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92-536: These vast limestone passages, that had formed during the Middle Triassic and are rich in speleothems were discovered in 1926 by Ján Majko. A 1,600 m (5,200 ft) section of its total length of 5,140 m (16,860 ft) has been publicly accessible since 1932. As an element of the Caves of Aggtelek Karst and Slovak Karst the site was listed as a UNESCO World Heritage Site list in 1995 because of

184-630: A bolide impact, for which an impact crater containing Manicouagan Reservoir in Quebec , Canada , has been singled out. However, the Manicouagan impact melt has been dated to 214±1 Mya. The date of the Triassic-Jurassic boundary has also been more accurately fixed recently, at 201.4 Mya. Both dates are gaining accuracy by using more accurate forms of radiometric dating, in particular the decay of uranium to lead in zircons formed at time of

276-626: A cosmopolitan distribution . Coelacanths show their highest post- Devonian diversity in the Early Triassic . Ray-finned fishes (actinopterygians) went through a remarkable diversification in the beginning of the Triassic, leading to peak diversity during the Middle Triassic; however, the pattern of this diversification is still not well understood due to a taphonomic megabias . The first stem-group teleosts appeared during

368-502: A disaster taxon (although this view was questioned ), Lystrosaurus had a wide range across Pangea. In the southern part of the supercontinent, it co-occurred with the non-mammalian cynodonts Galesaurus and Thrinaxodon , early relatives of mammals . The first archosauriforms appeared, such as Erythrosuchus ( Olenekian - Ladinian ). This group includes the ancestors of crocodiles and dinosaurs (including birds ). Fossilized foot prints of dinosauromorphs are known from

460-555: A chain of mountain ranges stretching from Turkey to Malaysia . Pangaea was fractured by widespread faulting and rift basins during the Triassic—especially late in that period—but had not yet separated. The first nonmarine sediments in the rift that marks the initial break-up of Pangaea, which separated eastern North America from Morocco , are of Late Triassic age; in the United States , these thick sediments comprise

552-474: A few exposures in the west. During the Triassic peneplains are thought to have formed in what is now Norway and southern Sweden. Remnants of this peneplain can be traced as a tilted summit accordance in the Swedish West Coast . In northern Norway Triassic peneplains may have been buried in sediments to be then re-exposed as coastal plains called strandflats . Dating of illite clay from

644-544: A fine-grained loam on the Styx bank with imprints of stone axes are evidences of ceramics manufacture in the cave. Irons, awls, arrows, the oldest comb in Europe, ring, decorated cylinder bracelet and fishhook represent the peak of Neolithic processing of bones. Also pendants from shells and animal teeth were preserved. Triassic The Triassic ( / t r aɪ ˈ æ s ɪ k / try- ASS -ik ; sometimes symbolized 🝈 )

736-471: A long beak-like snout), and Shringasaurus (a horned herbivore which reached a body length of 3–4 metres (9.8–13.1 ft)). One group of archosauromorphs, the archosauriforms , were distinguished by their active predatory lifestyle, with serrated teeth and upright limb postures. Archosauriforms were diverse in the Triassic, including various terrestrial and semiaquatic predators of all shapes and sizes. The large-headed and robust erythrosuchids were among

828-522: A number of the local caves are simply parts of a bigger entity were corroborated by the explorer I.Vass, who surveyed the Baradla cave in 1821. E.Nyari took up archaeological work in 1881 and suggested a connection of the Certova diera and Baradla Cave as did K. Siegmenth in 1891. Only the extensive work of Jan Majko in 1926 confirmed the interconnection of several local caves and lead to the discovery of

920-617: A pseudosuchian. Pseudosuchians were far more ecologically dominant in the Triassic, including large herbivores (such as aetosaurs ), large carnivores (" rauisuchians "), and the first crocodylomorphs (" sphenosuchians "). Aetosaurs were heavily-armored reptiles that were common during the last 30 million years of the Late Triassic until they died out at the Triassic-Jurassic extinction. Most aetosaurs were herbivorous and fed on low-growing plants, but some may have eaten meat. " rauisuchians " (formally known as paracrocodylomorphs ) were

1012-466: A rich set of various conditions, that affect sedimentation, evolution and fossilization in a geological time frame. Jaroslav Böhm directed excavations during the 1930s. Human presence has been documented dating back to the Paleolithic . The cave had presumably been a casual refuge for the oldest Neolithic inhabitants of Eastern Slovakia. However, the cave has been frequented by Neolithic humans of

