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101-584: By earliest Permian time, during the Wolfcampian Epoch , the ovoid shaped subsiding Delaware Basin extended over 10,000 square miles (26,000 km) in what is now western Texas and southeast New Mexico . This period of deposition left a thickness of 1,600 to 2,200 feet (490 to 670 m) of limestone interbedded with dark-colored shale . A narrow outlet that geologists call the Hovey Channel periodically supplied new seawater from

202-901: A century after its original naming, with the United States Geological Survey until 1941 considering the Permian a subsystem of the Carboniferous equivalent to the Mississippian and Pennsylvanian . The Permian Period is divided into three epochs , from oldest to youngest, the Cisuralian, Guadalupian, and Lopingian. Geologists divide the rocks of the Permian into a stratigraphic set of smaller units called stages , each formed during corresponding time intervals called ages. Stages can be defined globally or regionally. For global stratigraphic correlation,

303-630: A city in Perm Krai. The stage was introduced by Alexandr Antonovich Stukenberg in 1890. The Kungurian currently lacks a defined GSSP. Recent proposals have suggested the appearance of Neostreptognathodus pnevi as the lower boundary. The Guadalupian Series is named after the Guadalupe Mountains in Texas and New Mexico, where extensive marine sequences of this age are exposed. It was named by George Herbert Girty in 1902. The Roadian

404-630: A coal swamp community, has an upper canopy consisting of lycopsid tree Sigillaria , with a lower canopy consisting of Marattialean tree ferns, and Noeggerathiales. Early conifers appeared in the Late Carboniferous, represented by primitive walchian conifers, but were replaced with more derived voltzialeans during the Permian. Permian conifers were very similar morphologically to their modern counterparts, and were adapted to stressed dry or seasonally dry climatic conditions. The increasing aridity, especially at low latitudes, facilitated

505-541: A dramatic increase in diversification during the Early Permian. Towards the end of the Permian, there was a substantial drop in both origination and extinction rates. The dominant insects during the Permian Period were early representatives of Paleoptera , Polyneoptera , and Paraneoptera . Palaeodictyopteroidea , which had represented the dominant group of insects during the Carboniferous, declined during

606-661: A few percent of evaporite minerals, the remainder being composed of the more typical detrital clastic rocks and carbonates . Examples of evaporite formations include occurrences of evaporite sulfur in Eastern Europe and West Asia. For a formation to be recognised as evaporitic it may simply require recognition of halite pseudomorphs , sequences composed of some proportion of evaporite minerals, and recognition of mud crack textures or other textures . Evaporites are important economically because of their mineralogy, their physical properties in-situ, and their behaviour within

707-481: A peak of diversity in the Cisuralian, with a substantial decline during the Guadalupian-Lopingian following Olson's extinction, with the family diversity dropping below Carboniferous levels. Embolomeres , a group of aquatic crocodile-like limbed vertebrates that are reptilliomorphs under some phylogenies. They previously had their last records in the Cisuralian, are now known to have persisted into

808-437: A picture into past Earth climates. Some particular deposits even show important tectonic and climatic changes. These deposits also may contain important minerals that help in today's economy. Thick non-marine deposits that accumulate tend to form where evaporation rates will exceed the inflow rate, and where there is sufficient soluble supplies. The inflow also has to occur in a closed basin, or one with restricted outflow, so that

909-525: A protective cover, over plants such as ferns that disperse spores in a wetter environment. The first modern trees ( conifers , ginkgos and cycads ) appeared in the Permian. Three general areas are especially noted for their extensive Permian deposits—the Ural Mountains (where Perm itself is located), China, and the southwest of North America, including the Texas red beds. The Permian Basin in

1010-491: Is a chart that shows minerals that form the marine evaporite rocks. They are usually the most common minerals that appear in this kind of deposit. Evaporite minerals start to precipitate when their concentration in water reaches such a level that they can no longer exist as solutes . The minerals precipitate out of solution in the reverse order of their solubilities, such that the order of precipitation from sea water is: The abundance of rocks formed by seawater precipitation

1111-642: Is a global hiatus in the terrestrial fossil record during the late Kungurian and early Roadian , referred to as "Olson's Gap" that obscures the nature of the transition. Other proposals have suggested that the North American and Russian records overlap, with the latest terrestrial North American deposition occurring during the Roadian, suggesting that there was an extinction event, dubbed " Olson's Extinction ". The Middle Permian faunas of South Africa and Russia are dominated by therapsids, most abundantly by

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1212-663: Is defined by the first appearance of the conodont Jinogondolella aserrata. The Capitanian is named after the Capitan Reef in the Guadalupe Mountains of Texas, named by George Burr Richardson in 1904, and first used in a chronostratigraphic sense by Glenister and Furnish in 1961 as a substage of the Guadalupian Stage. The Capitanian was ratified as an international stage by the ICS in 2001. The GSSP for

