123-671: The Eichstätter Garten or Hortus Eystettensis ("Eichstätt Garden"), also known as the Bastionsgarten (Bastion garden) was a botanical garden that was created during the Renaissance period under Prince-Bishop Johann Konrad von Gemmingen on the bastions of the Willibaldsburg in Eichstätt , Upper Bavaria. The plants in the garden were described in the magnificent botanical work Hortus Eystettensis , which
246-491: A botanical garden is defined by its scientific or academic connection, then the first true botanical gardens were established with the revival of learning that occurred in the European Renaissance . These were secular gardens attached to universities and medical schools, used as resources for teaching and research. The superintendents of these gardens were often professors of botany with international reputations,
369-547: A case of convergent evolution ) in the cultivated tomato. The analysis predicts that var. cerasiforme appeared around 78,000 years ago, while the cultivated tomato originated around 7,000 years ago (5,000 BCE), with substantial uncertainty, making it unclear how humans may have been involved in the process. The Spanish first introduced tomatoes to Europe, where they became used in Spanish food. Elsewhere in Europe, its first use
492-602: A column surrounding the pistil 's style. The anthers bend into a cone-like structure, surrounding the stigma. The flowers are 1–2 cm (0.4–0.8 in) across, yellow, with five pointed lobes on the corolla ; they are borne in a cyme of three to twelve together. The fruit develops from the ovary of the plant after fertilization, its flesh comprising the pericarp walls. The fruit contains locules , hollow spaces full of seeds. These vary among cultivated varieties. Some smaller varieties have two locules; globe-shaped varieties typically have three to five; beefsteak tomatoes have
615-403: A department of an educational institution, it may be related to a teaching program. In any case, it exists for scientific ends and is not to be restricted or diverted by other demands. It is not merely a landscaped or ornamental garden, although it may be artistic, nor is it an experiment station or yet a park with labels on the plants. The essential element is the intention of the enterprise, which
738-528: A factor that probably contributed to the creation of botany as an independent discipline rather than a descriptive adjunct to medicine. The botanical gardens of Southern Europe were associated with university faculties of medicine and were founded in Italy at Orto botanico di Pisa (1544), Orto botanico di Padova (1545), Orto Botanico di Firenze (1545), Orto Botanico dell'Università di Pavia (1558) and Orto Botanico dell'Università di Bologna (1568). Here
861-587: A few from the Americas, but very few from Africa. The garden is best known for the florilegium commissioned by Gemmingen and prepared and published by Besler in Latin in 1613 as Hortus Eystettensis . Some paintings from the staircase appeared in this work. After the death of the Prince-Bishop in 1612, the garden was neglected. Parts of the garden were lost in favor of extensions to the bastion. The garden
984-456: A great number of small locules; and plum tomatoes have very few, very small locules. For propagation, the seeds need to come from a mature fruit, and must be lightly fermented to remove the gelatinous outer coating and then dried before use. The tomato has a mutualistic relationship with arbuscular mycorrhizal fungi such as Rhizophagus irregularis . Scientists use the tomato as a model species for investigating such symbioses. Like
1107-402: A lighter green, and reducing sugar in the resulting ripe fruit by 10–15%. Perhaps more importantly, the fruit chloroplasts are remodelled during ripening into chlorophyll-free chromoplasts that synthesize and accumulate the carotenoids lycopene , β-carotene , and other metabolites that are sensory and nutritional assets of the ripe fruit. The potent chloroplasts in the dark-green shoulders of
1230-978: A name for the fruit. The usual pronunciations of tomato are / t ə ˈ m eɪ t oʊ / (in North American English ) and / t ə ˈ m ɑː t oʊ / (in British English ). The British pronunciation was like the American until the Great Vowel Shift . The word's dual pronunciations were immortalized in Ira and George Gershwin 's 1937 song " Let's Call the Whole Thing Off " ("You like / p ə ˈ t eɪ t oʊ / and I like / p ə ˈ t ɑː t oʊ / / You like / t ə ˈ m eɪ t oʊ / and I like / t ə ˈ m ɑː t oʊ / "). The likely wild ancestor of
1353-538: A period of prosperity when the city was a trading centre for the Dutch East India Company . Other gardens were constructed in Brazil ( Rio de Janeiro Botanical Garden , 1808), Sri Lanka ( Botanic Gardens of Peradeniya , 1821 and on a site dating back to 1371), Indonesia ( Bogor Botanical Gardens , 1817 and Kebun Raya Cibodas , 1852), and Singapore ( Singapore Botanical Gardens , 1822). These had
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#17327905010321476-901: A process that has impaired the fruit's sweetness and flavor. There are thousands of cultivars, varying in size, color, shape, and flavor. Tomatoes are attacked by many insect pests and nematodes, and are subject to diseases caused by viruses and by mildew and blight fungi. The tomato has a strong savoury umami flavor, and is an important ingredient in cuisines around the world. It is used in pizzas , pasta sauces, soups such as gazpacho , curries including dhansak and rogan josh , as juice, and in Bloody Mary cocktails . Tomato festivals are held annually in Buñol , Spain, in Reynoldsburg, Ohio , and in Närpes , Finland. The word tomato comes from
1599-608: A profound effect on the economy of the countries, especially in relation to the foods and medicines introduced. The importation of rubber trees to the Singapore Botanic Garden initiated the important rubber industry of the Malay Peninsula . At this time also, teak and tea were introduced to India and breadfruit , pepper and starfruit to the Caribbean. Included in the charter of these gardens
1722-451: A round ebony table, the leaf and foot of which are inlaid with silver engraved flowers and insects ... the gardens are all going to be turned round as well and levelled with each other around the castle. The gardens were created from stones cut from a quarry below the castle. Gemmingen described the pre-existing garden as meinem wenig enges Gärtlein (my narrow little garden)) which he envisaged as becoming much more spacious and integrated. By
1845-412: A serrated margin; both the stem and leaves are densely glandular-hairy. Tomato flowers are bisexual and are able to self fertilize. As tomatoes were moved from their native areas, their traditional pollinators (probably a species of halictid bee ) did not move with them. The trait of self-fertility became an advantage, and domestic cultivars of tomato have been selected to maximize this trait. This
1968-455: A staple of the peasant population because they were not as filling as other crops. Additionally, both toxic and inedible varieties discouraged many people from attempting to consume or prepare any other varieties. In certain areas of Italy, such as Florence, the fruit was used solely as a tabletop decoration, until it was incorporated into the local cuisine in the late 17th or early 18th century. The earliest discovered cookbook with tomato recipes