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1104-458: A short period of time, becoming extinct about 220 million years ago. They were exceptionally abundant in the middle of the Triassic, as the primary large herbivores in many Carnian-age ecosystems. They sheared plants with premaxillary beaks and plates along the upper jaw with multiple rows of teeth. Allokotosaurs were iguana-like reptiles, including Trilophosaurus (a common Late Triassic reptile with three-crowned teeth), Teraterpeton (which had

1196-567: A specialized subgroup of cynodonts, appeared during the Triassic and would survive the extinction event, allowing them to radiate during the Jurassic. Amphibians were primarily represented by the temnospondyls , giant aquatic predators that had survived the end-Permian extinction and saw a new burst of diversification in the Triassic, before going extinct by the end; however, early crown-group lissamphibians (including stem-group frogs , salamanders and caecilians ) also became more common during

1288-449: A strandflat of Bømlo , southern Norway, have shown that landscape there became weathered in Late Triassic times ( c. 210 million years ago) with the landscape likely also being shaped during that time. Eustatic sea level in the Triassic was consistently low compared to the other geological periods. The beginning of the Triassic was around present sea level, rising to about 10–20 metres (33–66 ft) above present-day sea level during

1380-585: A supercontinent has less shoreline compared to a series of smaller continents, Triassic marine deposits are relatively uncommon on a global scale. A major exception is in Western Europe , where the Triassic was first studied. The northeastern margin of Gondwana was a stable passive margin along the Neo-Tethys Ocean, and marine sediments have been preserved in parts of northern India and Arabia . In North America , marine deposits are limited to

1472-659: Is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya ), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is the first and shortest period of the Mesozoic Era and the seventh period of the Phanerozoic Eon . Both the start and end of the period are marked by major extinction events . The Triassic Period

1564-468: Is a recent study of North American faunas. In the Petrified Forest of northeast Arizona there is a unique sequence of late Carnian-early Norian terrestrial sediments. An analysis in 2002 found no significant change in the paleoenvironment. Phytosaurs , the most common fossils there, experienced a change-over only at the genus level, and the number of species remained the same. Some aetosaurs ,

1656-508: Is likely a paraphyletic group rather than a true clade. Tanystropheids were a family of protorosaurs which elevated their neck size to extremes, with the largest genus Tanystropheus having a neck longer than its body. The protorosaur family Sharovipterygidae used their elongated hindlimbs for gliding. Other archosauromorphs, such as rhynchosaurs and allokotosaurs , were mostly stocky-bodied herbivores with specialized jaw structures. Rhynchosaurs, barrel-gutted herbivores, thrived for only

1748-469: Is no evidence of glaciation at or near either pole; in fact, the polar regions were apparently moist and temperate , providing a climate suitable for forests and vertebrates, including reptiles. Pangaea's large size limited the moderating effect of the global ocean; its continental climate was highly seasonal, with very hot summers and cold winters. The strong contrast between the Pangea supercontinent and

1840-663: Is omnipresent and very impressive, including cascades of stalactite lakes, variously shaped stalactites domed temples and notably bat guano , that had diffused into sinter crusts accounts for unique forms and textures of surface decoration. Karst processes have over tens of millions of years created diverse structures and habitats that date back to the Late Cretaceous. Subsequent periods of temperate, glacial, subtropical and tropical climatic conditions excellently demonstrate karst formation under varying conditions. Biologists, geologists and paleontologists are confronted with

1932-834: Is preceded by the Lopingian Epoch (late Permian , Paleozoic Era ) and followed by the Middle Triassic Epoch. The Early Triassic is divided into the Induan and Olenekian ages . The Induan is subdivided into the Griesbachian and Dienerian subages and the Olenekian is subdivided into the Smithian and Spathian subages. The Lower Triassic series is coeval with the Scythian Stage, which

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2024-605: Is subdivided into three epochs: Early Triassic , Middle Triassic and Late Triassic . The Triassic began in the wake of the Permian–Triassic extinction event , which left the Earth's biosphere impoverished; it was well into the middle of the Triassic before life recovered its former diversity. Three categories of organisms can be distinguished in the Triassic record: survivors from the extinction event, new groups that flourished briefly, and other new groups that went on to dominate

2116-460: Is superimposed by 22 sea level drop events widespread in the geologic record, mostly of minor (less than 25-metre (82 ft)) and medium (25–75-metre (82–246 ft)) magnitudes. A lack of evidence for Triassic continental ice sheets suggest that glacial eustasy is unlikely to be the cause of these changes. The Triassic continental interior climate was generally hot and dry, so that typical deposits are red bed sandstones and evaporites . There