1313-466: Is in the same order as the precipitation given above. Thus, limestone (dolomite are more common than gypsum , which is more common than halite, which is more common than potassium and magnesium salts. Evaporites can also be easily recrystallized in laboratories in order to investigate the conditions and characteristics of their formation. Recent evidence from satellite observations and laboratory experiments suggest evaporites are likely present on

1414-452: Is thought to have existed during the Early Permian. Though the fossil record is fragmentary, lungfish appear to have undergone an evolutionary diversification and size increase in freshwater habitats during the Early Permian, but subsequently declined during the middle and late Permian. Conodonts experienced their lowest diversity of their entire evolutionary history during the Permian. Permian chondrichthyan faunas are poorly known. Members of

1515-611: The Panthalassa Ocean to the west. The somewhat smaller and shallower Midland Basin was just east and the much smaller Marfa Basin was to the southwest. All three basins were south of the equator , north of the Ouachita Mountains of mid-Texas, and part of the northern continent Laurasia . Structurally the Delaware, Midland and Marfa were foreland basins created when the Ouachita Mountains were uplifted as

1616-700: The Salado Formation . The Delaware Basin was filled at least to the top of Capitan Reef and mostly covered by dry land before the end of the Ochoan Epoch. Rivers migrated over its surface and deposited the red silt and sand that now constitute the siltstone and sandstone of the Rustler and Dewey Lake Formations . A karst topography developed as groundwater circulated in the buried limestone formations, dissolving away caverns which were later destroyed by infill and erosion. Uplift associated with

1717-533: The Apache Mountains and Glass Mountains to the south. Streams eroded the softer sediment away, lowering the ground level to its current position. Acidic groundwater excavated caves in the limestone of the higher areas and eroded sediment helped fill any remaining Permian-aged caves. Unlike most other caves in limestone, in this case the acid was likely derived from hydrogen sulfide and sulfide -rich brines that were freed by tectonic activity in

1818-481: The Castile Formation . The Castile consists of 1 ⁄ 16 inch (1.6 mm) thick laminae of alternating gray anhydrite and gypsum , brown calcite , and halite . As the salt concentration increased, halite and potassium -rich salt precipitated from the briny body of water on its margin and on nearshore areas. This salt layer covered an increasingly large area as the water level dropped, forming

1919-722: The Great Salt Lake in Utah and the Dead Sea , which lies between Jordan and Israel. Evaporite depositional environments that meet the above conditions include: The most significant known evaporite depositions happened during the Messinian salinity crisis in the basin of the Mediterranean . Evaporite formations need not be composed entirely of halite salt. In fact, most evaporite formations do not contain more than

2020-599: The International Commission on Stratigraphy (ICS) ratify global stages based on a Global Boundary Stratotype Section and Point (GSSP) from a single formation (a stratotype ) identifying the lower boundary of the stage. The ages of the Permian, from youngest to oldest, are: For most of the 20th century, the Permian was divided into the Early and Late Permian, with the Kungurian being the last stage of

2121-554: The Laramide orogeny in the late Mesozoic and early Cenozoic created a major fault along which the Guadalupe Mountains were thrust into existence. The range forms the tilted upthrown part of the system and the Salt Flat Bolson forms the downfallen block. Capitan Reef limestone was exposed above the surface with the 1000-foot-high (300 m) El Capitan being its most prominent feature. Other large outcrops compose

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2222-580: The Leonardian Epoch at the start of the mid- Permian . Small banks along its margin developed along with small discontinuous patch reefs in the shallow water just offshore. The first formation that resulted was the Yeso and consists of alternating beds of dolomitic limestone, gypsum , and sandstone . The sediments responsible for creating the Yeso were deposited in nearshore areas that graded into

2323-726: The North China Craton , the South China Block and Indochina fused to each other and Pangea by the end of the Permian. The Zechstein Sea , a hypersaline epicontinental sea , existed in what is now northwestern Europe. Large continental landmass interiors experience climates with extreme variations of heat and cold (" continental climate ") and monsoon conditions with highly seasonal rainfall patterns. Deserts seem to have been widespread on Pangaea. Such dry conditions favored gymnosperms , plants with seeds enclosed in

2424-704: The Paleo-Tethys Ocean , a large ocean that existed between Asia and Gondwana. The Cimmeria continent rifted away from Gondwana and drifted north to Laurasia , causing the Paleo-Tethys Ocean to shrink. A new ocean was growing on its southern end, the Neotethys Ocean , an ocean that would dominate much of the Mesozoic Era. A magmatic arc, containing Hainan on its southwesternmost end, began to form as Panthalassa subducted under

2525-825: The Permian–Triassic extinction event (colloquially known as the Great Dying), the largest mass extinction in Earth's history (which is the last of the three or four crises that occurred in the Permian), in which nearly 81% of marine species and 70% of terrestrial species died out, associated with the eruption of the Siberian Traps . It took well into the Triassic for life to recover from this catastrophe; on land, ecosystems took 30 million years to recover. Prior to

