63-412: Elapidae ( / ə ˈ l æ p ə d iː / , commonly known as elapids / ˈ ɛ l ə p ə d z / , from Ancient Greek : ἔλαψ élaps , variant of ἔλλοψ éllops "sea-fish") is a family of snakes characterized by their permanently erect fangs at the front of the mouth. Most elapids are venomous , with the exception of the genus Emydocephalus . Many members of this family exhibit
126-543: A pitch accent . In Modern Greek, all vowels and consonants are short. Many vowels and diphthongs once pronounced distinctly are pronounced as /i/ ( iotacism ). Some of the stops and glides in diphthongs have become fricatives , and the pitch accent has changed to a stress accent . Many of the changes took place in the Koine Greek period. The writing system of Modern Greek, however, does not reflect all pronunciation changes. The examples below represent Attic Greek in
189-635: A threat display of rearing upwards while spreading out a neck flap. Elapids are endemic to tropical and subtropical regions around the world, with terrestrial forms in Asia, Australia, Africa, and the Americas and marine forms in the Pacific and Indian Oceans . Members of the family have a wide range of sizes, from the 18 cm (7.1 in) white-lipped snake to the 5.85 m (19 ft 2 in) king cobra . Most species have neurotoxic venom that
252-409: A bite can still result in a permanent scar , and in the worst cases, the affected limb may even have to be amputated . A victim's fate is impossible to predict, as this depends on many factors, including the species and size of the snake involved, how much venom was injected (if any), and the size and condition of the patient before being bitten. Viper bite victims may also be allergic to the venom or
315-525: A fifth major dialect group, or it is Mycenaean Greek overlaid by Doric, with a non-Greek native influence. Regarding the speech of the ancient Macedonians diverse theories have been put forward, but the epigraphic activity and the archaeological discoveries in the Greek region of Macedonia during the last decades has brought to light documents, among which the first texts written in Macedonian , such as
378-435: A pair of proteroglyphous fangs to inject venom from glands located towards the rear of the upper jaw (except for the genus Emydocephalus , in which fangs are present as a vestigial feature but without venom production, as they have specialized toward a fish egg diet, making them the only non-venomous elapids). The fangs, which are enlarged and hollow, are the first two teeth on each maxillary bone. Usually only one fang
441-550: A prefix /e-/, called the augment . This was probably originally a separate word, meaning something like "then", added because tenses in PIE had primarily aspectual meaning. The augment is added to the indicative of the aorist, imperfect, and pluperfect, but not to any of the other forms of the aorist (no other forms of the imperfect and pluperfect exist). The two kinds of augment in Greek are syllabic and quantitative. The syllabic augment
504-519: A separate family, have adapted to a marine way of life in different ways and to various degrees. All have evolved paddle-like tails for swimming and the ability to excrete salt. Most also have laterally compressed bodies, their ventral scales are much reduced in size, their nostrils are located dorsally (no internasal scales ), and they give birth to live young ( viviparity ). The reduction in ventral scaling has greatly diminished their terrestrial mobility, but aids in swimming. Members of this family have
567-415: A short time. In predatory bites, factors that influence the amount of venom injected include the size of the prey, the species of prey, and whether the prey item is held or released. The need to label prey for chemosensory relocation after a bite and release may also play a role. In defensive bites, the amount of venom injected may be determined by the size or species of the predator (or antagonist), as well as
630-542: A strong Northwest Greek influence, and can in some respects be considered a transitional dialect, as exemplified in the poems of the Boeotian poet Pindar who wrote in Doric with a small Aeolic admixture. Thessalian likewise had come under Northwest Greek influence, though to a lesser degree. Pamphylian Greek , spoken in a small area on the southwestern coast of Anatolia and little preserved in inscriptions, may be either
693-510: A vowel or /n s r/ ; final stops were lost, as in γάλα "milk", compared with γάλακτος "of milk" (genitive). Ancient Greek of the classical period also differed in both the inventory and distribution of original PIE phonemes due to numerous sound changes, notably the following: The pronunciation of Ancient Greek was very different from that of Modern Greek . Ancient Greek had long and short vowels ; many diphthongs ; double and single consonants; voiced, voiceless, and aspirated stops ; and
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#1732772158143756-476: A wide range of light levels. Typically, vipers are nocturnal and ambush their prey . Compared to many other snakes, vipers often appear rather sluggish. Most are ovoviviparous : the eggs are retained inside the mother's body, and the young emerge living. However, a few lay eggs in nests. Typically, the number of young in a clutch remains constant, but as the weight of the mother increases, larger eggs are produced, yielding larger young. Viperid snakes are found in