2208-638: Is today not included in the official timescales but can be found in older literature. In Europe, most of the Lower Triassic is composed of Buntsandstein , a lithostratigraphic unit of continental red beds . The Early Triassic and partly also the Middle Triassic span the interval of biotic recovery from the Permian-Triassic extinction event , the most severe mass extinction event in Earth's history. A second extinction event,

2300-411: Is usually divided into Early , Middle , and Late Triassic Epochs , and the corresponding rocks are referred to as Lower, Middle, or Upper Triassic. The faunal stages from the youngest to oldest are: During the Triassic, almost all the Earth's land mass was concentrated into a single supercontinent , Pangaea ( lit.   ' entire land ' ). This supercontinent was more-or-less centered on

2392-529: The Bükk culture , who might even have manufactured their distinct thin-walled pottery inside the cave. The original entrance of the Domica Cave was blocked by debris after Paleo-humans had abandoned it and the cave became inaccessible. Post holes from dwelling objects and fireplaces were discovered in several places of the cave. More than 200 reconstructed containers from sherds as well as a terrace-dug slope in

2484-578: The Carnian (early part of the Late Triassic), some advanced cynodonts gave rise to the first mammals . During the Triassic, archosaurs displaced therapsids as the largest and most ecologically prolific terrestrial amniotes. This "Triassic Takeover" may have contributed to the evolution of mammals by forcing the surviving therapsids and their mammaliaform successors to live as small, mainly nocturnal insectivores . Nocturnal life may have forced

2576-614: The Jurassic , when the temnospondyls had become very rare. Most of the Reptiliomorpha , stem-amniotes that gave rise to the amniotes, disappeared in the Triassic, but two water-dwelling groups survived: Embolomeri that only survived into the early part of the period, and the Chroniosuchia , which survived until the end of the Triassic. The Permian–Triassic extinction devastated terrestrial life. Biodiversity rebounded as

2668-504: The Lake Lugano region of northern Italy and southern Switzerland , was in Middle Triassic times a lagoon behind reefs with an anoxic bottom layer, so there were no scavengers and little turbulence to disturb fossilization, a situation that can be compared to the better-known Jurassic Solnhofen Limestone lagerstätte . The remains of fish and various marine reptiles (including the common pachypleurosaur Neusticosaurus , and

2760-509: The Mesozoic Era. Reptiles , especially archosaurs , were the chief terrestrial vertebrates during this time. A specialized group of archosaurs, called dinosaurs , first appeared in the Late Triassic but did not become dominant until the succeeding Jurassic Period. Archosaurs that became dominant in this period were primarily pseudosuchians , relatives and ancestors of modern crocodilians , while some archosaurs specialized in flight,

2852-658: The Newark Supergroup . Rift basins are also common in South America, Europe, and Africa. Terrestrial environments are particularly well-represented in the South Africa, Russia, central Europe, and the southwest United States. Terrestrial Triassic biostratigraphy is mostly based on terrestrial and freshwater tetrapods, as well as conchostracans ("clam shrimps"), a type of fast-breeding crustacean which lived in lakes and hypersaline environments. Because

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2944-474: The Olenekian and Anisian of Gondwana . Both kannemeyeriiform dicynodonts and gomphodont cynodonts remained important herbivores during much of the period. Therocephalians included both large predators ( Moschorhinus ) and herbivorous forms ( bauriids ) until their extinction midway through the period. Ecteniniid cynodonts played a role as large-sized, cursorial predators in the Late Triassic. During

3036-547: The Permian-Triassic extinction event . On the other hand, an alternative hypothesis proposes these Early Triassic climatic perturbations and biotic upheavals that inhibited the recovery of life following the P-T mass extinction to have been linked to forcing driven by changes in the Earth's obliquity defined by a roughly 32.8 thousand year periodicity with strong 1.2 million year modulations. According to proponents of this hypothesis, radiometric dating indicates that major activity from

3128-474: The Siberian Traps , which had probably triggered the Permian-Triassic extinction event and accelerated the rate of global warming into the Triassic. Studies suggest that Early Triassic climate was very volatile, punctuated by a number of relatively rapid global temperature changes, marine anoxic events, and carbon cycle disturbances, which led to subsequent extinction events in the aftermath of