2626-549: The Sydney Basin , and palaeoclimatic models of the Earth's climate based on the behaviour of modern weather patterns showing that such a megamonsoon would occur given the continental arrangement of the Permian. The aforementioned increasing equatorial aridity was likely driven by the development and intensification of this Pangaean megamonsoon. Permian marine deposits are rich in fossil mollusks , brachiopods , and echinoderms . Brachiopods were highly diverse during

2727-447: The U.S. states of Texas and New Mexico is so named because it has one of the thickest deposits of Permian rocks in the world. Sea levels dropped slightly during the earliest Permian (Asselian). The sea level was stable at several tens of metres above present during the Early Permian, but there was a sharp drop beginning during the Roadian, culminating in the lowest sea level of the entire Palaeozoic at around present sea level during

2828-399: The Ural Mountains in the years 1840 and 1841. Murchison identified "vast series of beds of marl , schist , limestone , sandstone and conglomerate" that succeeded Carboniferous strata in the region. Murchison, in collaboration with Russian geologists, named the period after the surrounding Russian region of Perm, which takes its name from the medieval kingdom of Permia that occupied

2929-618: The cherty dolomites of the San Andres Formation while deposition a little further out forms the quartz sandstone and scattered patch reefs of the Brushy Canyon Formation . Rapid subsidence of the basin started in the middle Guadalupian. Patch reefs responded by rapid (mostly vertical) growth, resulting in the Goat Seep Reef. Three facies developed: Subsidence of the basin stopped for good by

3030-577: The pseudosuchians , dinosaurs , and pterosaurs in the following Triassic, first appeared and diversified during the Late Permian, including the first appearance of the Archosauriformes during the latest Permian. Cynodonts , the group of therapsids ancestral to modern mammals , first appeared and gained a worldwide distribution during the Late Permian. Another group of therapsids, the therocephalians (such as Lycosuchus ), arose in

3131-566: The sauropsids ( reptiles ). The world at the time was dominated by the supercontinent Pangaea , which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa . The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their amphibian ancestors. Various authors recognise at least three, and possibly four extinction events in

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3232-605: The Asselian, Sakmarian, and Artinskian stages. The Kungurian was later added to conform to the Russian "Lower Permian". Albert Auguste Cochon de Lapparent in 1900 had proposed the "Uralian Series", but the subsequent inconsistent usage of this term meant that it was later abandoned. The Asselian was named by the Russian stratigrapher V.E. Ruzhenchev in 1954, after the Assel River in the southern Ural Mountains. The GSSP for

3333-691: The Cisuralian. Another cool period began around the middle Capitanian. This cool period, lasting for 3-4 Myr, was known as the Kamura Event. It was interrupted by the Emeishan Thermal Excursion in the late part of the Capitanian, around 260 million years ago, corresponding to the eruption of the Emeishan Traps . This interval of rapid climate change was responsible for the Capitanian mass extinction event. During

3434-450: The Cisuralian. Permian synapsids included some large members such as Dimetrodon . The special adaptations of synapsids enabled them to flourish in the drier climate of the Permian and they grew to dominate the vertebrates. A faunal turnover occurred around the transition between the Cisuralian and Guadalupian, with the decline of amphibians and the replacement of pelycosaurs (a paraphyletic group) with more advanced therapsids , although

3535-500: The Early Permian Chemnitz petrified forest of Germany demonstrates that they had complex branching patterns similar to modern angiosperm trees. By the Late Permian, high thin forests had become widespread across the globe, as evidenced by the global distribution of weigeltisaurids. The oldest likely record of Ginkgoales (the group containing Ginkgo and its close relatives) is Trichopitys heteromorpha from

3636-599: The Early Permian. Glenister and colleagues in 1992 proposed a tripartite scheme, advocating that the Roadian-Capitanian was distinct from the rest of the Late Permian, and should be regarded as a separate epoch. The tripartite split was adopted after a formal proposal by Glenister et al. (1999). Historically, most marine biostratigraphy of the Permian was based on ammonoids ; however, ammonoid localities are rare in Permian stratigraphic sections, and species characterise relatively long periods of time. All GSSPs for

3737-568: The Early Triassic, but a Permian origin is suspected. The diversity of coelacanths is relatively low throughout the Permian in comparison to other marine fishes, though there is an increase in diversity during the terminal Permian (Changhsingian), corresponding with the highest diversity in their evolutionary history during the Early Triassic. Diversity of freshwater fish faunas was generally low and dominated by lungfish and "Paleopterygians". The last common ancestor of all living lungfish

3838-611: The Guadalupe Mountains in the Pliocene and early Pleistocene epochs enlarged Carlsbad Cavern and nearby caves. Parts of the major caves emerged from the saturated phreatic zone into the vadose with temporary periods of repose during which additional solutional excavation occurred in the phreatic zone. These pauses in emergence are thought to be responsible for creating the different levels in Carlsbad Caverns. Precipitation of carbon dioxide -rich water that infiltrated into

