is from bōc that
728-490: A long history of cultivation, Bulgaria for example, do not exhibit this pattern, so how much human activity has influenced the spread of beech trees is as yet unclear. The primeval beech forests of the Carpathians are also an example of a singular, complete, and comprehensive forest dominated by a single tree species - the beech tree. Forest dynamics here were allowed to proceed without interruption or interference since
819-687: A naturally growing forest tree, beech marks the important border between the European deciduous forest zone and the northern pine forest zone. This border is important for wildlife and fauna. In Denmark and Scania at the southernmost peak of the Scandinavian peninsula, southwest of the natural spruce boundary, it is the most common forest tree. It grows naturally in Denmark and southern Norway and Sweden up to about 57–59°N. The most northern known naturally growing (not planted) beech trees are found in
910-515: A petiole like structure. Pseudopetioles occur in some monocotyledons including bananas , palms and bamboos . Stipules may be conspicuous (e.g. beans and roses ), soon falling or otherwise not obvious as in Moraceae or absent altogether as in the Magnoliaceae . A petiole may be absent (apetiolate), or the blade may not be laminar (flattened). The petiole mechanically links the leaf to
1001-463: A plant matures; as a case in point Eucalyptus species commonly have isobilateral, pendent leaves when mature and dominating their neighbors; however, such trees tend to have erect or horizontal dorsiventral leaves as seedlings, when their growth is limited by the available light. Other factors include the need to balance water loss at high temperature and low humidity against the need to absorb atmospheric carbon dioxide. In most plants, leaves also are
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#17327795892201092-419: A regular organization at the cellular scale. Specialized cells that differ markedly from surrounding cells, and which often synthesize specialized products such as crystals, are termed idioblasts . The epidermis is the outer layer of cells covering the leaf. It is covered with a waxy cuticle which is impermeable to liquid water and water vapor and forms the boundary separating the plant's inner cells from
1183-475: A scaffolding matrix imparting mechanical rigidity to leaves. Leaves are normally extensively vascularized and typically have networks of vascular bundles containing xylem , which supplies water for photosynthesis , and phloem , which transports the sugars produced by photosynthesis. Many leaves are covered in trichomes (small hairs) which have diverse structures and functions. The major tissue systems present are These three tissue systems typically form
1274-789: A separate family, the Nothofagaceae (which remains a member of the order Fagales ). They are found throughout the Southern Hemisphere in Australia , New Zealand , New Guinea , New Caledonia , as well as Argentina and Chile (principally Patagonia and Tierra del Fuego ). Species treated in Denk et al. (2024) and listed in Plants of the World Online (POWO) : Erroneously synonymized by some authors (e.g. POWO) with
1365-541: A severe dry season, some plants may shed their leaves until the dry season ends. In either case, the shed leaves may be expected to contribute their retained nutrients to the soil where they fall. In contrast, many other non-seasonal plants, such as palms and conifers, retain their leaves for long periods; Welwitschia retains its two main leaves throughout a lifetime that may exceed a thousand years. The leaf-like organs of bryophytes (e.g., mosses and liverworts ), known as phyllids , differ heavily morphologically from
1456-402: A single (sometimes more) primary vein in the centre of the leaf, referred to as the midrib or costa, which is continuous with the vasculature of the petiole. The secondary veins, also known as second order veins or lateral veins, branch off from the midrib and extend toward the leaf margins. These often terminate in a hydathode , a secretory organ, at the margin. In turn, smaller veins branch from
1547-578: A single leaf grows from each node, and when the stem is held straight, the leaves form a helix . The divergence angle is often represented as a fraction of a full rotation around the stem. A rotation fraction of 1/2 (a divergence angle of 180°) produces an alternate arrangement, such as in Gasteria or the fan-aloe Kumara plicatilis . Rotation fractions of 1/3 (divergence angles of 120°) occur in beech and hazel . Oak and apricot rotate by 2/5, sunflowers, poplar, and pear by 3/8, and in willow and almond
1638-533: A single point. In evolutionary terms, early emerging taxa tend to have dichotomous branching with reticulate systems emerging later. Veins appeared in the Permian period (299–252 mya), prior to the appearance of angiosperms in the Triassic (252–201 mya), during which vein hierarchy appeared enabling higher function, larger leaf size and adaption to a wider variety of climatic conditions. Although it