2091-552: A tribute to Besler, and as in the Hortus , is organised by season. In the north-west is Spring, and moves from there through to winter. About half of the 1,048 plants that were illustrated in Besler's book are found in the modern garden. In 2013, an exhibition opened at the garden featuring the "exotic" plants of the historic garden to be found in the modern reconstruction. Botanical garden A botanical garden or botanic garden
2214-457: A wide influence on both botany and horticulture, as plants poured into it from around the world. The garden's golden age came in the 18th century, when it became the world's most richly stocked botanical garden. Its seed-exchange programme was established in 1682 and still continues today. With the increase in maritime trade , ever more plants were being brought back to Europe as trophies from distant lands, and these were triumphantly displayed in
2337-470: A year. Historically, botanical gardens exchanged plants through the publication of seed lists (these were called Latin : Indices Seminae in the 18th century). This was a means of transferring both plants and information between botanical gardens. This system continues today, although the possibility of genetic piracy and the transmission of invasive species has received greater attention in recent times. The International Association of Botanic Gardens
2460-462: Is tobacco mosaic virus . Handling cigarettes and other infected tobacco products can transmit the virus to tomato plants. A serious disease is curly top , carried by the beet leafhopper , which interrupts the lifecycle. As the name implies, it has the symptom of making the top leaves of the plant wrinkle up and grow abnormally. Bacterial wilt is another common disease impacting yield. Wang et al. , 2019 found phage combination therapies to reduce
2583-615: Is a garden with a documented collection of living plants for the purpose of scientific research, conservation, display, and education. It is their mandate as a botanical garden that plants are labelled with their botanical names . It may contain specialist plant collections such as cacti and other succulent plants , herb gardens , plants from particular parts of the world, and so on; there may be glasshouses or shadehouses , again with special collections such as tropical plants , alpine plants , or other exotic plants that are not native to that region. Most are at least partly open to
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#17327905010322706-590: Is a member of the nightshade family that includes tobacco , potato , and chili peppers . It originated from and was domesticated in western South America . It was introduced to the Old World by the Spanish in the Columbian exchange in the 16th century. Tomato plants are vines, largely annual and vulnerable to frost, though sometimes living longer in greenhouses. The flowers are able to self-fertilise. Modern varieties have been bred to ripen uniformly red, in
2829-488: Is closely linked to the history of botany itself. The botanical gardens of the 16th and 17th centuries were medicinal gardens, but the idea of a botanical garden changed to encompass displays of the beautiful, strange, new and sometimes economically important plant trophies being returned from the European colonies and other distant lands. Later, in the 18th century, they became more educational in function, demonstrating
2952-486: Is increasing, providing fruit during those times of the year when field-grown fruit is not readily available. Smaller fruit (cherry and grape), or cluster tomatoes (fruit-on-the-vine) are the fruit of choice for the large commercial greenhouse operators while the beefsteak varieties are the choice of owner-operator growers. Tomatoes are also grown using hydroponics . To facilitate transportation and storage, tomatoes are often picked unripe (green) and ripened in storage with
3075-523: Is inside the Parque La Carolina is a 165.5-acre (670,000 m ) park in the centre of the Quito central business district , bordered by the avenues Río Amazonas, de los Shyris, Naciones Unidas, Eloy Alfaro, and de la República. The botanical garden of Quito is a park, a botanical garden, an arboretum and greenhouses of 18,600 square meters that is planned to increase, maintain the plants of
3198-401: Is not the same as self-pollination , despite the common claim that tomatoes do so. That tomatoes pollinate themselves poorly without outside aid is clearly shown in greenhouse situations, where pollination must be aided by artificial wind, vibration of the plants, or by cultured bumblebees . The flowers develop on the apical meristem . They have the anthers fused along the edges, forming
3321-494: Is not used because it violates the International Code of Nomenclature barring the use of tautonyms in botanical nomenclature. The corrected name Lycopersicon lycopersicum (Nicolson 1974) was technically valid, because Miller's genus name and Linnaeus's species name differ in exact spelling. As Lycopersicon esculentum has become so well known, it was officially listed as a nomen conservandum in 1983, and would be
3444-669: Is respected worldwide for the published work of its scientists, the education of horticultural students, its public programmes, and the scientific underpinning of its horticulture. In 1728, John Bartram founded Bartram's Garden in Philadelphia , one of the continent's first botanical gardens. The garden is now managed as a historical site that includes a few original and many modern specimens as well as extensive archives and restored historical farm buildings. The large number of plants needing description were often listed in garden catalogues; and at this time Carl Linnaeus established
3567-1101: Is the acquisition and dissemination of botanical knowledge. A contemporary botanic garden is a strictly protected green area, where a managing organization creates landscaped gardens and holds documented collections of living plants and/or preserved plant accessions containing functional units of heredity of actual or potential value for purposes such as scientific research, education, public display, conservation, sustainable use, tourism and recreational activities, production of marketable plant-based products and services for improvement of human well-being. Worldwide, there are now about 1800 botanical gardens and arboreta in about 150 countries (mostly in temperate regions) of which about 550 are in Europe (150 of which are in Russia ), 200 in North America , and an increasing number in East Asia. These gardens attract about 300 million visitors
3690-546: Is to maintain documented collections of living plants for the purposes of scientific research, conservation, display, and education, although this will depend on the resources available and the special interests pursued at each particular garden. The staff will normally include botanists as well as gardeners. Many botanical gardens offer diploma/certificate programs in horticulture, botany and taxonomy. There are many internship opportunities offered to aspiring horticulturists. As well as opportunities for students/researchers to use