3939-472: The Late Permian. Members of the modern orders Archostemata and Adephaga are known from the Late Permian. Complex wood boring traces found in the Late Permian of China suggest that members of Polyphaga , the most diverse group of modern beetles, were also present by the Late Permian. The terrestrial fossil record of the Permian is patchy and temporally discontinuous. Early Permian records are dominated by equatorial Europe and North America, while those of

4040-617: The Late Permian. By the Changhsingian, only a handful (4-6) genera remained. Corals exhibited a decline in diversity over the course of the Middle and Late Permian. Terrestrial life in the Permian included diverse plants, fungi , arthropods , and various types of tetrapods . The period saw a massive desert covering the interior of Pangaea . The warm zone spread in the northern hemisphere, where extensive dry desert appeared. The rocks formed at that time were stained red by iron oxides,

4141-623: The Lopingian in China. Modern amphibians ( lissamphibians ) are suggested to have originated during Permian, descending from a lineage of dissorophoid temnospondyls or lepospondyls . The diversity of fish during the Permian is relatively low compared to the following Triassic. The dominant group of bony fishes during the Permian were the " Paleopterygii " a paraphyletic grouping of Actinopterygii that lie outside of Neopterygii . The earliest unequivocal members of Neopterygii appear during

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4242-605: The Lopingian series. The GSSP for the base of the Wuchiapingian is located at Penglaitan, Guangxi , China and was ratified in 2004. The boundary is defined by the first appearance of Clarkina postbitteri postbitteri The Changhsingian was originally derived from the Changxing Limestone, a geological unit first named by the Grabau in 1923, ultimately deriving from Changxing County , Zhejiang .The GSSP for

4343-598: The Lopingian to a series, including all Permian deposits in South China that overlie the Maokou Limestone. In 1995, a vote by the Subcommission on Permian Stratigraphy of the ICS adopted the Lopingian as an international standard chronostratigraphic unit. The Wuchiapinginan and Changhsingian were first introduced in 1962, by J. Z. Sheng as the "Wuchiaping Formation" and "Changhsing Formation" within

4444-514: The Middle Permian. There were no flying vertebrates, though the extinct lizard-like reptile family Weigeltisauridae from the Late Permian had extendable wings like modern gliding lizards , and are the oldest known gliding vertebrates. Permian stem-amniotes consisted of lepospondyli and batrachosaurs , according to some phylogenies; according to others, stem-amniotes are represented only by diadectomorphs . Temnospondyls reached

4545-598: The Middle and Late Permian are dominated by temperate Karoo Supergroup sediments of South Africa and the Ural region of European Russia. Early Permian terrestrial faunas of North America and Europe were dominated by primitive pelycosaur synapsids including the herbivorous edaphosaurids , and carnivorous sphenacodontids , diadectids and amphibians . Early Permian reptiles, such as acleistorhinids , were mostly small insectivores. Synapsids (the group that would later include mammals) thrived and diversified greatly during

4646-514: The Ochoan, corresponding to the Lopingian. During the Permian, all the Earth 's major landmasses were collected into a single supercontinent known as Pangaea , with the microcontinental terranes of Cathaysia to the east. Pangaea straddled the equator and extended toward the poles, with a corresponding effect on ocean currents in the single great ocean (" Panthalassa ", the "universal sea"), and

4747-620: The Permian are based around the first appearance datum of specific species of conodont , an enigmatic group of jawless chordates with hard tooth-like oral elements. Conodonts are used as index fossils for most of the Palaeozoic and the Triassic. The Cisuralian Series is named after the strata exposed on the western slopes of the Ural Mountains in Russia and Kazakhstan. The name was proposed by J. B. Waterhouse in 1982 to comprise

4848-533: The Permian) reefs are mainly composed of corals . Sea level dropped as the late Permian glaciation intensified and locked increasing amounts of water in distant ice caps . Sedimentation continued to fill the Delaware Basin into the Ochoan Epoch of the upper Permian, periodically cutting the basin off from its source of seawater. Part of the resulting brine became the deep-water evaporites of

4949-582: The Permian, representing up to a third of all insects at some localities. Mecoptera (sometimes known as scorpionflies) first appeared during the Early Permian, going on to become diverse during the Late Permian. Some Permian mecopterans, like Mesopsychidae have long proboscis that suggest they may have pollinated gymnosperms. The earliest known beetles appeared at the beginning of the Permian. Early beetles such as members of Permocupedidae were likely xylophagous , feeding on decaying wood. Several lineages such as Schizophoridae expanded into aquatic habitats by

5050-540: The Permian-Triassic boundary, corresponding to the eruption of the Siberian Traps , which released more than 5 teratonnes of CO 2 , more than doubling the atmospheric carbon dioxide concentration. A -2% δ O excursion signifies the extreme magnitude of this climatic shift. This extremely rapid interval of greenhouse gas release caused the Permian-Triassic mass extinction, as well as ushering in an extreme hothouse that persisted for several million years into