1729-560: A small grove north of Bergen on the west coast of Norway. Near the city of Larvik is the largest naturally occurring beech forest in Norway, Bøkeskogen . Some research suggests that early agriculture patterns supported the spread of beech in continental Europe. Research has linked the establishment of beech stands in Scandinavia and Germany with cultivation and fire disturbance, i.e. early agricultural practices. Other areas which have
1820-409: A small leaf. Stipules may be lasting and not be shed (a stipulate leaf, such as in roses and beans ), or be shed as the leaf expands, leaving a stipule scar on the twig (an exstipulate leaf). The situation, arrangement, and structure of the stipules is called the "stipulation". Veins (sometimes referred to as nerves) constitute one of the most visible features of leaves. The veins in a leaf represent
1911-405: A specialized cell group known as the stomatal complex. The opening and closing of the stomatal aperture is controlled by the stomatal complex and regulates the exchange of gases and water vapor between the outside air and the interior of the leaf. Stomata therefore play the important role in allowing photosynthesis without letting the leaf dry out. In a typical leaf, the stomata are more numerous over
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#17327795892202002-578: A substantially different nucleome (nuclear DNA) , especially in noncoding, highly variable gene regions such as the spacers of the nuclear-encoded ribosomal RNA genes (ribosomal DNA) . Further differentiating characteristics include the whitish bloom on the underside of the leaves, the visible tertiary leaf veins, and a long, smooth cupule-peduncle. Originally proposed but not formalized by botanist Chung-Fu Shen in 1992, this group comprised two Japanese species, F. japonica and F. okamotoi, and one Chinese species, F. engleriana . While
2093-440: A tone between those of maple and birch , the two most popular drum woods. The textile modal is a kind of rayon often made wholly from reconstituted cellulose of pulped beech wood. The European species Fagus sylvatica yields a tough, utility timber. It weighs about 720 kg per cubic metre and is widely used for furniture construction, flooring, and engineering purposes, in plywood and household items, but rarely as
2184-483: A variety of patterns (venation) and form cylindrical bundles, usually lying in the median plane of the mesophyll , between the two layers of epidermis . This pattern is often specific to taxa, and of which angiosperms possess two main types, parallel and reticulate (net like). In general, parallel venation is typical of monocots, while reticulate is more typical of eudicots and magnoliids (" dicots "), though there are many exceptions. The vein or veins entering
2275-700: Is a genus of deciduous trees in the family Fagaceae , native to subtropical (accessory forest element) and temperate (as dominant element of mesophytic forests) Eurasia and North America . There are 14 accepted species in two distinct subgenera, Englerianae Denk & G.W.Grimm and Fagus . The subgenus Englerianae is found only in East Asia, distinctive for its low branches, often made up of several major trunks with yellowish bark. The better known species of subgenus Fagus are native to Europe , western and eastern Asia and eastern North America. They are high-branching trees with tall, stout trunks and smooth silver-grey bark. The European beech Fagus sylvatica
2366-439: Is also used to smoke Westphalian ham , andouille sausage, and some cheeses. Beeches are monoecious , bearing both male and female flowers on the same plant. The small flowers are unisexual, the female flowers borne in pairs, the male flowers wind-pollinating catkins . They are produced in spring shortly after the new leaves appear. The fruit of the beech tree, known as beechnuts or mast, is found in small burrs that drop from
2457-460: Is called a stipe in ferns . The lamina is the expanded, flat component of the leaf which contains the chloroplasts . The sheath is a structure, typically at the base that fully or partially clasps the stem above the node, where the leaf is attached. Leaf sheathes typically occur in Poaceae (grasses) and Apiaceae (umbellifers). Between the sheath and the lamina, there may be a pseudopetiole ,
2548-428: Is not planted much as an ornamental due to slower growth and less resistance to urban pollution. It most commonly occurs as an overstory component in the northern part of its range with sugar maple, transitioning to other forest types further south such as beech-magnolia. American beech is rarely encountered in developed areas except as a remnant of a forest that was cut down for land development. The dead brown leaves of
2639-476: Is smooth and light gray. The fruit is a small, sharply three-angled nut 10–15 mm ( 3 ⁄ 8 – 5 ⁄ 8 in) long, borne singly or in pairs in soft-spined husks 1.5–2.5 cm ( 5 ⁄ 8 –1 in) long, known as cupules. The husk can have a variety of spine- to scale-like appendages, the character of which is, in addition to leaf shape, one of the primary ways beeches are differentiated. The nuts are called beechnuts or beech mast and have