3220-723: The Smithian-Spathian boundary event , occurred during the Olenekian. A third extinction event occurred at the Olenekian-Anisian boundary, marking the end of the Early Triassic epoch. The climate during the Early Triassic Epoch (especially in the interior of the supercontinent Pangaea ) was generally arid, rainless and dry and deserts were widespread; however the poles possessed a temperate climate . The pole-to-equator temperature gradient

3312-531: The Spathian subage, the flora changed back to gymnosperm and pteridophyte dominated. These shifts reflect global changes in precipitation and temperature. Floral diversity was overall very low during the Early Triassic, as plant life had yet to fully recover from the Permian-Triassic extinction. Microbially induced sedimentary structures (MISS) are common in the fossil record of North China in

3404-604: The surviving species repopulated empty terrain, but these were short-lived. Diverse communities with complex food-web structures took 30 million years to reestablish. Archosauromorph reptiles, which had already appeared and diversified to an extent in the Permian Period, exploded in diversity as an adaptive radiation in response to the Permian-Triassic mass extinction. By the Early Triassic, several major archosauromorph groups had appeared. Long-necked, lizard-like early archosauromorphs were known as protorosaurs , which

3496-468: The thecodonts ) disappeared, as did most of the large labyrinthodont amphibians, groups of small reptiles, and most synapsids. Some of the early, primitive dinosaurs also became extinct, but more adaptive ones survived to evolve into the Jurassic. Surviving plants that went on to dominate the Mesozoic world included modern conifers and cycadeoids. The cause of the Late Triassic extinction is uncertain. It

3588-437: The traversodont cynodonts—were much reduced in the northern half of Pangaea ( Laurasia ). These extinctions within the Triassic and at its end allowed the dinosaurs to expand into many niches that had become unoccupied. Dinosaurs became increasingly dominant, abundant and diverse, and remained that way for the next 150 million years. The true "Age of Dinosaurs" is during the following Jurassic and Cretaceous periods, rather than

3680-832: The Anisian to Ladinian of the Tethysian domain, and from the Carnian and Rhaetian of a larger area that includes also the Boreal domain (e.g., Svalbard Islands), the North American continent, the South China block and Argentina . The best-studied of such episodes of humid climate, and probably the most intense and widespread, was the Carnian Pluvial Event . The Early Triassic was the hottest portion of

3772-405: The Carnian and include early sauropodomorphs and theropods. Most Triassic dinosaurs were small predators and only a few were common, such as Coelophysis , which was 1 to 2 metres (3.3 to 6.6 ft) long. Triassic sauropodomorphs primarily inhabited cooler regions of the world. The large predator Smok was most likely also an archosaur, but it is uncertain if it was a primitive dinosaur or

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3864-513: The Domica cave on October 3, 1926. J. Majko penetrated from its bottom through a 15 m deep abyss into large underground spaces in 1926 where plenty of archaeological discoveries were made later. The Domica Cave is situated in the Slovak karst and consists of light limestone of the Middle Triassic . The passages were shaped by the waters of an underground river, called Styx . Speleothem decoration

3956-483: The Early Triassic has been interpreted as a signal of increased bioturbation and recovery of terrestrial ecosystems. In the oceans, the most common Early Triassic hard-shelled marine invertebrates were bivalves , gastropods , ammonoids , echinoids , and a few articulate brachiopods . Conodonts experienced a revival in diversity following a nadir during the Permian. The first oysters ( Liostrea ) appeared in

4048-508: The Early Triassic, forming small patches of reefs of modest extent compared to the great reef systems of Devonian or modern times. At the end of the Carnian, a reef crisis occurred in South China. Serpulids appeared in the Middle Triassic. Microconchids were abundant. The shelled cephalopods called ammonites recovered, diversifying from a single line that survived the Permian extinction. Bivalves began to rapidly diversify during

4140-402: The Early Triassic, while others (e.g. capitosaurs ) remained successful throughout the whole period, or only came to prominence in the Late Triassic (e.g. Plagiosaurus , metoposaurs ). The first Lissamphibians (modern amphibians) appear in the Triassic, with the progenitors of the first frogs already present by the Early Triassic. However, the group as a whole did not become common until

4232-439: The Early Triassic. They grew on the shells of living ammonoids as epizoans. Microbial reefs were common, possibly due to lack of competition with metazoan reef builders as a result of the extinction. However, transient metazoan reefs reoccurred during the Olenekian wherever permitted by environmental conditions. Ammonoids show blooms followed by extinctions during the Early Triassic. Aquatic vertebrates diversified after

