A compound eye is a visual organ found in arthropods such as insects and crustaceans . It may consist of thousands of ommatidia , which are tiny independent photoreception units that consist of a cornea , lens , and photoreceptor cells which distinguish brightness and color. The image perceived by this arthropod eye is a combination of inputs from the numerous ommatidia, which are oriented to point in slightly different directions. Compared with single-aperture eyes , compound eyes have poor image resolution ; however, they possess a very large view angle and the ability to detect fast movement and, in some cases, the polarization of light. Because a compound eye is made up of a collection of ommatidia, each with its own lens, light will enter each ommatidium instead of using a single entrance point. The individual light receptors behind each lens are then turned on and off due to a series of changes in the light intensity during movement or when an object is moving, creating a flicker-effect known as the flicker frequency, which is the rate at which the ommatidia are turned on and off– this facilitates faster reaction to movement; honey bees respond in 0.01s compared with 0.05s for humans.
29-494: Trialeurodes vaporariorum , commonly known as the glasshouse whitefly or greenhouse whitefly , is an insect that inhabits the world's temperate regions. Like various other whiteflies , it is a primary insect pest of many fruit, vegetable and ornamental crops . It is frequently found in glasshouses ( greenhouses ), polytunnels , and other protected horticultural environments. Adults are 1–2 mm in length, with yellowish bodies and four wax-coated wings held near parallel to
58-563: A hundred species, are marine. Insects such as snow scorpionflies flourish in cold habitats including the Arctic and at high altitude. Insects such as desert locusts , ants, beetles, and termites are adapted to some of the hottest and driest environments on earth, such as the Sonoran Desert . Insects form a clade , a natural group with a common ancestor, among the arthropods . A phylogenetic analysis by Kjer et al. (2016) places
87-421: A pair of antennae . Insects are the most diverse group of animals, with more than a million described species ; they represent more than half of all animal species. The insect nervous system consists of a brain and a ventral nerve cord . Most insects reproduce by laying eggs . Insects breathe air through a system of paired openings along their sides, connected to small tubes that take air directly to
116-489: A single erect image. Apposition eyes can be divided into two groups. The typical apposition eye has a lens focusing light from one direction on the rhabdom , while light from other directions is absorbed by the dark wall of the ommatidium . The mantis shrimp is the most advanced example of an animal with this type of eye. In the other kind of apposition eye, found in the Strepsiptera , each lens forms an image, and
145-454: A transparent gap but use corner mirrors instead of lenses. Good fliers like flies or honey bees, or prey-catching insects like praying mantises or dragonflies , have specialized zones of ommatidia organized into a fovea area which gives acute vision. In the acute zone the eye is flattened and the facets larger. The flattening allows more ommatidia to receive light from a spot and therefore higher resolution. There are some exceptions from
174-408: A way that resembles a true compound eye. Asymmetries in compound eyes may be associated with asymmetries in behaviour. For example, Temnothorax albipennis ant scouts show behavioural lateralization when exploring unknown nest sites, showing a population-level bias to prefer left turns. One possible reason for this is that its environment is partly maze-like and consistently turning in one direction
203-491: Is a good way to search and exit mazes without getting lost. This turning bias is correlated with slight asymmetries in the ants' compound eyes (differential ommatidia count). The body of Ophiomastix wendtii , a type of brittle star , was previously thought to be covered with ommatidia, turning its whole skin into a compound eye, but this has since been found to be erroneous; the system does not rely on lenses or image formation. "Dragonfly eyes" (Chinese: 蜻蜓眼 qingting yan ]
232-475: Is a single large facet that is three times in diameter the others in the eye and behind this is an enlarged crystalline cone. This projects an upright image on a specialized retina. The resulting eye is a mixture of a simple eye within a compound eye. Another version is the pseudofaceted eye, as seen in Scutigera . This type of eye consists of a cluster of numerous ocelli on each side of the head, organized in
261-423: Is constrained by the inelastic exoskeleton, so development involves a series of molts . The immature stages often differ from the adults in structure, habit and habitat. Groups that undergo four-stage metamorphosis often have a nearly immobile pupa . Insects that undergo three-stage metamorphosis lack a pupa, developing through a series of increasingly adult-like nymphal stages. The higher level relationship of