3813-670: The Botanical Garden of the University of Vienna and Hortus Botanicus Leiden . Many plants were being collected from the Near East , especially bulbous plants from Turkey . Clusius laid the foundations of Dutch tulip breeding and the bulb industry, and he helped create one of the earliest formal botanical gardens of Europe at Leyden where his detailed planting lists have made it possible to recreate this garden near its original site. The hortus medicus of Leyden in 1601
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3936-583: The Columbian exchange ; certainly the tomato was being grown in Europe within a few years of that event. The earliest discussion of the tomato in European literature appeared in Pietro Andrea Mattioli 's 1544 herbal . He suggested that a new type of eggplant had been brought to Italy. He stated that it was blood red or golden color when mature, and could be divided into segments and eaten like an eggplant—that is, cooked and seasoned with salt, black pepper, and oil. Ten years later Mattioli named
4059-885: The Global Strategy for Plant Conservation by producing a range resources and publications, and by organizing international conferences and conservation programs. Communication also happens regionally. In the United States, there is the American Public Gardens Association (formerly the American Association of Botanic Gardens and Arboreta), and in Australasia there is the Botanic Gardens of Australia and New Zealand (BGANZ). The history of botanical gardens
4182-842: The Government Hill in Victoria City , Hong Kong Island . The Koishikawa Botanical Garden in Tokyo, with its origin going back to the Tokugawa shogunate 's ownership, became in 1877 part of the Tokyo Imperial University . In Sri Lanka major botanical gardens include the Royal Botanic Gardens, Peradeniya (formally established in 1843), Hakgala Botanical Gardens (1861) and Henarathgoda Botanical Garden (1876). Jardín Botánico de Quito
4305-1001: The Royal Tasmanian Botanical Gardens , 1818; the Royal Botanic Gardens, Melbourne , 1845; Adelaide Botanic Gardens , 1854; and Brisbane Botanic Gardens , 1855. These were established essentially as colonial gardens of economic botany and acclimatisation. The Auburn Botanical Gardens , 1977, located in Sydney's western suburbs , are one of the popular and diverse botanical gardens in the Greater Western Sydney area. Major botanical gardens in New Zealand include Dunedin Botanic Gardens , 1863; Christchurch Botanic Gardens , 1863; Ōtari-Wilton's Bush , 1926; and Wellington Botanic Gardens , 1868. Hong Kong Botanic Gardens , 1871 (renamed Hong Kong Zoological and Botanical Gardens in 1975), up from
4428-595: The Schmiede-Bastion , opened in 1998 and testifies to the former splendor of the Prince-Bishop's garden. It is unclear where the historic garden actually was. It serves as an information resource for both the historic garden and florilegium. The reconstruction was based on the account by the French botanist Gérard G. Aymonin in the 1988 facsimile edition of the Hortus Eystettensis . The project
4551-654: The potato , tomatoes belong to the genus Solanum , which is a member of the nightshade family, the Solanaceae . That is a diverse family of flowering plants, often poisonous, that includes the mandrake ( Mandragora ), deadly nightshade ( Atropa ), and tobacco ( Nicotiana ), as shown in the outline phylogenetic tree (many branches omitted). many garden flowers and other species Nicotiana (tobacco) Atropa (nightshades) Mandragora (mandrake) (sweet and bell peppers) S. lycopersicum (tomato) S. tuberosum (potato) In 1753, Linnaeus placed
4674-412: The tomato bug , stink bugs , cutworms , tomato hornworms and tobacco hornworms , aphids , cabbage loopers , whiteflies , tomato fruitworms , flea beetles , red spider mite , slugs , and Colorado potato beetles . The tomato russet mite , Aculops lycopersici , feeds on foliage and young fruit of tomato plants, causing shrivelling and necrosis of leaves, flowers, and fruit, possibly killing
4797-453: The "Father of Botany". There is some debate among science historians whether this garden was ordered and scientific enough to be considered "botanical", and suggest it more appropriate to attribute the earliest known botanical garden in Europe to the botanist and pharmacologist Antonius Castor , mentioned by Pliny the Elder in the 1st century. Though these ancient gardens shared some of
4920-697: The "U" phenotype are beneficial here, but have the disadvantage of leaving green shoulders near the stems of the ripe fruit, and even cracked yellow shoulders. This is apparently because of oxidative stress due to overload of the photosynthetic chain in direct sunlight at high temperatures. Hence, genetic design of a commercial variety that combines the advantages of types "u" and "U" requires fine tuning, but may be feasible. Breeders strive to produce tomato plants with improved yield, shelf life, size, and resistance to environmental pressures, including disease. These efforts have yielded unintended negative consequences on various fruit attributes. For instance, linkage drag ,
5043-552: The "botanical staircase" because it was fitted with wooden panels painted with flowers. It was constructed before Elias Holl 's new building, the Gemmingenbau. Gemmingen has planned for his garden to be the first to include all the plants known at the time, including many imported from distant lands. One of the reasons that the garden became famous was the variety of what were then considered exotic plants, including potatoes, sunflowers and tomatoes , tulips and figs. A number of
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5166-523: The 1540s. It was probably eaten shortly after it was introduced, and was certainly being used as food by the early 17th century in Spain, as documented in the 1618 play La octava maravilla by Lope de Vega with "lovelier than ... a tomato in season". The tomato was introduced to China, likely via the Philippines or Macau, in the 16th century. It was given the name 番茄 fānqié (foreign eggplant), as
5289-408: The 16th and 17th centuries, the first plants were being imported to these major Western European gardens from Eastern Europe and nearby Asia (which provided many bulbs ), and these found a place in the new gardens, where they could be conveniently studied by the plant experts of the day. For example, Asian introductions were described by Carolus Clusius (1526–1609), who was director, in turn, of
5412-685: The British and Dutch, in India , South-east Asia and the Caribbean . This was also the time of Sir Joseph Banks 's botanical collections during Captain James Cook 's circumnavigations of the planet and his explorations of Oceania , which formed the last phase of plant introduction on a grand scale. There are currently about 230 tropical botanical gardens with a concentration in southern and south-eastern Asia. The first botanical garden founded in