5151-508: The Permian. Xenacanthiformes , another extinct group of shark-like chondrichthyans, were common in freshwater habitats, and represented the apex predators of freshwater ecosystems. Four floristic provinces in the Permian are recognised, the Angaran , Euramerican, Gondwanan, and Cathaysian realms. The Carboniferous Rainforest Collapse would result in the replacement of lycopsid -dominated forests with tree-fern dominated ones during

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5252-543: The Permian. The end of the Early Permian ( Cisuralian ) saw a major faunal turnover, with most lineages of primitive " pelycosaur " synapsids becoming extinct, being replaced by more advanced therapsids . The end of the Capitanian Stage of the Permian was marked by the major Capitanian mass extinction event , associated with the eruption of the Emeishan Traps . The Permian (along with the Paleozoic) ended with

5353-524: The Permian. The extinct order Productida was the predominant group of Permian brachiopods, accounting for up to about half of all Permian brachiopod genera. Brachiopods also served as important ecosystem engineers in Permian reef complexes. Amongst ammonoids , Goniatitida were a major group during the Early-Mid Permian, but declined during the Late Permian. Members of the order Prolecanitida were less diverse. The Ceratitida originated from

5454-515: The Permian. This is likely due to competition by Hemiptera , due to their similar mouthparts and therefore ecology. Primitive relatives of damselflies and dragonflies ( Meganisoptera ), which include the largest flying insects of all time, also declined during the Permian. Holometabola , the largest group of modern insects, also diversified during this time. " Grylloblattidans ", an extinct group of winged insects thought to be related to modern ice crawlers , reached their apex of diversity during

5555-654: The Usolka section in the southern Urals, which was ratified in 2018. The GSSP is defined by the first appearance of Sweetognathus binodosus . The Artinskian was named after the city of Arti in Sverdlovsk Oblast , Russia. It was named by Karpinsky in 1874. The Artinskian currently lacks a defined GSSP. The proposed definition for the base of the Artinskian is the first appearance of Sweetognathus aff. S. whitei. The Kungurian takes its name after Kungur ,

5656-401: The Word Formation by Johan August Udden in 1916, Glenister and Furnish in 1961 was the first publication to use it as a chronostratigraphic term as a substage of the Guadalupian Stage. The GSSP for the base of the Wordian is located in Guadalupe Pass, Texas, within the sediments of the Getaway Limestone Member of the Cherry Canyon Formation , which was ratified in 2001. The base of the Wordian

5757-413: The Wuchiapingian, followed by a slight rise during the Changhsingian. The Permian was cool in comparison to most other geologic time periods, with modest pole to Equator temperature gradients. At the start of the Permian, the Earth was still in the Late Paleozoic icehouse (LPIA), which began in the latest Devonian and spanned the entire Carboniferous period, with its most intense phase occurring during

5858-506: The amount of lava estimated to have been produced during this period, the worst-case scenario is the release of enough carbon dioxide from the eruptions to raise world temperatures five degrees Celsius. Another hypothesis involves ocean venting of hydrogen sulfide gas. Portions of the deep ocean will periodically lose all of its dissolved oxygen allowing bacteria that live without oxygen to flourish and produce hydrogen sulfide gas. If enough hydrogen sulfide accumulates in an anoxic zone ,

5959-411: The ancestors of many present-day families. Rich forests were present in many areas, with a diverse mix of plant groups. The southern continent saw extensive seed fern forests of the Glossopteris flora. Oxygen levels were probably high there. The ginkgos and cycads also appeared during this period. Insects, which had first appeared and become abundant during the preceding Carboniferous, experienced

6060-405: The base of the Asselian is located in the Aidaralash River valley near Aqtöbe , Kazakhstan, which was ratified in 1996. The beginning of the stage is defined by the first appearance of Streptognathodus postfusus . The Sakmarian is named in reference to the Sakmara River in the southern Urals, and was coined by Alexander Karpinsky in 1874. The GSSP for the base of the Sakmarian is located at

6161-418: The base of the Capitanian is located at Nipple Hill in the southeast Guadalupe Mountains of Texas, and was ratified in 2001, the beginning of the stage is defined by the first appearance of Jinogondolella postserrata. The Lopingian was first introduced by Amadeus William Grabau in 1923 as the "Loping Series" after Leping , Jiangxi , China. Originally used as a lithostraphic unit, T.K. Huang in 1932 raised

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6262-404: The base of the Changhsingian is located 88 cm above the base of the Changxing Limestone in the Meishan D section, Zhejiang, China and was ratified in 2005, the boundary is defined by the first appearance of Clarkina wangi. The GSSP for the base of the Triassic is located at the base of Bed 27c at the Meishan D section, and was ratified in 2001. The GSSP is defined by the first appearance of