2730-406: Is the more complex pattern, branching veins appear to be plesiomorphic and in some form were present in ancient seed plants as long as 250 million years ago. A pseudo-reticulate venation that is actually a highly modified penniparallel one is an autapomorphy of some Melanthiaceae , which are monocots; e.g., Paris quadrifolia (True-lover's Knot). In leaves with reticulate venation, veins form
2821-426: Is the most commonly cultivated species, yielding a utility timber used for furniture construction, flooring and engineering purposes, in plywood, and household items. The timber can be used to build homes. Beechwood makes excellent firewood . Slats of washed beech wood are spread around the bottom of fermentation tanks for Budweiser beer. Beech logs are burned to dry the malt used in some German smoked beers . Beech
Granitz - Misplaced Pages Continue
2912-688: The Granitz hunting lodge constructed in the 19th century. Other landmarks include Granitz House , the grave site of Finnish warriors and the Cross Oak ( Kreuzeiche ). The name is probably of Slavic origin and comes from the personal name, Granza , used by the Rani tribe, which first appears as a prince's name in the Saxo Grammaticus in 1168. In 1888, the Waldhalle , a tourist restaurant,
3003-653: The cryptic sister species F. hayatae and F. pashanica ) , two pseudo-cryptic species in eastern North America ( F. grandifolia , F. mexicana ), and a species complex of at least four species ( F. caspica , F. hohenackeriana , F. orientalis , F. sylvatica ) in Western Eurasia . Their genetics are highly complex and include both species-unique alleles as well as alleles and ribosomal DNA spacers that are shared between two or more species. The western Eurasian species are characterized by morphological and genetical gradients. Research suggests that
3094-494: The diet of many animals . Correspondingly, leaves represent heavy investment on the part of the plants bearing them, and their retention or disposition are the subject of elaborate strategies for dealing with pest pressures, seasonal conditions, and protective measures such as the growth of thorns and the production of phytoliths , lignins , tannins and poisons . Deciduous plants in frigid or cold temperate regions typically shed their leaves in autumn, whereas in areas with
3185-424: The gymnosperms and angiosperms . Euphylls are also referred to as macrophylls or megaphylls (large leaves). A structurally complete leaf of an angiosperm consists of a petiole (leaf stalk), a lamina (leaf blade), stipules (small structures located to either side of the base of the petiole) and a sheath. Not every species produces leaves with all of these structural components. The proximal stalk or petiole
3276-551: The phyllids of mosses and liverworts . Leaves are the most important organs of most vascular plants. Green plants are autotrophic , meaning that they do not obtain food from other living things but instead create their own food by photosynthesis . They capture the energy in sunlight and use it to make simple sugars , such as glucose and sucrose , from carbon dioxide and water. The sugars are then stored as starch , further processed by chemical synthesis into more complex organic molecules such as proteins or cellulose ,
3367-399: The plant shoots and roots . Vascular plants transport sucrose in a special tissue called the phloem . The phloem and xylem are parallel to each other, but the transport of materials is usually in opposite directions. Within the leaf these vascular systems branch (ramify) to form veins which supply as much of the leaf as possible, ensuring that cells carrying out photosynthesis are close to
3458-423: The 38 plants whose flowers are used to prepare Bach flower remedies . Leaf A leaf ( pl. : leaves ) is a principal appendage of the stem of a vascular plant , usually borne laterally above ground and specialized for photosynthesis . Leaves are collectively called foliage , as in "autumn foliage", while the leaves, stem, flower, and fruit collectively form the shoot system. In most leaves,
3549-636: The American beech remain on the branches until well into the following spring, when the new buds finally push them off. East Asia is home to eight species of Fagus , only one of which ( F. crenata ) is occasionally planted in Western countries. Smaller than F. sylvatica and F. grandifolia , this beech is one of the most common hardwoods in its native range. Beech grows on a wide range of soil types, acidic or basic, provided they are not waterlogged. The tree canopy casts dense shade and thickens
3640-723: The Crimean F. × taurica, from which it differs morphologically and genetically. Numerous species have been named globally from the fossil record spanning from the Cretaceous to the Pleistocene . Fossil species formerly placed in Fagus include: The name of the tree in Latin, fagus (from whence the generic epithet ), is cognate with English "beech" and of Indo-European origin, and played an important role in early debates on