4324-562: The Early and Middle Triassic. Sea level rise accelerated in the Ladinian, culminating with a sea level up to 50 metres (164 ft) above present-day levels during the Carnian. Sea level began to decline in the Norian, reaching a low of 50 metres (164 ft) below present sea level during the mid-Rhaetian. Low global sea levels persisted into the earliest Jurassic. The long-term sea level trend

4416-403: The Jurassic. The Triassic was named in 1834 by Friedrich August von Alberti , after a succession of three distinct rock layers (Greek triás meaning 'triad') that are widespread in southern Germany : the lower Buntsandstein (colourful sandstone ) , the middle Muschelkalk (shell-bearing limestone ) and the upper Keuper (coloured clay ). On the geologic time scale , the Triassic

4508-613: The Jurassic. There were many types of marine reptiles. These included the Sauropterygia , which featured pachypleurosaurus and nothosaurs (both common during the Middle Triassic, especially in the Tethys region), placodonts , the earliest known herbivorous marine reptile Atopodentatus , and the first plesiosaurs . The first of the lizardlike Thalattosauria ( askeptosaurs ) and the highly successful ichthyopterygians , which appeared in Early Triassic seas, soon diversified. By

4600-600: The Latest Olenekian Cooling (LOC), from 248 to 247 Ma, temperatures cooled by about 6 °C. The Middle Triassic was cooler than the Early Triassic, with temperatures falling over most of the Anisian, with the exception of a warming spike in the latter portion of the stage. From 242 to 233 Ma, the Ladinian-Carnian Cooling (LCC) ensued. At the beginning of the Carnian, global temperatures continued to be relatively cool. The eruption of

4692-420: The Middle Triassic, becoming highly abundant in the oceans. Aquatic insects rapidly diversified during the Middle Triassic, with this time interval representing a crucial diversification for Holometabola , the clade containing the majority of modern insect species. In the wake of the Permian-Triassic mass extinction event , the fish fauna was remarkably uniform, with many families and genera exhibiting

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4784-618: The Middle Triassic, some ichthyopterygians were achieving very large body masses. Among other reptiles, the earliest turtles , like Proganochelys and Proterochersis , appeared during the Norian Age (Stage) of the Late Triassic Period. The Lepidosauromorpha , specifically the Sphenodontia , are first found in the fossil record of the earlier Carnian Age, though the earliest lepidosauromorphs likely occurred in

4876-562: The Olenekian. The Early Triassic entomofauna is very poorly understood because of the paucity of insect fossils from this epoch. The flora was gymnosperm -dominated at the onset of the Triassic, but changed rapidly and became lycopod -dominated (e.g. Pleuromeia ) during the Griesbachian-Dienerian ecological crisis . This change coincided with the extinction of the Permian Glossopteris flora. In

4968-450: The Permian extinction, Archaeplastida (red and green algae) had been the major marine phytoplanktons since about 659–645 million years ago, when they replaced marine planktonic cyanobacteria , which first appeared about 800 million years ago, as the dominant phytoplankton in the oceans. In the Triassic, secondary endosymbiotic algae became the most important plankton. In marine environments , new modern types of corals appeared in

5060-475: The Permian. The Procolophonidae , the last surviving parareptiles , were an important group of small lizard-like herbivores. The drepanosaurs were a clade of unusual, chameleon-like arboreal reptiles with birdlike heads and specialised claws. Three therapsid groups survived into the Triassic: dicynodonts , therocephalians , and cynodonts . The cynodont Cynognathus was a characteristic top predator in

5152-608: The Siberian Traps ended very shortly after the end-Permian extinction and did not span the entire Early Triassic epoch, thus not being the primary culprit for the climatic changes throughout this epoch. The Triassic Period opened in the aftermath of the Permian–Triassic extinction event . The massive extinctions that ended the Permian Period (and with that the Paleozoic Era ) caused extreme hardships for

5244-524: The Southern Hemisphere, ceased to exist. Other groups, such as Actinopterygii , appear to have been less affected by this extinction event and body size was not a selective factor during the extinction event. Animals that were most successful in the Early Triassic were those with high metabolisms. Different patterns of recovery are evident on land and in the sea. Early Triassic faunas lacked biodiversity and were relatively homogeneous due to