5535-618: The Chelsea Physic Garden to the Province of Georgia in 1732 and tea into India by Calcutta Botanic Garden. The transfer of germplasm between the temperate and tropical botanical gardens was undoubtedly responsible for the range of agricultural crops currently used in several regions of the tropics. The first botanical gardens in Australia were founded early in the 19th century. The Royal Botanic Gardens, Sydney , 1816;
5658-619: The Chinese named many foodstuffs introduced from abroad, but referring specifically to early introductions. In 1548, when the house steward of Cosimo de' Medici , the grand duke of Tuscany , wrote to the Medici private secretary informing him that the basket of tomatoes sent from the grand duke's Florentine estate at Torre del Gallo "had arrived safely". Tomatoes were grown mainly as ornamentals early on after their arrival in Italy. For example,
5781-521: The Christian conquest in 1085 CE. Ibn Bassal then founded a garden in Seville, most of its plants being collected on a botanical expedition that included Morocco, Persia, Sicily, and Egypt. The medical school of Montpelier was also founded by Spanish Arab physicians, and by 1250 CE, it included a physic garden, but the site was not given botanic garden status until 1593. Botanical gardens, in
5904-499: The Florentine aristocrat Giovanvettorio Soderini wrote how they "were to be sought only for their beauty", and were grown only in gardens or flower beds. The tomato's ability to mutate and create new and different varieties helped contribute to its success and spread throughout Italy. However, in areas where the climate supported growing tomatoes, their habit of growing close to the ground suggested low status. They were not adopted as
6027-1023: The Netherlands ( Hortus Botanicus Leiden , 1590; Hortus Botanicus (Amsterdam) , 1638), Germany ( Alter Botanischer Garten Tübingen , 1535; Leipzig Botanical Garden , 1580; Botanischer Garten Jena , 1586; Botanischer Garten Heidelberg , 1593; Herrenhäuser Gärten, Hanover , 1666; Botanischer Garten der Christian-Albrechts-Universität zu Kiel , 1669; Botanical Garden in Berlin , 1672), Switzerland ( Old Botanical Garden, Zürich , 1560; Basel , 1589); England ( University of Oxford Botanic Garden , 1621; Chelsea Physic Garden , 1673); Scotland ( Royal Botanic Garden Edinburgh , 1670); and in France ( Jardin des plantes de Montpellier , 1593; Faculty of Medicine Garden, Paris, 1597; Jardin des Plantes , Paris, 1635), Denmark ( University of Copenhagen Botanical Garden , 1600); Sweden ( Uppsala University , 1655). During
6150-743: The Royal Botanic Gardens, Kew (1759) and Orotava Acclimatization Garden (in Spanish) , Tenerife (1788) and the Real Jardín Botánico de Madrid (1755) were set up to cultivate new species returned from expeditions to the tropics; they also helped found new tropical botanical gardens. From the 1770s, following the example of the French and Spanish , amateur collectors were supplemented by official horticultural and botanical plant hunters. These botanical gardens were boosted by
6273-534: The Royal Garden set aside as a physic garden. William Aiton (1741–1793), the first curator, was taught by garden chronicler Philip Miller of the Chelsea Physic Garden whose son Charles became first curator of the original Cambridge Botanic Garden (1762). In 1759, the "Physick Garden" was planted, and by 1767, it was claimed that "the Exotick Garden is by far the richest in Europe". Gardens such as
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#17327905010326396-523: The Spanish tomate , which in turn comes from the Nahuatl word tomatl [ˈtomat͡ɬ] pronunciation . The specific name lycopersicum , meaning "wolf peach", originated with Galen , who used it to denote a plant that has never been identified. Luigi Anguillara speculated in the 16th century that Galen's lycopersicum might be the tomato, and despite the impossibility of this identification, lycopersicum entered scientific use as
6519-575: The Spanish invaders, not only with their appearance, but also because the indigenous Aztecs employed many more medicinal plants than did the classical world of Europe. Early medieval gardens in Islamic Spain resembled botanic gardens of the future, an example being the 11th-century Huerta del Rey garden of physician and author Ibn Wafid (999–1075 CE) in Toledo . This was later taken over by garden chronicler Ibn Bassal (fl. 1085 CE) until
6642-486: The Vatican grounds in 1447, for a garden of medicinal plants that were used to promote the teaching of botany, and this was a forerunner to the University gardens at Padua and Pisa established in the 1540s. Certainly the founding of many early botanic gardens was instigated by members of the medical profession. In the 17th century, botanical gardens began their contribution to a deeper scientific curiosity about plants. If
6765-452: The biochemistry (many of the alkaloids common to other Solanum species are conspicuously absent from the tomato). On the other hand, hybrids of tomato and diploid potato can be created in the lab by somatic fusion , and are partially fertile, providing evidence of the close relationship between these species. An international consortium of researchers from 10 countries began sequencing the tomato genome in 2004. A prerelease version of
6888-415: The brightest red to the deepest yellow. Sahagún mentioned Aztecs cooking various sauces, some with tomatoes of different sizes, serving them in city markets: "foods sauces, hot sauces; ... with tomatoes, ... sauce of large tomatoes, sauce of ordinary tomatoes, ..." The Spanish conquistador Hernán Cortés 's capture of Tenochtitlan in 1521 initiated the widespread cultural and biological interchange called
7011-549: The castle on Mondscheinweg has been preserved as a stable house, the renovation of which was completed by the Bavarian State in 2007. A fountain bowl from the 17th century has been preserved from the stone garden decoration of the Prince-Bishop's garden. The modern Bastionsgarten (Bastion Garden), newly built by the Bavarian Palace Administration on the northern bastion of the outer bailey,
7134-451: The castle, and had made all the preparations, borders, steps, pathways and turns this necessitated". He filled in the moat, which was spanned by a bridge between the forecastle and the outerworks, with vault constructions. He also created a level area for a new building. At his funeral, the oration recognised him for having "built the new gardens, summerhouses and summer palaces so splendidly", while his obituary, by Philipp Menzel , referred to
7257-481: The characteristics of present-day botanical gardens, the forerunners of modern botanical gardens are generally regarded as being the medieval monastic physic gardens that originated after the decline of the Roman Empire at the time of Emperor Charlemagne (742–789 CE). These contained a hortus , a garden used mostly for vegetables, and another section set aside for specially labelled medicinal plants and this
7380-442: The classification systems being developed by botanists in the gardens' museums and herbaria. Botanical gardens had now become scientific collections, as botanists published their descriptions of the new exotic plants, and these were also recorded for posterity in detail by superb botanical illustrations. In this century, botanical gardens effectively dropped their medicinal function in favour of scientific and aesthetic priorities, and