3731-607: The Granitz as do several kettle bogs . A number of non-native stands of conifers are being turned into a near-natural forest. No roads of any description run through the Granitz, but there are many cycle and footpaths. Local transport and access to the area is provided by the Rügen Light Railway and Binz Seaside Railway ( Binzer Bäderbahn ). On the highest point, the Tempelberg ( 107 m above sea level (NN) ), Prince Wilhelm Malte I of Putbus had
Granitz - Misplaced Pages Continue
3822-679: The amount of light they absorb to avoid or mitigate excessive heat, ultraviolet damage, or desiccation, or to sacrifice light-absorption efficiency in favor of protection from herbivory. For xerophytes the major constraint is not light flux or intensity , but drought. Some window plants such as Fenestraria species and some Haworthia species such as Haworthia tesselata and Haworthia truncata are examples of xerophytes. and Bulbine mesembryanthemoides . Leaves also function to store chemical energy and water (especially in succulents ) and may become specialized organs serving other functions, such as tendrils of peas and other legumes,
3913-583: The basic structural material in plant cell walls, or metabolized by cellular respiration to provide chemical energy to run cellular processes. The leaves draw water from the ground in the transpiration stream through a vascular conducting system known as xylem and obtain carbon dioxide from the atmosphere by diffusion through openings called stomata in the outer covering layer of the leaf ( epidermis ), while leaves are orientated to maximize their exposure to sunlight. Once sugar has been synthesized, it needs to be transported to areas of active growth such as
4004-458: The blade attaches directly to the stem. Subpetiolate leaves are nearly petiolate or have an extremely short petiole and may appear to be sessile. In clasping or decurrent leaves, the blade partially surrounds the stem. When the leaf base completely surrounds the stem, the leaves are said to be perfoliate , such as in Eupatorium perfoliatum . In peltate leaves, the petiole attaches to
4095-406: The blade inside the blade margin. In some Acacia species, such as the koa tree ( Acacia koa ), the petioles are expanded or broadened and function like leaf blades; these are called phyllodes . There may or may not be normal pinnate leaves at the tip of the phyllode. A stipule , present on the leaves of many dicotyledons , is an appendage on each side at the base of the petiole, resembling
4186-521: The bottom of fermentation tanks for Budweiser beer. This provides a complex surface on which the yeast can settle, so that it does not pile up, preventing yeast autolysis which would contribute off-flavours to the beer. Beech logs are burned to dry the malt used in German smoked beers . Beech is also used to smoke Westphalian ham , traditional andouille (an offal sausage) from Normandy, and some cheeses. Some drums are made from beech, which has
4277-578: The disease has become widespread in Connecticut, Massachusetts and Rhode Island, and in portions of coastal New Hampshire and coastal and central Maine. The beech most commonly grown as an ornamental tree is the European beech ( Fagus sylvatica ), widely cultivated in North America as well as its native Europe. Many varieties are in cultivation, notably the weeping beech F. sylvatica 'Pendula', several varieties of copper or purple beech,
4368-606: The early Devonian lycopsid Baragwanathia , first evolved as enations, extensions of the stem. True leaves or euphylls of larger size and with more complex venation did not become widespread in other groups until the Devonian period , by which time the carbon dioxide concentration in the atmosphere had dropped significantly. This occurred independently in several separate lineages of vascular plants, in progymnosperms like Archaeopteris , in Sphenopsida , ferns and later in
4459-821: The eastern United States and southeastern Canada, with a disjunct sister species in Mexico ( F. mexicana ). There are the only extant (surviving) Fagus species in the Western Hemisphere. Before the Pleistocene Ice Age, it is believed to have spanned the entire width of the continent from the Atlantic Ocean to the Pacific but now is confined to the east of the Great Plains. F. grandifolia tolerates hotter climates than European species but
4550-558: The equivalents of the petioles and stipules of leaves. Because each leaflet can appear to be a simple leaf, it is important to recognize where the petiole occurs to identify a compound leaf. Compound leaves are a characteristic of some families of higher plants, such as the Fabaceae . The middle vein of a compound leaf or a frond , when it is present, is called a rachis . Leaves which have a petiole (leaf stalk) are said to be petiolate . Sessile (epetiolate) leaves have no petiole and
4641-662: The exception of the Western Eurasian and North American species. Fagus is the first diverging lineage in the evolution of the Fagaceae family, which also includes oaks and chestnuts . The oldest fossils that can be assigned to the beech lineage are 81–82 million years old pollen from the Late Cretaceous of Wyoming , United States. The southern beeches (genus Nothofagus ) historically thought closely related to beeches, are treated as members of