5336-408: The Triassic (teleosts are by far the most diverse group of fish today). Predatory actinopterygians such as saurichthyids and birgeriids , some of which grew over 1.2 m (3.9 ft) in length, appeared in the Early Triassic and became widespread and successful during the period as a whole. Lakes and rivers were populated by lungfish (Dipnoi), such as Ceratodus , which are mainly known from

5428-497: The Triassic and survived the extinction event. The earliest known neopterygian fish, including early holosteans and teleosts , appeared near the beginning of the Triassic, and quickly diversified to become among the dominant groups of fish in both freshwater and marine habitats. The vast supercontinent of Pangaea dominated the globe during the Triassic, but in the latest Triassic ( Rhaetian ) and Early Jurassic it began to gradually rift into two separate landmasses: Laurasia to

5520-609: The Triassic, enlarging the Neo-Tethys Ocean which formed in their wake. At the same time, they forced the Paleo-Tethys Ocean to shrink as it was being subducted under Asia. By the end of the Triassic, the Paleo-Tethys Ocean occupied a small area and the Cimmerian terranes began to collide with southern Asia. This collision, known as the Cimmerian Orogeny , continued into the Jurassic and Cretaceous to produce

5612-601: The Triassic. Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale . It spans the time between 251.9 Ma and 247.2 Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series , which is a unit in chronostratigraphy . The Early Triassic is the oldest epoch of the Mesozoic Era . It

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5704-839: The Wrangellia Large Igneous Province around 234 Ma caused abrupt global warming, terminating the cooling trend of the LCC. This warming was responsible for the Carnian Pluvial Event and resulted in an episode of widespread global humidity. The CPE ushered in the Mid-Carnian Warm Interval (MCWI), which lasted from 234 to 227 Ma. At the Carnian-Norian boundary occurred a positive δ C excursion believed to signify an increase in organic carbon burial. From 227 to 217 Ma, there

5796-401: The bizarre long-necked archosauromorph Tanystropheus ), along with some terrestrial forms like Ticinosuchus and Macrocnemus , have been recovered from this locality. All these fossils date from the Anisian and Ladinian ages (about 242 Ma ago). The Triassic Period ended with a mass extinction, which was particularly severe in the oceans; the conodonts disappeared, as did all

5888-447: The dental plates, abundant in the fossils record. Hybodonts , a group of shark-like cartilaginous fish , were dominant in both freshwater and marine environments throughout the Triassic. Last survivors of the mainly Palaeozoic Eugeneodontida are known from the Early Triassic. Temnospondyl amphibians were among those groups that survived the Permian–Triassic extinction. Once abundant in both terrestrial and aquatic environments,

5980-427: The diversity of cave features found within it and its record of both tropical and temperate effects on karst cave formation. The site belongs to a complex variety of karstic formations that include 712 caves, many of which were well known among the local population for centuries. Bartholomeides, a priest from Ochtiná , documented his first systematic excursions into the Certova diera Cave in 1801. His conclusions that

6072-428: The dominant carnivores in the early Triassic. Phytosaurs were a particularly common group which prospered during the Late Triassic. These long-snouted and semiaquatic predators resemble living crocodiles and probably had a similar lifestyle, hunting for fish and small reptiles around the water's edge. However, this resemblance is only superficial and is a prime-case of convergent evolution. True archosaurs appeared in

6164-405: The early Triassic, splitting into two branches: Avemetatarsalia (the ancestors to birds) and Pseudosuchia (the ancestors to crocodilians). Avemetatarsalians were a minor component of their ecosystems, but eventually produced the earliest pterosaurs and dinosaurs in the Late Triassic. Early long-tailed pterosaurs appeared in the Norian and quickly spread worldwide. Triassic dinosaurs evolved in

6256-408: The effects of the extinction. The ecological recovery on land took 30 million years, well into the Late Triassic . Two Early Triassic lagerstätten stand out due to their exceptionally high biodiversity , the Dienerian aged Guiyang biota and the earliest Spathian aged Paris biota . The most common land vertebrate was the small herbivorous synapsid Lystrosaurus . Often interpreted as

6348-401: The entire Phanerozoic, seeing as it occurred during and immediately after the discharge of titanic volumes of greenhouse gases from the Siberian Traps. The Early Triassic began with the Permian-Triassic Thermal Maximum (PTTM) and was followed by the brief Dienerian Cooling (DC) from 251 to 249 Ma, which was in turn followed by the Latest Smithian Thermal Maximum (LSTT) around 249 to 248 Ma. During