6363-412: The beginning of the Triassic Period 251.902 Mya. It is the sixth and last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison , who named it after the region of Perm in Russia . The Permian witnessed the diversification of the two groups of amniotes , the synapsids and

6464-399: The carbonate banks of the Victorio Peak Formation in the deeper waters. Thin-bedded limestones of the Bone Spring Formation accumulated as limy ooze in the stagnant deepest part of the basin. Subsidence of the Delaware Basin restarted later in the mid Permian and by the Guadalupian Epoch of the upper Permian the patch reefs had grown larger. Sediments deposited close to the shore are now

6565-434: The cavern created speleothems , especially in the humid parts of the Pleistocene. Speleothems found in the "Big Room" of Carlsbad were dated using electron spin resonance dating and were found to be 500,000 to 600,000 years old. This indicates that the Big Room level was dry by that time. The soft and easily eroded gypsum of the Castile Formation was removed, exposing the Guadalupe Escarpment. Additional erosion intersected

6666-429: The chondrichthyan clade Holocephali , which contains living chimaeras , reached their apex of diversity during the Carboniferous-Permian, the most famous Permian representative being the "buzz-saw shark" Helicoprion , known for its unusual spiral shaped spiral tooth whorl in the lower jaw. Hybodonts , a group of shark-like chondrichthyans, were widespread and abundant members of marine and freshwater faunas throughout

6767-449: The climate became notably more arid at the end of the Carboniferous and beginning of the Permian. Nonetheless, temperatures continued to cool during most of the Asselian and Sakmarian, during which the LPIA peaked. By 287 million years ago, temperatures warmed and the South Pole ice cap retreated in what was known as the Artinskian Warming Event (AWE), though glaciers remained present in the uplands of eastern Australia, and perhaps also

6868-399: The conodont Hindeodus parvus . The Russian Tatarian Stage includes the Lopingian, Capitanian and part of the Wordian, while the underlying Kazanian includes the rest of the Wordian as well as the Roadian. In North America, the Permian is divided into the Wolfcampian (which includes the Nealian and the Lenoxian stages); the Leonardian (Hessian and Cathedralian stages); the Guadalupian; and

6969-434: The continental interior by the more advanced seed ferns and early conifers as a result of the Carboniferous rainforest collapse . At the close of the Permian, lycopod and equisete swamps reminiscent of Carboniferous flora survived only in Cathaysia , a series of equatorial islands in the Paleo-Tethys Ocean that later would become South China . The Permian saw the radiation of many important conifer groups, including

7070-458: The decline of early synapsid clades was apparently a slow event that lasted about 20 Ma, from the Sakmarian to the end of the Kungurian . Predator-prey interactions among terrestrial synapsids became more dynamic. If terrestrial deposition ended around the end of the Cisuralian in North America and began in Russia during the early Guadalupian, a continuous record of the transition is not preserved. Uncertain dating has led to suggestions that there

7171-412: The diverse Dinocephalia . Dinocephalians become extinct at the end of the Middle Permian, during the Capitanian mass extinction event . Late Permian faunas are dominated by advanced therapsids such as the predatory sabertoothed gorgonopsians and herbivorous beaked dicynodonts , alongside large herbivorous pareiasaur parareptiles . The Archosauromorpha , the group of reptiles that would give rise to

7272-465: The earliest Permian of France. The oldest known fossils definitively assignable to modern cycads are known from the Late Permian. In Cathaysia, where a wet tropical frost-free climate prevailed, the Noeggerathiales , an extinct group of tree fern-like progymnosperms were a common component of the flora The earliest Permian (~ 298 million years ago) Cathyasian Wuda Tuff flora, representing

7373-573: The early Wuchiapingian, following the emplacement of the Emeishan Traps, global temperatures declined as carbon dioxide was weathered out of the atmosphere by the large igneous province's emplaced basalts. The late Wuchiapingian saw the finale of the Late Palaeozoic Ice Age, when the last Australian glaciers melted. The end of the Permian is marked by a temperature excursion, much larger than the Emeishan Thermal Excursion, at

7474-431: The end of the Permian. Nautiloids , a subclass of cephalopods, surprisingly survived this occurrence. There is evidence that magma , in the form of flood basalt , poured onto the Earth's surface in what is now called the Siberian Traps , for thousands of years, contributing to the environmental stress that led to mass extinction. The reduced coastal habitat and highly increased aridity probably also contributed. Based on

7575-580: The family Daraelitidae within Prolecanitida during the mid-Permian, and extensively diversified during the Late Permian. Only three families of trilobite are known from the Permian, Proetidae , Brachymetopidae and Phillipsiidae . Diversity, origination and extinction rates during the Early Permian were low. Trilobites underwent a diversification during the Kungurian-Wordian, the last in their evolutionary history, before declining during