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#17327795892204732-657: The external world. The cuticle is in some cases thinner on the lower epidermis than on the upper epidermis, and is generally thicker on leaves from dry climates as compared with those from wet climates. The epidermis serves several functions: protection against water loss by way of transpiration , regulation of gas exchange and secretion of metabolic compounds. Most leaves show dorsoventral anatomy: The upper (adaxial) and lower (abaxial) surfaces have somewhat different construction and may serve different functions. The epidermis tissue includes several differentiated cell types; epidermal cells, epidermal hair cells ( trichomes ), cells in
4823-728: The fern-leaved beech F. sylvatica 'Asplenifolia', and the tricolour beech F. sylvatica 'Roseomarginata'. The columnar Dawyck beech ( F. sylvatica 'Dawyck') occurs in green, gold, and purple forms, named after Dawyck Botanic Garden in the Scottish Borders, one of the four garden sites of the Royal Botanic Garden Edinburgh . Beech wood is an excellent firewood , easily split and burning for many hours with bright but calm flames. Slats of beech wood are washed in caustic soda to leach out any flavour or aroma characteristics and are spread around
4914-759: The first representatives of the modern-day genus were already present in the Paleocene of Arctic North America (western Greenland ) and quickly radiated across the high latitudes of the Northern Hemisphere , with a first diversity peak in the Miocene of northeastern Asia . The contemporary species are the product of past, repeated reticulate evolutionary processes ( outbreeding , introgression , hybridization ). As far as studied, heterozygosity and intragenomic variation are common in beech species, and their chloroplast genomes are nonspecific with
5005-406: The fraction is 5/13. These arrangements are periodic. The denominator of the rotation fraction indicates the number of leaves in one period, while the numerator indicates the number of complete turns or gyres made in one period. For example: Most divergence angles are related to the sequence of Fibonacci numbers F n . This sequence begins 1, 1, 2, 3, 5, 8, 13; each term is the sum of
5096-491: The geographical origins of the Indo-European people , the beech argument . Greek φηγός (figós) is from the same root, but the word was transferred to the oak tree (e.g. Iliad 16.767) as a result of the absence of beech trees in southern Greece . Fagus sylvatica was a late entrant to Great Britain after the last glaciation, and may have been restricted to basic soils in the south of England. Some suggest that it
5187-479: The ground with leaf litter . In North America, they can form beech-maple climax forests by partnering with the sugar maple . The beech blight aphid ( Grylloprociphilus imbricator ) is a common pest of American beech trees. Beeches are also used as food plants by some species of Lepidoptera . Beech bark is extremely thin and scars easily. Since the beech tree has such delicate bark, carvings, such as lovers' initials and other forms of graffiti, remain because
5278-866: The ground, they are referred to as prostrate . Perennial plants whose leaves are shed annually are said to have deciduous leaves, while leaves that remain through winter are evergreens . Leaves attached to stems by stalks (known as petioles ) are called petiolate, and if attached directly to the stem with no petiole they are called sessile. Dicot leaves have blades with pinnate venation (where major veins diverge from one large mid-vein and have smaller connecting networks between them). Less commonly, dicot leaf blades may have palmate venation (several large veins diverging from petiole to leaf edges). Finally, some exhibit parallel venation. Monocot leaves in temperate climates usually have narrow blades, and usually parallel venation converging at leaf tips or edges. Some also have pinnate venation. The arrangement of leaves on
5369-572: The last ice age. Nowadays, they are amongst the last pure beech forests in Europe to document the undisturbed postglacial repopulation of the species, which also includes the unbroken existence of typical animals and plants. These virgin beech forests and similar forests across 12 countries in continental Europe were inscribed on the UNESCO World Heritage List in 2007. The American beech ( Fagus grandifolia ) occurs across much of
5460-443: The leaf from the petiole are called primary or first-order veins. The veins branching from these are secondary or second-order veins. These primary and secondary veins are considered major veins or lower order veins, though some authors include third order. Each subsequent branching is sequentially numbered, and these are the higher order veins, each branching being associated with a narrower vein diameter. In parallel veined leaves,