6440-467: The equator and extended between the poles, though it did drift northwards as the period progressed. Southern Pangea, also known as Gondwana , was made up by closely-appressed cratons corresponding to modern South America , Africa , Madagascar , India , Antarctica , and Australia . North Pangea, also known as Laurussia or Laurasia , corresponds to modern-day North America and the fragmented predecessors of Eurasia . The western edge of Pangea lay at

6532-415: The extinct family Cheirolepidiaceae , which first appeared in the Late Triassic, and would be prominent throughout most of the rest of the Mesozoic. No known coal deposits date from the start of the Triassic Period. This is known as the Early Triassic "coal gap" and can be seen as part of the Permian–Triassic extinction event . Possible explanations for the coal gap include sharp drops in sea level at

6624-406: The first time among vertebrates, becoming the pterosaurs . Therapsids , the dominant vertebrates of the preceding Permian period, saw a brief surge in diversification in the Triassic, with dicynodonts and cynodonts quickly becoming dominant, but they declined throughout the period with the majority becoming extinct by the end. However, the first stem-group mammals ( mammaliamorphs ), themselves

6716-521: The global ocean triggered intense cross-equatorial monsoons , sometimes referred to as the Pangean megamonsoons . The Triassic may have mostly been a dry period, but evidence exists that it was punctuated by several episodes of increased rainfall in tropical and subtropical latitudes of the Tethys Sea and its surrounding land. Sediments and fossils suggestive of a more humid climate are known from

6808-509: The immediate aftermath of the Permian-Triassic extinction, indicating that microbial mats dominated local terrestrial ecosystems following the Permian-Triassic boundary. The regional prevalence of MISS is attributable to a decrease in bioturbation and grazing pressure as a result of aridification and temperature increase. MISS have also been reported from Early Triassic fossil deposits in Arctic Canada. The disappearance of MISS later in

6900-420: The impact. So, the evidence suggests the Manicouagan impact preceded the end of the Triassic by approximately 10±2 Ma. It could not therefore be the immediate cause of the observed mass extinction. The number of Late Triassic extinctions is disputed. Some studies suggest that there are at least two periods of extinction towards the end of the Triassic, separated by 12 to 17 million years. But arguing against this

6992-401: The keystone predators of most Triassic terrestrial ecosystems. Over 25 species have been found, including giant quadrupedal hunters, sleek bipedal omnivores, and lumbering beasts with deep sails on their backs. They probably occupied the large-predator niche later filled by theropods. "Rauisuchians" were ancestral to small, lightly-built crocodylomorphs, the only pseudosuchians which survived into

7084-545: The mammaliaforms to develop fur and a higher metabolic rate . Two Early Triassic lagerstätten (high-quality fossil beds), the Dienerian aged Guiyang biota and the earliest Spathian aged Paris biota stand out due to their exceptional preservation and diversity . They represent the earliest lagerstätten of the Mesozoic era and provide insight into the biotic recovery from the Permian-Triassic mass extinction event. The Monte San Giorgio lagerstätte, now in

7176-573: The margin of an enormous ocean, Panthalassa ( lit.   ' entire sea ' ), which roughly corresponds to the modern Pacific Ocean . Practically all deep-ocean crust present during the Triassic has been recycled through the subduction of oceanic plates, so very little is known about the open ocean from this time period. Most information on Panthalassan geology and marine life is derived from island arcs and rare seafloor sediments accreted onto surrounding land masses, such as present-day Japan and western North America. The eastern edge of Pangea

7268-453: The marine reptiles except ichthyosaurs and plesiosaurs . Invertebrates like brachiopods and molluscs (such as gastropods ) were severely affected. In the oceans, 22% of marine families and possibly about half of marine genera went missing. Though the end-Triassic extinction event was not equally devastating in all terrestrial ecosystems, several important clades of crurotarsans (large archosaurian reptiles previously grouped together as

7360-430: The next most common tetrapods, and early dinosaurs, passed through unchanged. However, both phytosaurs and aetosaurs were among the groups of archosaur reptiles completely wiped out by the end-Triassic extinction event. It seems likely then that there was some sort of end-Carnian extinction, when several herbivorous archosauromorph groups died out, while the large herbivorous therapsids —the kannemeyeriid dicynodonts and

7452-512: The north and Gondwana to the south. The global climate during the Triassic was mostly hot and dry, with deserts spanning much of Pangaea's interior. However, the climate shifted and became more humid as Pangaea began to drift apart. The end of the period was marked by yet another major mass extinction, the Triassic–Jurassic extinction event , that wiped out many groups, including most pseudosuchians, and allowed dinosaurs to assume dominance in