7503-405: The collection for their studies. The origin of modern botanical gardens is generally traced to the appointment of botany professors to the medical faculties of universities in 16th-century Renaissance Italy, which also entailed curating a medicinal garden . However, the objectives, content, and audience of today's botanic gardens more closely resembles that of the grandiose gardens of antiquity and
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#17327905010327626-490: The complex with the precious and exotic plants was also a representation of his princely power. Philipp Hainhofer provided a detailed description of the garden on his visit there in 1611. He mentions how he; ... went into eight gardens around the castle, which is situated on rock ... all of which are arranged differently with flowerbeds, flowers, especially beautiful roses, lilies, tulips ... some of which are embellished with painted rooms and summerhouses, including in one room
7749-475: The correct name for the tomato in classifications which do not place the tomato in the genus Solanum . Genetic evidence shows that Linnaeus was correct to put the tomato in the genus Solanum , making S. lycopersicum the correct name. Both names, however, will probably be found in the literature for some time. Two of the major reasons for considering the genera separate are the leaf structure (tomato leaves are markedly different from any other Solanum ), and
7872-414: The country (Ecuador is among the 17 richest countries in the world in the native species, a study on this matter). The Ecuadorian flora classified, determines the existence of 17,000 species) Tomato The tomato ( US : / t ə m eɪ t oʊ / , UK : / t ə m ɑː t oʊ / ), Solanum lycopersicum , is a plant whose fruit is an edible berry that is eaten as a vegetable. The tomato
7995-440: The culinary use of the tomato included Michele Felice Cornè and Robert Gibbon Johnson . Many Americans considered tomatoes to be poisonous at this time and, in general, they were grown more as ornamental plants than as food. In 1897, W. H. Garrison stated, "The belief was once transmitted that the tomato was sinisterly dangerous." He recalled in his youth tomatoes were dubbed "love-apples or wolf-apples" and shunned as "globes of
8118-502: The devil". When Alexander W. Livingston (1821–1898) began developing the tomato as a commercial crop, his aim had been to grow tomatoes smooth in contour, uniform in size, and sweet in flavor. He eventually developed over seventeen varieties. The U.S. Department of Agriculture's 1937 yearbook declared that "half of the major varieties were a result of the abilities of the Livingstons to evaluate and perpetuate superior material in
8241-525: The direct ancestor of some modern cultivated tomatoes. The Aztecs raised several varieties of tomato, with red tomatoes called xitomatl and green tomatoes (physalis) called tomatl ( tomatillo ). Bernardino de Sahagún reported seeing a great variety of tomatoes in the Aztec market at Tenochtitlán (Mexico City): "large tomatoes, small tomatoes, leaf tomatoes, sweet tomatoes, large serpent tomatoes, nipple-shaped tomatoes", and tomatoes of all colors from
8364-431: The early blight pathogen Alternaria solani . However, this tactic has limitations, since selection for traits such as pathogen resistance can negatively impact other favorable traits such as fruit production. The tomato is grown worldwide for its edible fruits , with thousands of cultivars . Greenhouse tomato production in large-acreage commercial greenhouses and owner-operator stand-alone or multiple-bay greenhouses
8487-481: The educational garden of Theophrastus in the Lyceum of ancient Athens. The early concern with medicinal plants changed in the 17th century to an interest in the new plant imports from explorations outside Europe as botany gradually established its independence from medicine. In the 18th century, systems of nomenclature and classification were devised by botanists working in the herbaria and universities associated with
8610-544: The end of the 18th century, Kew, under the directorship of Sir Joseph Banks , enjoyed a golden age of plant hunting, sending out collectors to the South African Cape , Australia , Chile , China , Ceylon , Brazil , and elsewhere, and acting as "the great botanical exchange house of the British Empire ". From its earliest days to the present, Kew has in many ways exemplified botanic garden ideals, and
8733-456: The flora being sent back to Europe from various European colonies around the globe . At this time, British horticulturalists were importing many woody plants from Britain's colonies in North America , and the popularity of horticulture had increased enormously, encouraged by the horticultural and botanical collecting expeditions overseas fostered by the directorship of Sir William Jackson Hooker and his keen interest in economic botany . At
8856-625: The fresh market and for canning and processing. The University of California, Davis 's C.M. Rick Tomato Genetics Resource Center maintains a gene bank of wild relatives, monogenic mutants and genetic stocks. Research on processing tomatoes is also conducted by the California Tomato Research Institute in Escalon, California . In California, growers have used a method of cultivation called dry-farming , especially with Early Girl tomatoes. This technique encourages
8979-590: The fruits in print as pomi d'oro , or "golden apples". After the Spanish colonization of the Americas , the Spanish distributed the tomato throughout their colonies in the Caribbean . They brought it to the Philippines , from where it spread to southeast Asia and then the whole of Asia. The Spanish brought the tomato to Europe, where it grew easily in Mediterranean climates ; cultivation began in
9102-434: The gardens, these systems often being displayed in the gardens as educational "order beds ". With the rapid expansion of European colonies around the globe in the late 18th century, botanic gardens were established in the tropics, and economic botany became a focus with the hub at the Royal Botanic Gardens, Kew , near London. Over the years, botanical gardens, as cultural and scientific organisations, have responded to
9225-595: The genome was made available in December 2009. The complete genome for the cultivar Heinz 1706 was published on 31 May 2012 in Nature . The latest reference genome published in 2021 had 799 MB and encodes 34,384 (predicted) proteins, spread over 12 chromosomes. The first commercially available genetically modified food was a tomato called Flavr Savr , which was engineered to have a longer shelf life. It could be vine ripened without compromising shelf life . However,
9348-488: The interests of botany and horticulture . Nowadays, most botanical gardens display a mix of the themes mentioned and more; having a strong connection with the general public, there is the opportunity to provide visitors with information relating to the environmental issues being faced at the start of the 21st century, especially those relating to plant conservation and sustainability . The "New Royal Horticultural Society Dictionary of Gardening" (1999) points out that among