5551-523: The leaf veins form, and these have functional implications. Of these, angiosperms have the greatest diversity. Within these the major veins function as the support and distribution network for leaves and are correlated with leaf shape. For instance, the parallel venation found in most monocots correlates with their elongated leaf shape and wide leaf base, while reticulate venation is seen in simple entire leaves, while digitate leaves typically have venation in which three or more primary veins diverge radially from
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#17327795892205642-551: The leaves of vascular plants . In most cases, they lack vascular tissue, are only a single cell thick, and have no cuticle , stomata, or internal system of intercellular spaces. (The phyllids of the moss family Polytrichaceae are notable exceptions.) The phyllids of bryophytes are only present on the gametophytes , while in contrast the leaves of vascular plants are only present on the sporophytes . These can further develop into either vegetative or reproductive structures. Simple, vascularized leaves ( microphylls ), such as those of
5733-429: The modern word derives. In modern German, the word for "book" is Buch, with Buche meaning "beech tree". In modern Dutch, the word for "book" is boek, with beuk meaning "beech tree". In Swedish, these words are the same, bok meaning both "beech tree" and "book". There is a similar relationship in some Slavic languages. In Russian and Bulgarian , the word for beech is бук ( buk ), while that for "letter" (as in
5824-552: The number of stomata (pores that intake and output gases), the amount and structure of epicuticular wax and other features. Leaves are mostly green in color due to the presence of a compound called chlorophyll which is essential for photosynthesis as it absorbs light energy from the sun . A leaf with lighter-colored or white patches or edges is called a variegated leaf . Leaves can have many different shapes, sizes, textures and colors. The broad, flat leaves with complex venation of flowering plants are known as megaphylls and
5915-404: The photosynthetic organelles , the chloroplasts , to light and to increase the absorption of carbon dioxide while at the same time controlling water loss. Their surfaces are waterproofed by the plant cuticle and gas exchange between the mesophyll cells and the atmosphere is controlled by minute (length and width measured in tens of μm) openings called stomata which open or close to regulate
6006-412: The plant and provides the route for transfer of water and sugars to and from the leaf. The lamina is typically the location of the majority of photosynthesis. The upper ( adaxial ) angle between a leaf and a stem is known as the axil of the leaf. It is often the location of a bud . Structures located there are called "axillary". External leaf characteristics, such as shape, margin, hairs, the petiole, and
6097-434: The presence of stipules and glands, are frequently important for identifying plants to family, genus or species levels, and botanists have developed a rich terminology for describing leaf characteristics. Leaves almost always have determinate growth. They grow to a specific pattern and shape and then stop. Other plant parts like stems or roots have non-determinate growth, and will usually continue to grow as long as they have
6188-410: The previous two. Rotation fractions are often quotients F n / F n + 2 of a Fibonacci number by the number two terms later in the sequence. This is the case for the fractions 1/2, 1/3, 2/5, 3/8, and 5/13. The ratio between successive Fibonacci numbers tends to the golden ratio φ = (1 + √5)/2 . When a circle is divided into two arcs whose lengths are in the ratio 1:φ , the angle formed by
6279-416: The primary photosynthetic tissue is the palisade mesophyll and is located on the upper side of the blade or lamina of the leaf but in some species, including the mature foliage of Eucalyptus , palisade mesophyll is present on both sides and the leaves are said to be isobilateral. Most leaves are flattened and have distinct upper ( adaxial ) and lower ( abaxial ) surfaces that differ in color, hairiness,
6370-465: The primary organs responsible for transpiration and guttation (beads of fluid forming at leaf margins). Leaves can also store food and water , and are modified accordingly to meet these functions, for example in the leaves of succulent plants and in bulb scales. The concentration of photosynthetic structures in leaves requires that they be richer in protein , minerals , and sugars than, say, woody stem tissues. Accordingly, leaves are prominent in
6461-429: The primary veins run parallel and equidistant to each other for most of the length of the leaf and then converge or fuse (anastomose) towards the apex. Usually, many smaller minor veins interconnect these primary veins, but may terminate with very fine vein endings in the mesophyll. Minor veins are more typical of angiosperms, which may have as many as four higher orders. In contrast, leaves with reticulate venation have