290-401: Is excreted and that alone can be a second, major source of damage. The third and potentially most harmful characteristic is the ability of adults to transmit several plant viruses . The crop hosts principally affected are vegetables such as cucurbits , potatoes and tomatoes , although a range of other crop and non-crop plants including weed species are susceptible, and can therefore harbour
319-853: The Latin word insectum from in , "cut up", as insects appear to be cut into three parts. The Latin word was introduced by Pliny the Elder who calqued the Ancient Greek word ἔντομον éntomon "insect" (as in entomology ) from ἔντομος éntomos "cut in pieces"; this was Aristotle 's term for this class of life in his biology , also in reference to their notched bodies. The English word insect first appears in 1601 in Philemon Holland 's translation of Pliny. In common speech, insects and other terrestrial arthropods are often called bugs . Entomologists to some extent reserve
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#1732798206868348-2342: The Paraneoptera , and Kjer et al. 2016 for the Holometabola . The numbers of described extant species (boldface for groups with over 100,000 species) are from Stork 2018. Archaeognatha (hump-backed/jumping bristletails, 513 spp) [REDACTED] Zygentoma (silverfish, firebrats, fishmoths, 560 spp) [REDACTED] Odonata (dragonflies and damselflies, 5,899 spp) [REDACTED] Ephemeroptera (mayflies, 3,240 spp) [REDACTED] Zoraptera (angel insects, 37 spp) [REDACTED] Dermaptera (earwigs, 1,978 spp) [REDACTED] Plecoptera (stoneflies, 3,743 spp) [REDACTED] Orthoptera (grasshoppers, crickets, katydids, 23,855 spp) [REDACTED] Grylloblattodea (ice crawlers, 34 spp) [REDACTED] Mantophasmatodea (gladiators, 15 spp) [REDACTED] Phasmatodea (stick insects, 3,014 spp) [REDACTED] Embioptera (webspinners, 463 spp) [REDACTED] Mantodea (mantises, 2,400 spp) [REDACTED] Blattodea (cockroaches and termites, 7,314 spp) [REDACTED] Psocodea (book lice, barklice and sucking lice, 11,000 spp) [REDACTED] [REDACTED] Hemiptera (true bugs, 103,590 spp) [REDACTED] Thysanoptera (thrips, 5,864 spp) [REDACTED] Hymenoptera (sawflies, wasps, bees, ants, 116,861 spp) [REDACTED] Strepsiptera (twisted-wing flies, 609 spp) [REDACTED] Coleoptera (beetles, 386,500 spp) [REDACTED] Raphidioptera (snakeflies, 254 spp) [REDACTED] Neuroptera (lacewings, 5,868 spp) [REDACTED] Megaloptera (alderflies and dobsonflies, 354 spp) [REDACTED] Lepidoptera (butterflies and moths, 157,338 spp) [REDACTED] Trichoptera (caddisflies, 14,391 spp) [REDACTED] Diptera (true flies, 155,477 spp) [REDACTED] Mecoptera (scorpionflies, 757 spp) [REDACTED] Siphonaptera (fleas, 2,075 spp) [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] Compound eye Compound eyes are typically classified as either apposition eyes, which form multiple inverted images, or superposition eyes, which form
377-623: The insects is unclear. Fossilized insects of enormous size have been found from the Paleozoic Era, including giant dragonfly-like insects with wingspans of 55 to 70 cm (22 to 28 in). The most diverse insect groups appear to have coevolved with flowering plants . Adult insects typically move about by walking and flying; some can swim. Insects are the only invertebrates that can achieve sustained powered flight; insect flight evolved just once. Many insects are at least partly aquatic , and have larvae with gills; in some species,
406-483: The pheromones of female moths over great distances. Other species communicate with sounds: crickets stridulate , or rub their wings together, to attract a mate and repel other males. Lampyrid beetles communicate with light. Humans regard many insects as pests , especially those that damage crops, and attempt to control them using insecticides and other techniques. Others are parasitic , and may act as vectors of diseases . Insect pollinators are essential to
435-623: The Hemiptera (true bugs), Lepidoptera (butterflies and moths), Diptera (true flies), Hymenoptera (wasps, ants, and bees), and Coleoptera (beetles), each with more than 100,000 described species. Insects are distributed over every continent and almost every terrestrial habitat. There are many more species in the tropics , especially in rainforests , than in temperate zones. The world's regions have received widely differing amounts of attention from entomologists. The British Isles have been thoroughly surveyed, so that Gullan and Cranston 2014 state that
464-404: The adults too are aquatic. Some species, such as water striders , can walk on the surface of water. Insects are mostly solitary, but some, such as bees , ants and termites , are social and live in large, well-organized colonies . Others, such as earwigs , provide maternal care, guarding their eggs and young. Insects can communicate with each other in a variety of ways. Male moths can sense
493-470: The images are combined in the brain. This is called the schizochroal compound eye or the neural superposition eye (which, despite its name, is a form of the apposition eye). The superposition eye is divided into three subtypes; the refracting , the reflecting , and the parabolic superposition eye. The refracting superposition eye has a gap between the lens and the rhabdom, and no side wall. Each lens takes light at an angle to its axis and reflects it to