9471-551: The introduction of an undesired trait during backcrossing , has altered the metabolism of the fruit. This trait is physically close to the desired allele along the chromosome. Breeding for traits like larger fruit has thus unintentionally altered nutritional value and flavor. Breeders have turned to wild tomato species as a source of alleles to introduce beneficial traits into modern varieties. For example, wild relatives may possess higher amounts of fruit solids (associated with greater sugar content), or resistance to diseases such as to
9594-585: The irrigation schemes and fragrance of exotic flowers. Prince-Bishop Johann Konrad von Gemmingen (1595–1612) immediately commenced plans to expand the existing structures and commissioned Joachim Camerarius the Younger (1534–1598), a physician and botanist, to supervise the project, which was implemented from 1597. After Camerarius died the following year, Gemmingen asked the Nuremberg pharmacist Basilius Besler (1561–1629) to take his place. Besler noted that
9717-599: The latest plant classification systems devised by botanists working in the associated herbaria as they tried to order these new treasures. Then, in the 19th and 20th centuries, the trend was towards a combination of specialist and eclectic collections demonstrating many aspects of both horticulture and botany. The idea of "scientific" gardens used specifically for the study of plants dates back to antiquity. Near-eastern royal gardens set aside for economic use or display and containing at least some plants gained by special collecting trips or military campaigns abroad, are known from
9840-508: The modern sense, developed from physic gardens , whose main purpose was to cultivate herbs for medical use as well as research and experimentation. Such gardens have a long history. In Europe, for example, Aristotle (384 BCE – 322 BCE) is said to have had a physic garden in the Lyceum at Athens, which was used for educational purposes and for the study of botany, and this was inherited, or possibly set up, by his pupil Theophrastus ,
9963-642: The name 'Love Apple' as a French misreading of the Italian pomo dei Mori ("the Moors' apple") as pomme d'amour , ("apple of love"). The tomato was introduced to cultivation in the Middle East by John Barker , British consul in Aleppo c. 1799 to 1825 . Nineteenth century descriptions of its consumption are uniformly as an ingredient in a cooked dish. In 1881, it is described as only eaten in
10086-724: The physicians (referred to in English as apothecaries ) delivered lectures on the Mediterranean "simples" or " officinals " that were being cultivated in the grounds. Student education was no doubt stimulated by the relatively recent advent of printing and the publication of the first herbals. All of these botanical gardens still exist, mostly in their original locations. The tradition of these Italian gardens passed into Spain Botanical Garden of Valencia , 1567) and Northern Europe , where similar gardens were established in
10209-403: The plant hormone ethylene . At industrial scale, such as for canning, tomatoes are picked mechanically. The machine cuts the whole vine and uses sensors to separate ripe tomatoes from the rest of the plant, which is returned to the farm for use either as green manure or to be grazed by livestock . In 2022, world production of tomatoes was 186 million tonnes , with China accounting for 37% of
10332-422: The plant to send roots deep to find existing moisture. Tomato plants are vines, becoming decumbent , and can grow up to 3 m (9.8 ft); bush varieties are generally no more than 100 cm (3 ft 3 in) tall. They are tender perennials, often grown as annuals. Tomato plants are dicots . They grow as a series of branching stems, with a terminal bud at the tip that does the actual growing. When
10455-430: The plant. After an insect attack tomato plants produce systemin , a plant peptide hormone . This activates defensive mechanisms, such as the production of protease inhibitors to slow the growth of insects. The hormone was first identified in tomatoes. Tomato cultivars vary widely in their resistance to disease. Modern hybrids focus on improving disease resistance over the heirloom plants . A common tomato disease
10578-534: The plants found their way from the Americas by way of Spanish and Portuguese sailors. Apart from Camerarius, other plants came from the collection of Carolus Clusius (1526–1609) in Leiden and from traders in the Netherlands. In Gemmingen's time, about half of the plants he introduced had become naturalised within Germany. A third came from the Mediterranean, about ten per cent from the Middle East, India and Asia, and
10701-447: The private estates of the wealthy, in commercial nurseries , and in the public botanical gardens. Heated conservatories called " orangeries ", such as the one at Kew, became a feature of many botanical gardens. Industrial expansion in Europe and North America resulted in new building skills, so plants sensitive to cold were kept over winter in progressively elaborate and expensive heated conservatories and glasshouses. The 18th century
10824-408: The product was not commercially successful, and was sold only until 1997. The poor taste and lack of sugar in modern garden and commercial tomato varieties resulted from breeding tomatoes to ripen uniformly red. This change occurred after discovery of a mutant "u" phenotype in the mid-20th century, so named because the fruits ripened uniformly. This was widely cross-bred to produce red fruit without
10947-557: The public for the purposes of recreation, education and research." The term tends to be used somewhat differently in different parts of the world. For example a large woodland garden with a good collection of rhododendron and other flowering tree and shrub species is very likely to present itself as a "botanical garden" if it is located in the US, but very unlikely to do so if in the UK (unless it also contains other relevant features). Very few of
11070-454: The public, and may offer guided tours, public programming such as workshops, courses, educational displays, art exhibitions , book rooms, open-air theatrical and musical performances, and other entertainment. Botanical gardens are often run by universities or other scientific research organizations, and often have associated herbaria and research programmes in plant taxonomy or some other aspect of botanical science. In principle, their role
11193-824: The purposes of scientific research, conservation, display and education." The following definition was produced by staff of the Liberty Hyde Bailey Hortorium of Cornell University in 1976. It covers in some detail the many functions and activities generally associated with botanical gardens: A botanical garden is a controlled and staffed institution for the maintenance of a living collection of plants under scientific management for purposes of education and research, together with such libraries, herbaria, laboratories, and museums as are essential to its particular undertakings. Each botanical garden naturally develops its own special fields of interests depending on its personnel, location, extent, available funds, and