6552-416: The products of photosynthesis (photosynthate) from the cells where it takes place, while major veins are responsible for its transport outside of the leaf. At the same time water is being transported in the opposite direction. The number of vein endings is very variable, as is whether second order veins end at the margin, or link back to other veins. There are many elaborate variations on the patterns that
6643-429: The protective spines of cacti and the insect traps in carnivorous plants such as Nepenthes and Sarracenia . Leaves are the fundamental structural units from which cones are constructed in gymnosperms (each cone scale is a modified megaphyll leaf known as a sporophyll) and from which flowers are constructed in flowering plants . The internal organization of most kinds of leaves has evolved to maximize exposure of
6734-714: The rate exchange of carbon dioxide (CO 2 ), oxygen (O 2 ) and water vapor into and out of the internal intercellular space system. Stomatal opening is controlled by the turgor pressure in a pair of guard cells that surround the stomatal aperture. In any square centimeter of a plant leaf, there may be from 1,000 to 100,000 stomata. The shape and structure of leaves vary considerably from species to species of plant, depending largely on their adaptation to climate and available light, but also to other factors such as grazing animals (such as deer), available nutrients, and ecological competition from other plants. Considerable changes in leaf type occur within species, too, for example as
6825-535: The resources to do so. The type of leaf is usually characteristic of a species (monomorphic), although some species produce more than one type of leaf (dimorphic or polymorphic ). The longest leaves are those of the Raffia palm , R. regalis which may be up to 25 m (82 ft) long and 3 m (9.8 ft) wide. The terminology associated with the description of leaf morphology is presented, in illustrated form, at Wikibooks . Where leaves are basal, and lie on
6916-446: The secondary veins, known as tertiary or third order (or higher order) veins, forming a dense reticulate pattern. The areas or islands of mesophyll lying between the higher order veins, are called areoles . Some of the smallest veins (veinlets) may have their endings in the areoles, a process known as areolation. These minor veins act as the sites of exchange between the mesophyll and the plant's vascular system. Thus, minor veins collect
7007-455: The sequence 180°, 90°, 180°, 270°. Two basic forms of leaves can be described considering the way the blade (lamina) is divided. A simple leaf has an undivided blade. However, the leaf may be dissected to form lobes, but the gaps between lobes do not reach to the main vein. A compound leaf has a fully subdivided blade, each leaflet of the blade being separated along a main or secondary vein. The leaflets may have petiolules and stipels,
7098-454: The smaller arc is the golden angle , which is 1/φ × 360° ≈ 137.5° . Because of this, many divergence angles are approximately 137.5° . In plants where a pair of opposite leaves grows from each node, the leaves form a double helix. If the nodes do not rotate (a rotation fraction of zero and a divergence angle of 0°), the two helices become a pair of parallel lines, creating a distichous arrangement as in maple or olive trees. More common in
7189-953: The species that bear them, the majority, as broad-leaved or megaphyllous plants, which also include acrogymnosperms and ferns . In the lycopods , with different evolutionary origins, the leaves are simple (with only a single vein) and are known as microphylls . Some leaves, such as bulb scales, are not above ground. In many aquatic species, the leaves are submerged in water. Succulent plants often have thick juicy leaves, but some leaves are without major photosynthetic function and may be dead at maturity, as in some cataphylls and spines . Furthermore, several kinds of leaf-like structures found in vascular plants are not totally homologous with them. Examples include flattened plant stems called phylloclades and cladodes , and flattened leaf stems called phyllodes which differ from leaves both in their structure and origin. Some structures of non-vascular plants look and function much like leaves. Examples include
7280-793: The status of F. okamotoi remains uncertain, the most recent systematic treatment based on morphological and genetic data confirmed a third species, F. multinervis, endemic to Ulleungdo , a South Korean island in the Sea of Japan . The beeches of Ulleungdo have been traditionally treated as a subspecies of F. engleriana, to which they are phenotypically identical, or as a variety of F. japonica. The differ from their siblings by their unique nuclear and plastid genotypes. The better known subgenus Fagus beeches are high-branching with tall, stout trunks and smooth silver-gray bark. This group includes five extant species in continental and insular East Asia ( F. crenata , F. longipetiolata , F. lucida , and