7676-429: The gas can rise into the atmosphere. Oxidizing gases in the atmosphere would destroy the toxic gas, but the hydrogen sulfide would soon consume all of the atmospheric gas available. Hydrogen sulfide levels might have increased dramatically over a few hundred years. Models of such an event indicate that the gas would destroy ozone in the upper atmosphere allowing ultraviolet radiation to kill off species that had survived

7777-721: The introduction of the term Permian , rocks of equivalent age in Germany had been named the Rotliegend and Zechstein , and in Great Britain as the New Red Sandstone . The term Permian was introduced into geology in 1841 by Sir Roderick Impey Murchison , president of the Geological Society of London , after extensive Russian explorations undertaken with Édouard de Verneuil in the vicinity of

7878-748: The late Carboniferous in Euramerica, and result in the differentiation of the Cathaysian floras from those of Euramerica. The Gondwanan floristic region was dominated by Glossopteridales , a group of woody gymnosperm plants, for most of the Permian, extending to high southern latitudes. The ecology of the most prominent glossopterid, Glossopteris , has been compared to that of bald cypress , living in mires with waterlogged soils. The tree-like calamites , distant relatives of modern horsetails , lived in coal swamps and grew in bamboo -like vertical thickets. A mostly complete specimen of Arthropitys from

7979-479: The later part of the Guadalupian. Capitan Reef was the largest in the basin, and it rapidly grew 350 miles (560 km) around it. The facies were: Capitan Reef was built primarily from calcareous sponges , encrusting algae such as stromatolites , and directly from seawater as a limey mud. In stark contrast, Cenozoic (current era), Mesozoic (age of the dinosaurs ), and even middle Paleozoic (well before

8080-664: The latter part of the Pennsylvanian epoch. A significant trend of increasing aridification can be observed over the course of the Cisuralian. Early Permian aridification was most notable in Pangaean localities at near-equatorial latitudes. Sea levels also rose notably in the Early Permian as the LPIA slowly waned. At the Carboniferous-Permian boundary, a warming event occurred. In addition to becoming warmer,

8181-461: The marine environments. Common minerals that are found in these deposits include blödite , borax , epsomite , gaylussite , glauberite , mirabilite , thenardite and trona . Non-marine deposits may also contain halite, gypsum, and anhydrite, and may in some cases even be dominated by these minerals, although they did not come from ocean deposits. This, however, does not make non-marine deposits any less important; these deposits often help to paint

8282-466: The mid- Tertiary and mixed with oxygenated groundwater, forming sulfuric acid . Carlsbad Caverns and nearby modern caves started to form at this time in the groundwater -saturated phreatic zone . Due to the semiarid climate , the karst topography that was created lacks the characteristic depressions, sink holes , pits, and solutional fissures on the surface. Mass wasting such as landslides further reduced topographic relief. Additional uplift of

8383-514: The most extensive extinction event recorded in paleontology : the Permian–Triassic extinction event . 90 to 95% of marine species became extinct , as well as 70% of all land organisms. It is also the only known mass extinction of insects. Recovery from the Permian–Triassic extinction event was protracted; on land, ecosystems took 30 million years to recover. Trilobites , which had thrived since Cambrian times, finally became extinct before

8484-473: The most potent greenhouse gases ) into the atmosphere to raise world temperatures an additional five degrees Celsius. The frozen methane hypothesis helps explain the increase in carbon-12 levels found midway in the Permian–Triassic boundary layer. It also helps explain why the first phase of the layer's extinctions was land-based, the second was marine-based (and starting right after the increase in C-12 levels), and

8585-414: The mountainous regions of far northern Siberia. Southern Africa also retained glaciers during the late Cisuralian in upland environments. The AWE also witnessed aridification of a particularly great magnitude. In the late Kungurian, cooling resumed, resulting in a cool glacial interval that lasted into the early Capitanian, though average temperatures were still much higher than during the beginning of

8686-713: The next geologic epoch, the Triassic. The Permian climate was also extremely seasonal and characterised by megamonsoons , which produced high aridity and extreme seasonality in Pangaea's interiors. Precipitation along the western margins of the Palaeo-Tethys Ocean was very high. Evidence for the megamonsoon includes the presence of megamonsoonal rainforests in the Qiangtang Basin of Tibet, enormous seasonal variation in sedimentation, bioturbation, and ichnofossil deposition recorded in sedimentary facies in

8787-591: The original water depth remains. At this point, minor carbonates begin to form. The next phase in the sequence comes when the experiment is left with about 20% of its original level. At this point, the mineral gypsum begins to form, which is then followed by halite at 10%, excluding carbonate minerals that tend not to be evaporites. The most common marine evaporites are calcite , gypsum and anhydrite , halite, sylvite , carnallite , langbeinite , polyhalite , and kainite . Kieserite (MgSO 4 ) may also be included, which often will make up less than four percent of

8888-458: The overall content. However, there are approximately 80 different minerals that have been reported found in evaporite deposits, though only about a dozen are common enough to be considered important rock formers. Non-marine evaporites are usually composed of minerals that are not common in marine environments because in general the water from which non-marine evaporite precipitates has proportions of chemical elements different from those found in