11316-665: The region "within the last forty years". The earliest reference to tomatoes being grown in British North America is from 1710, when herbalist William Salmon saw them in what is today South Carolina , perhaps introduced from the Caribbean. By the mid-18th century, they were cultivated on some Carolina plantations, and probably in other parts of the Southeast. Thomas Jefferson , who ate tomatoes in Paris, sent some seeds back to America. Some early American advocates of
11439-544: The rubber plant was introduced to Singapore. Especially in the tropics, the larger gardens were frequently associated with a herbarium and museum of economy. The Botanical Garden of Peradeniya had considerable influence on the development of agriculture in Ceylon where the Para rubber tree ( Hevea brasiliensis ) was introduced from Kew, which had itself imported the plant from South America . Other examples include cotton from
11562-689: The second millennium BCE in ancient Egypt , Mesopotamia , Crete , Mexico and China . In about 2800 BCE, the Chinese Emperor Shen Nung sent collectors to distant regions searching for plants with economic or medicinal value. It has also been suggested that the Spanish colonization of Mesoamerica influenced the history of the botanical garden as gardens in Tenochtitlan established by king Nezahualcoyotl , also gardens in Chalco (altépetl) and elsewhere, greatly impressed
11685-428: The sites used for the UK's dispersed National Plant Collection , usually holding large collections of a particular taxonomic group, would call themselves "botanic gardens". This has been further reduced by Botanic Gardens Conservation International to the following definition which "encompasses the spirit of a true botanic garden": "A botanic garden is an institution holding documented collections of living plants for
11808-467: The soil surrounding the castle had been much improved with earth from the valley below. Records of the expenses relating to the garden and the round tower pavilion, dating back to 1599, survive. The Hortus Eystettensis consisted of eight garden areas around the residence on the Willibaldsberg. The Prince-Bishop's love of nature was probably not the only decisive factor in laying out the garden,
11931-400: The sound of running water could be heard. This led to a complex irrigation system of pipes, pillars and a brook bringing water to the entire garden. Beneath the belvedere were two stories of treasure vaults, accessed from the Prince-Bishop's ante-chamber via a spiral staircase from the belvedere which continued down to the hare pit and over a bridge to the outer ward. This staircase was called
12054-404: The system of binomial nomenclature which greatly facilitated the listing process. Names of plants were authenticated by dried plant specimens mounted on card (a hortus siccus or garden of dried plants) that were stored in buildings called herbaria , these taxonomic research institutions being frequently associated with the botanical gardens, many of which by then had "order beds" to display
12177-417: The term "botanic garden" came to be more closely associated with the herbarium, library (and later laboratories) housed there than with the living collections – on which little research was undertaken. The late 18th and early 19th centuries were marked by the establishment of tropical botanical gardens as a tool of colonial expansion (for trade and commerce and, secondarily, science) mainly by
12300-401: The terms of its charter. It may include greenhouses, test grounds, an herbarium, an arboretum, and other departments. It maintains a scientific as well as a plant-growing staff, and publication is one of its major modes of expression. This broad outline is then expanded: The botanic garden may be an independent institution, a governmental operation, or affiliated to a college or university. If
12423-403: The time that Hainhofer visited them, they extended within the ramparts, around the castle as well as the inner courtyards. The Prince-Bishop had a belvedere balcony (altane) constructed in front of his room full of plants and small trees in containers, visible through the glass windows. Here he would scatter food to attract birds. Water to irrigate also flowed through a pipe in the room, so that
12546-590: The tip eventually stops growing, whether because of pruning or flowering, lateral buds take over and grow into new, fully functional, vines. Tomato vines are typically pubescent, meaning covered with fine short hairs. The hairs facilitate the vining process, turning into roots wherever the plant is in contact with the ground and moisture, especially if the vine's connection to its original root has been damaged or severed. The leaves are 10–25 cm (4–10 in) long, odd pinnate , with five to nine leaflets on petioles , each leaflet up to 8 cm (3 in) long, with
12669-518: The tomato in England. Gerard knew the tomato was eaten in Spain and Italy. Nonetheless, he believed it was poisonous. Gerard's views were influential, and the tomato was considered unfit for eating for many years in Britain and its North American colonies . By 1820, tomatoes were described as "to be seen in great abundance in all our vegetable markets" and to be "used by all our best cooks", reference
12792-411: The tomato in the genus Solanum (alongside the potato) as Solanum lycopersicum . In 1768, Philip Miller moved it to its own genus, naming it Lycopersicon esculentum . The name came into wide use, but was technically in breach of the plant naming rules because Linnaeus's species name lycopersicum still had priority. Although the name Lycopersicum lycopersicum was suggested by Karsten (1888), it
12915-588: The tomato, the red-fruited Solanum pimpinellifolium , is native to western South America, where it was probably first domesticated. The resulting domesticated plant, ancestral to the modern large-fruited tomato varieties, was probably the cherry tomato, S. lycopersicum var. cerasiforme . However, genomic analysis suggests that the domestication process may have been more complex than this. S. lycopersicon var. cerasiforme may have existed before domestication, while traits supposedly typical of domestication may have been reduced in that variety and then reselected (in
13038-773: The tomato." Livingston's first breed of tomato, the Paragon, was introduced in 1870. In 1875, he introduced the Acme, said to be in the parentage of most cultivars for the next twenty-five years. Other early breeders included Henry Tilden in Iowa and a Dr. Hand in Baltimore. Because of the tomato's need for heat and a long growing season, several states in the Sun Belt became major producers, particularly Florida and California . In California, tomatoes are grown under irrigation for both
13161-666: The total, followed by India , Turkey , and the United States as major producers (table). The world dedicated 4.8 million hectares in 2012 for tomato cultivation and the total production was about 161.8 million tonnes . The average world farm yield for tomato was 33.6 tonnes per hectare in 2012. Tomato farms in the Netherlands were the most productive in 2012, with a nationwide average of 476 tonnes per hectare, followed by Belgium (463 tonnes per hectare) and Iceland (429 tonnes per hectare). Common tomato pests include