7371-421: The stem is known as phyllotaxis . A large variety of phyllotactic patterns occur in nature: In the simplest mathematical models of phyllotaxis , the apex of the stem is represented as a circle. Each new node is formed at the apex, and it is rotated by a constant angle from the previous node. This angle is called the divergence angle . The number of leaves that grow from a node depends on the plant species. When
7462-547: The stomatal complex; guard cells and subsidiary cells. The epidermal cells are the most numerous, largest, and least specialized and form the majority of the epidermis. They are typically more elongated in the leaves of monocots than in those of dicots . Chloroplasts are generally absent in epidermal cells, the exception being the guard cells of the stomata . The stomatal pores perforate the epidermis and are surrounded on each side by chloroplast-containing guard cells, and two to four subsidiary cells that lack chloroplasts, forming
7553-511: The surrounding air , promoting cooling. Functionally, in addition to carrying out photosynthesis, the leaf is the principal site of transpiration , providing the energy required to draw the transpiration stream up from the roots, and guttation . Many conifers have thin needle-like or scale-like leaves that can be advantageous in cold climates with frequent snow and frost. These are interpreted as reduced from megaphyllous leaves of their Devonian ancestors. Some leaf forms are adapted to modulate
7644-632: The transportation system. Typically leaves are broad, flat and thin (dorsiventrally flattened), thereby maximising the surface area directly exposed to light and enabling the light to penetrate the tissues and reach the chloroplasts , thus promoting photosynthesis. They are arranged on the plant so as to expose their surfaces to light as efficiently as possible without shading each other, but there are many exceptions and complications. For instance, plants adapted to windy conditions may have pendent leaves, such as in many willows and eucalypts . The flat, or laminar, shape also maximizes thermal contact with
7735-453: The tree in autumn. They are small, roughly triangular, and edible, with a bitter, astringent, or mild and nut-like taste. The European beech ( Fagus sylvatica ) is the most commonly cultivated, although few important differences are seen between species aside from detail elements such as leaf shape. The leaves of beech trees are entire or sparsely toothed, from 5–15 centimetres (2–6 inches) long and 4–10 cm (2–4 in) broad. The bark
7826-622: The tree is unable to heal itself. Beech bark disease is a fungal infection that attacks the American beech through damage caused by scale insects. Infection can lead to the death of the tree. Beech leaf disease is a disease that affects American beeches spread by the newly discovered nematode, Litylenchus crenatae mccannii . This disease was first discovered in Lake County, Ohio, in 2012 and has now spread to over 41 counties in Ohio, Pennsylvania, New York, and Ontario, Canada. As of 2024,
7917-462: The vascular structure of the organ, extending into the leaf via the petiole and providing transportation of water and nutrients between leaf and stem, and play a crucial role in the maintenance of leaf water status and photosynthetic capacity. They also play a role in the mechanical support of the leaf. Within the lamina of the leaf, while some vascular plants possess only a single vein, in most this vasculature generally divides (ramifies) according to
8008-583: The world (according to Guinness World Records ) is the Meikleour Beech Hedge in Meikleour , Perth and Kinross , Scotland. Fagus sylvatica is one of the most common hardwood trees in north-central Europe, in France constituting alone about 15% of all nonconifers. The Balkans are also home to the lesser-known oriental beech ( F. orientalis ) and Crimean beech ( F. taurica ). As
8099-495: Was built on the hill of Falkenberg on the clifftop path from Binz to Sellin near the coast. The building had to be abandoned and demolished in the 1980s as a result of coast collapse. Only a few remains of foundations on the cliff face of the Baltic Sea coast recall the existence of the building. 54°23′13.03906″N 13°38′56.90″E / 54.3869552944°N 13.6491389°E / 54.3869552944; 13.6491389 Beech See text Beech ( Fagus )
8190-513: Was designated for its beech woodlands, which are believed to be on the western edge of their natural range in this steep limestone gorge. Beech is not native to Ireland; however, it was widely planted in the 18th century and can become a problem shading out the native woodland understory. Beech is widely planted for hedging and in deciduous woodlands, and mature, regenerating stands occur throughout mainland Britain at elevations below about 650 m (2,100 ft). The tallest and longest hedge in
8281-459: Was introduced by Neolithic tribes who planted the trees for their edible nuts. The beech is classified as a native in the south of England and as a non-native in the north where it is often removed from 'native' woods. Large areas of the Chilterns are covered with beech woods, which are habitat to the common bluebell and other flora. The Cwm Clydach National Nature Reserve in southeast Wales
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