8989-414: The remaining water is enriched in salts, and they precipitate when the water becomes supersaturated. Marine evaporites tend to have thicker deposits and are usually the focus of more extensive research. When scientists evaporate ocean water in a laboratory, the minerals are deposited in a defined order that was first demonstrated by Usiglio in 1884. The first phase of precipitation begins when about 50% of

9090-439: The result of intense heating by the sun of a surface devoid of vegetation cover. A number of older types of plants and animals died out or became marginal elements. The Permian began with the Carboniferous flora still flourishing. About the middle of the Permian a major transition in vegetation began. The swamp -loving lycopod trees of the Carboniferous, such as Lepidodendron and Sigillaria , were progressively replaced in

9191-587: The same area hundreds of years prior, and which is now located in the Perm Krai administrative region. Between 1853 and 1867, Jules Marcou recognised Permian strata in a large area of North America from the Mississippi River to the Colorado River and proposed the name Dyassic , from Dyas and Trias , though Murchison rejected this in 1871. The Permian system was controversial for over

9292-489: The sediment has time to pool and form in a lake or other standing body of water. Primary examples of this are called "saline lake deposits". Saline lakes includes things such as perennial lakes, which are lakes that are there year-round, playa lakes, which are lakes that appear only during certain seasons, or any other terms that are used to define places that hold standing bodies of water intermittently or year-round. Examples of modern non-marine depositional environments include

9393-408: The southeastern South China. The Central Pangean Mountains , which began forming due to the collision of Laurasia and Gondwana during the Carboniferous, reached their maximum height during the early Permian around 295 million years ago, comparable to the present Himalayas , but became heavily eroded as the Permian progressed. The Kazakhstania block collided with Baltica during the Cisuralian, while

9494-559: The southern continent Gondwana collided with Laurasia, forming the supercontinent Pangea in the Late Carboniferous ( Pennsylvanian ). The Ouachita Mountains formed a rainshadow over the basins, and a warm shallow sea flooded the surrounding area. On the other side of the equator, the Ancestral Rocky Mountains formed a large mountainous island . The Delaware Basin temporarily stopped subsiding in

9595-532: The spread of conifers and their increasing prevalence throughout terrestrial ecosystems. Bennettitales , which would go on to become in widespread the Mesozoic, first appeared during the Cisuralian in China. Lyginopterids , which had declined in the late Pennsylvanian and subsequently have a patchy fossil record, survived into the Late Permian in Cathaysia and equatorial east Gondwana. The Permian ended with

9696-674: The subsurface. Evaporite minerals, especially nitrate minerals, are economically important in Peru and Chile. Nitrate minerals are often mined for use in the production on fertilizer and explosives . Thick halite deposits are expected to become an important location for the disposal of nuclear waste because of their geologic stability, predictable engineering and physical behaviour, and imperviousness to groundwater. Halite formations are famous for their ability to form diapirs , which produce ideal locations for trapping petroleum deposits. Halite deposits are often mined for use as salt . This

9797-399: The surface and in aquifers contain dissolved salts, the water must evaporate into the atmosphere for the minerals to precipitate. For this to happen, the water body must enter a restricted environment where water input into this environment remains below the net rate of evaporation. This is usually an arid environment with a small basin fed by a limited input of water. When evaporation occurs,

9898-551: The third land-based again. Evaporite An evaporite ( / ɪ ˈ v æ p ə ˌ r aɪ t / ) is a water- soluble sedimentary mineral deposit that results from concentration and crystallization by evaporation from an aqueous solution . There are two types of evaporite deposits: marine, which can also be described as ocean deposits, and non-marine, which are found in standing bodies of water such as lakes. Evaporites are considered sedimentary rocks and are formed by chemical sediments . Although all water bodies on

9999-413: The toxic gas. There are species that can metabolize hydrogen sulfide. Another hypothesis builds on the flood basalt eruption theory. An increase in temperature of five degrees Celsius would not be enough to explain the death of 95% of life. But such warming could slowly raise ocean temperatures until frozen methane reservoirs below the ocean floor near coastlines melted, expelling enough methane (among

10100-527: The upper part of Carlsbad Cavern and other caves, forming their entrances. Drying of cave air has reduced the growth rate of speleothems and encouraged the development of nodular travertine ("cave popcorn"). Permian The Permian ( / ˈ p ɜːr m i . ə n / PUR -mee-ən ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period 298.9 million years ago (Mya), to

10201-667: Was named in 1968 in reference to the Road Canyon Member of the Word Formation in Texas. The GSSP for the base of the Roadian is located 42.7m above the base of the Cutoff Formation in Stratotype Canyon, Guadalupe Mountains, Texas, and was ratified in 2001. The beginning of the stage is defined by the first appearance of Jinogondolella nankingensis . The Wordian was named in reference to

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