522-443: The infection. Effective control has been provided for many years through the release of beneficial insects, such as the aphelinid parasitoid, Encarsia formosa ( Gahan ). If required, integrated pest management strategies can incorporate applications of selective chemical insecticides or biopesticides such as Lecanicillium muscarium that complement these natural enemies. For the majority of outdoor crops chemicals are still
551-608: The insects among the Hexapoda , six-legged animals with segmented bodies; their closest relatives are the Diplura (bristletails). Collembola (springtails) [REDACTED] Protura (coneheads) [REDACTED] Diplura (two-pronged bristletails) [REDACTED] Insecta (=Ectognatha) [REDACTED] The internal phylogeny is based on the works of Wipfler et al. 2019 for the Polyneoptera , Johnson et al. 2018 for
580-461: The leaf surface. Females are capable of mating less than 24 hours after emergence and most frequently lay their eggs on the undersides of leaves. Eggs are pale yellow in colour, before turning grey prior to hatching. Newly hatched nymphs , often known as crawlers, are the only mobile immature life-stage. During the first and second nymph instars, the appearance is that of a pale yellow/translucent, flat scale which can be difficult to distinguish with
609-466: The most widely used method of control. To study pest resistance management a 787‐Mb high‐quality draft genome has been sequenced and assembled. Insect Insects (from Latin insectum ) are hexapod invertebrates of the class Insecta . They are the largest group within the arthropod phylum . Insects have a chitinous exoskeleton , a three-part body ( head , thorax and abdomen ), three pairs of jointed legs , compound eyes , and
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#1732798206868638-414: The naked eye. During the fourth and final immature life-stage, referred to as the " pupa ", compound eyes and other body tissues become visible as the nymph thickens and rises from the leaf-surface. All life-stages apart from eggs and "pupae" cause crop damage through direct feeding, inserting their stylet into leaf veins and extracting nourishment from the phloem sap. As a by-product of feeding, honeydew
667-402: The name "bugs" for a narrow category of " true bugs ", insects of the order Hemiptera , such as cicadas and shield bugs . Other terrestrial arthropods, such as centipedes , millipedes , woodlice , spiders , mites and scorpions , are sometimes confused with insects, since they have a jointed exoskeleton. Adult insects are the only arthropods that ever have wings, with up to two pairs on
696-535: The reproduction of many flowering plants and so to their ecosystems. Many insects are ecologically beneficial as predators of pest insects, while a few provide direct economic benefit. Two species in particular are economically important and were domesticated many centuries ago: silkworms for silk and honey bees for honey . Insects are consumed as food in 80% of the world's nations, by people in roughly 3000 ethnic groups. Human activities are having serious effects on insect biodiversity . The word insect comes from
725-512: The same angle on the other side. The result is an image at half the radius of the eye, which is where the tips of the rhabdoms are. This kind is used mostly by nocturnal insects. In the parabolic superposition eye , seen in arthropods such as mayflies , the parabolic surfaces of the inside of each facet focus light from a reflector to a sensor array . Long-bodied decapod crustaceans such as shrimp , prawns , crayfish and lobsters are alone in having reflecting superposition eyes , which also have
754-527: The thorax. Whether winged or not, adult insects can be distinguished by their three-part body plan, with head, thorax, and abdomen; they have three pairs of legs on the thorax. Estimates of the total number of insect species vary considerably, suggesting that there are perhaps some 5.5 million insect species in existence, of which about one million have been described and named. These constitute around half of all eukaryote species, including animals , plants , and fungi . The most diverse insect orders are
783-466: The tissues. The blood therefore does not carry oxygen; it is only partly contained in vessels, and some circulates in an open hemocoel . Insect vision is mainly through their compound eyes , with additional small ocelli . Many insects can hear, using tympanal organs , which may be on the legs or other parts of the body. Their sense of smell is via receptors, usually on the antennae and the mouthparts. Nearly all insects hatch from eggs . Insect growth
812-525: The total of around 22,500 species is probably within 5% of the actual number there; they comment that Canada's list of 30,000 described species is surely over half of the actual total. They add that the 3000 species of the American Arctic must be broadly accurate. In contrast, a large majority of the insect species of the tropics and the southern hemisphere are probably undescribed. Some 30–40,000 species inhabit freshwater ; very few insects, perhaps
841-405: The types mentioned above. Some insects have a so-called single lens compound eye, a transitional type which is something between a superposition type of the multi-lens compound eye and the single lens eye found in animals with simple eyes. Then there is the mysid shrimp, Dioptromysis paucispinosa . The shrimp has an eye of the refracting superposition type, in the rear behind this in each eye there
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