13284-847: The tropics was the Pamplemousses Botanical Garden in Mauritius , established in 1735 to provide food for ships using the port, but later trialling and distributing many plants of economic importance. This was followed by the West Indies ( Saint Vincent and the Grenadines Botanic Gardens , 1764) and in 1786 by the Acharya Jagadish Chandra Bose Botanical Garden in Calcutta , India founded during
13407-424: The typical green ring around the stem on un-crossbred varieties. Before this, most tomatoes produced more sugar during ripening, and were sweeter and more flavorful. 10–20% of the total carbon fixed in the fruit can be produced by photosynthesis in the developing fruit of the normal U phenotype. The u mutation encodes a factor that produces defective chloroplasts with lower density in developing fruit, making them
13530-519: The various kinds of organizations known as botanical gardens, there are many that are in modern times public gardens with little scientific activity, and it cited a tighter definition published by the World Wildlife Fund and IUCN when launching the "Botanic Gardens Conservation Strategy" in 1989: "A botanic garden is a garden containing scientifically ordered and maintained collections of plants, usually documented and labelled, and open to
13653-399: The well known and highly prized San Marzano tomato grown in that region, with a European protected designation of origin certification. Tomatoes were not grown in England until the 1590s. One of the earliest cultivators was John Gerard , a barber-surgeon . Gerard's Herbal , published in 1597, and largely plagiarized from continental sources, is also one of the earliest discussions of
13776-439: Was a perfect square divided into quarters for the four continents, but by 1720, though, it was a rambling system of beds, struggling to contain the novelties rushing in, and it became better known as the hortus academicus . His Exoticorum libri decem (1605) is an important survey of exotic plants and animals that is still consulted today. The inclusion of new plant introductions in botanic gardens meant their scientific role
13899-558: Was called the herbularis or hortus medicus —more generally known as a physic garden, and a viridarium or orchard. These gardens were probably given impetus when Charlemagne issued a capitulary , the Capitulary de Villis, which listed 73 herbs to be used in the physic gardens of his dominions. Many of these were found in British gardens even though they only occurred naturally in continental Europe, demonstrating earlier plant introduction. Pope Nicholas V set aside part of
14022-469: Was damaged during the Thirty Years' War (1618-1648). Restoration began in 1648 under Prince-Bishop Marquard II Schenk von Castell (1637–1685), and the last time the garden was intact was under Prince-Bishop Johann Anton I Knebel von Katzenelnbogen (1705–1725). After that, there was only a kitchen garden. The gardeners lived outside the castle. From ca. 1710–12, the so-called botanist's house below
14145-427: Was first published in 1613 on behalf of the Prince-Bishop. Gardens at Willibaldsburg date back at least as early as Prince Bishop Martin von Schaumberg (1560–1590) who laid out "new gardens […] behind the palace". The plan of Schaumberg's garden was described by stating that he had "run walls around the castle and the gardens, also renovated the fountains and water-works and put them to better use in various places in
14268-556: Was formed in 1954 as a worldwide organisation affiliated to the International Union of Biological Sciences . More recently, coordination has also been provided by Botanic Gardens Conservation International (BGCI), which has the mission "To mobilise botanic gardens and engage partners in securing plant diversity for the well-being of people and the planet". BGCI has over 700 members – mostly botanic gardens – in 118 countries, and strongly supports
14391-485: Was led by Bernd Ringholz and the planning and layout as well as procurement of historical varieties and heir planting took almost five years. The area around the Willibaldsburg has undergone major changes since the original garden fell into neglect in the 18th century, making it impossible to reconstruct the original. The current garden occupies about 1500 sq m. and is laid out in the shape of an open book, as
14514-517: Was made to their cultivation in gardens still "for the singularity of their appearance", while their use in cooking was associated with exotic Italian or Jewish cuisine. For example, in Elizabeth Blackwell 's A Curious Herbal , it is described under the name "Love Apple ( Amoris Pomum )" as being consumed with oil and vinegar in Italy, similar to consumption of cucumbers in the UK. In 1963, The New York Times gave an explanation of
14637-590: Was marked by introductions from the Cape of South Africa – including ericas , geraniums , pelargoniums , succulents, and proteaceous plants – while the Dutch trade with the Dutch East Indies resulted in a golden era for the Leiden and Amsterdam botanical gardens and a boom in the construction of conservatories. The Royal Gardens at Kew were founded in 1759, initially as part of
14760-658: Was now widening, as botany gradually asserted its independence from medicine. In the mid to late 17th century, the Paris Jardin des Plantes was a centre of interest with the greatest number of new introductions to attract the public. In England , the Chelsea Physic Garden was founded in 1673 as the "Garden of the Society of Apothecaries". The Chelsea garden had heated greenhouses , and in 1723 appointed Philip Miller (1691–1771) as head gardener . He had
14883-514: Was ornamental, not least because it was understood to be related to the nightshades and assumed to be poisonous. The exact date of domestication is unknown; by 500 BC, it was already being cultivated in southern Mexico and probably other areas. The Pueblo people believed that tomato seeds could confer powers of divination . The large, lumpy variety of tomato, a mutation from a smoother, smaller fruit, originated in Mesoamerica, and may be
15006-536: Was published in Naples in 1692, though the author had apparently obtained these recipes from Spanish sources. Varieties were developed over the following centuries for drying, for sauce, for pizzas, and for long-term storage. These varieties are usually known for their place of origin as much as by a variety name. For example, there is the Pomodorino del Piennolo del Vesuvio , the "hanging tomato of Vesuvius", and
15129-442: Was the investigation of the local flora for its economic potential to both the colonists and the local people. Many crop plants were introduced by or through these gardens – often in association with European botanical gardens such as Kew or Amsterdam – and included cloves , tea , coffee , breadfruit, cinchona , sugar , cotton , palm oil and Theobroma cacao (for chocolate). During these times,
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