7544-402: The order Isoetales (which contains living quillworts ), rose to prominence due to the environmental instability following the Permian-Triassic extinction, with one particularly notable example being the genus Pleuromeia , which grew in columnar like fashion, sometimes reaching a height of 2 metres (6.6 ft). The relevance of lycophytes declined from the Middle Triassic onwards, following

7636-416: The return of more stable environmental conditions. While having first appeared during the Permian, the extinct seed plant group Bennettitales first became a prominent element in global floras during the Late Triassic, a position they would hold for much of the Mesozoic. In the Southern Hemisphere landmasses of Gondwana, the tree Dicroidium , an extinct " seed fern " belong to the order Corystospermales

7728-412: The surviving species. The Early Triassic Epoch saw the biotic recovery of life after the biggest mass extinction event of the past, which took millions of years due to the severity of the event and the harsh Early Triassic climate. Many types of corals , brachiopods , molluscs , echinoderms , and other invertebrates had disappeared. The Permian vegetation, which was dominated by Glossopteris in

7820-676: The terminus of the Triassic, there was an extreme warming event referred to as the End-Triassic Thermal Event (ETTE), which was responsible for the Triassic-Jurassic mass extinction. Bubbles of carbon dioxide in basaltic rocks dating back to the end of the Triassic indicate that volcanic activity from the Central Atlantic Magmatic Province helped trigger climate change in the ETTE. During the Early Triassic, lycophytes , particularly those of

7912-595: The terrestrial species had mostly died out during the extinction event. The Triassic survivors were aquatic or semi-aquatic, and were represented by Tupilakosaurus , Thabanchuia , Branchiosauridae and Micropholis , all of which died out in Early Triassic, and the successful Stereospondyli , with survivors into the Cretaceous Period. The largest Triassic stereospondyls, such as Mastodonsaurus , were up to 4 to 6 metres (13 to 20 ft) in length. Some lineages (e.g. trematosaurs ) flourished briefly in

8004-577: The time of the Permo-Triassic boundary; acid rain from the Siberian Traps eruptions or from an impact event that overwhelmed acidic swamps; climate shift to a greenhouse climate that was too hot and dry for peat accumulation; evolution of fungi or herbivores that were more destructive of wetlands; the extinction of all plants adapted to peat swamps, with a hiatus of several million years before new plant species evolved that were adapted to peat swamps; or soil anoxia as oxygen levels plummeted. Before

8096-615: Was a dominant element in forest habitats across the region during the Middle-Late Triassic. During the Late Triassic, the Ginkgoales (which today are represented by only a single species, Ginkgo biloba ) underwent considerable diversification. Conifers were abundant during the Triassic, and included the Voltziales (which contains various lineages, probably including those ancestral to modern conifers), as well as

8188-673: Was a relatively cool period known as the Early Norian Cool Interval (ENCI), after which occurred the Mid-Norian Warm Interval (MNWI) from 217 to 209 Ma. The MNWI was briefly interrupted around 214 Ma by a cooling possibly related to the Manicouagan impact . Around 212 Ma, a 10 Myr eccentricity maximum caused a paludification of Pangaea and a reduction in the size of arid climatic zones. The Rhaetian Cool Interval (RCI) lasted from 209 to 201 Ma. At

8280-505: Was accompanied by huge volcanic eruptions that occurred as the supercontinent Pangaea began to break apart about 202 to 191 million years ago (40Ar/39Ar dates), forming the Central Atlantic Magmatic Province (CAMP), one of the largest known inland volcanic events since the planet had first cooled and stabilized. Other possible but less likely causes for the extinction events include global cooling or even

8372-484: Was encroached upon by a pair of extensive oceanic basins: The Neo-Tethys (or simply Tethys) and Paleo-Tethys Oceans . These extended from China to Iberia, hosting abundant marine life along their shallow tropical peripheries. They were divided from each other by a long string of microcontinents known as the Cimmerian terranes . Cimmerian crust had detached from Gondwana in the early Permian and drifted northwards during

8464-409: Was temporally flat during the Early Triassic and may have allowed tropical species to extend their distribution poleward. This is evidenced by the global distribution of ammonoids . The extremely hot ocean temperatures facilitated extremely powerful hurricanes that frequently hit the coast of North China. The mostly hot climate of the Early Triassic may have been caused by late volcanic eruptions of

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