819-440: A wide range of sizes. Drysdalia species are small serpents typically 50 cm (20 in) and down to 18 cm (7.1 in) in length. Cobras , mambas , and taipans are mid- to large sized snakes which can reach 2 m (6 ft 7 in) or above. The king cobra is the world's longest venomous snake with a maximum length of 5.85 m (19.2 ft) and an average mass of 6 kg (13 lb). All elapids have
882-418: Is added to stems beginning with consonants, and simply prefixes e (stems beginning with r , however, add er ). The quantitative augment is added to stems beginning with vowels, and involves lengthening the vowel: Some verbs augment irregularly; the most common variation is e → ei . The irregularity can be explained diachronically by the loss of s between vowels, or that of the letter w , which affected
945-536: Is also dual-purpose: first, it is used for defense and to immobilize prey, as with neurotoxic venoms; second, many of the venom's enzymes have a digestive function, breaking down molecules such as lipids , nucleic acids , and proteins. This is an important adaptation, as many vipers have inefficient digestive systems. Due to the nature of proteolytic venom, a viperid bite is often a very painful experience and should always be taken seriously, though it may not necessarily prove fatal. Even with prompt and proper treatment,
1008-523: Is believed that the Laticauda and the 'true sea snakes' evolved separately from Australasian land snakes. Asian cobras, coral snakes, and American coral snakes also appear to be monophyletic, while African cobras do not. The type genus for the Elapidae was originally Elaps , but the group was moved to another family. In contrast to what is typical of botany , the family Elapidae was not renamed. In
1071-500: Is channeled by their hollow fangs, and some may contain other toxic components in varying proportions. The family includes 55 genera with around 360 species and over 170 subspecies. Terrestrial elapids look similar to the Colubridae ; almost all have long, slender bodies with smooth scales, a head covered with large shields (and not always distinct from the neck), and eyes with rounded pupils. Also like colubrids, their behavior
1134-448: Is considered by some linguists to have been closely related to Greek . Among Indo-European branches with living descendants, Greek is often argued to have the closest genetic ties with Armenian (see also Graeco-Armenian ) and Indo-Iranian languages (see Graeco-Aryan ). Ancient Greek differs from Proto-Indo-European (PIE) and other Indo-European languages in certain ways. In phonotactics , ancient Greek words could end only in
1197-469: Is derived from the Latin word vipera , - ae , also meaning viper, possibly from vivus ("living") and parere ("to beget"), referring to the trait viviparity (giving live birth) common in vipers like most of the species of Boidae . All viperids have a pair of relatively long solenoglyphous (hollow) fangs that are used to inject venom from glands located towards the rear of the upper jaws, just behind
1260-1161: Is in place on each side at any time. The maxilla is intermediate in both length and mobility between typical colubrids (long, less mobile) and viperids (very short, highly mobile). When the mouth is closed, the fangs fit into grooved slots in the buccal floor and usually below the front edge of the eye and are angled backwards; some elapids ( Acanthophis , taipan, mamba, and king cobra) have long fangs on quite mobile maxillae and can make fast strikes. A few species are capable of spraying their venom from forward-facing holes in their fangs for defense, as exemplified by spitting cobras . Most elapids are terrestrial , while some are strongly arboreal (African Pseudohaje and Dendroaspis , Australian Hoplocephalus ). Many species are more or less specialized burrowers (e.g. Ogmodon , Parapistocalamus , Simoselaps , Toxicocalamus , and Vermicella ) in either humid or arid environments. Some species have very generalised diets ( euryphagy ), but many taxa have narrow prey preferences (stenophagy) and correlated morphological specializations, for example feeding almost exclusively on other serpents (especially
1323-688: Is usually quite active and fast, with most of the females being oviparous (egg-layers). Exceptions to these generalizations occur; for example, certain adders ( Acanthophis ) have commonalities with the Viperidae family, such as shorter, stout bodies, rough/keeled scales, broad heads, cat-like pupils and ovoviviparous (internal hatchings with live births). Furthermore, they can also be sluggish, ambush predators with partially fragmented head shields, similar to rattlesnakes or Gaboon vipers . Sea snakes (the Hydrophiinae ), sometimes considered to be
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#17327721581431386-647: The Classical period ( c. 500–300 BC ). Ancient Greek was the language of Homer and of fifth-century Athenian historians, playwrights, and philosophers . It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance . This article primarily contains information about the Epic and Classical periods of
1449-501: The Pella curse tablet , as Hatzopoulos and other scholars note. Based on the conclusions drawn by several studies and findings such as Pella curse tablet , Emilio Crespo and other scholars suggest that ancient Macedonian was a Northwest Doric dialect , which shares isoglosses with its neighboring Thessalian dialects spoken in northeastern Thessaly . Some have also suggested an Aeolic Greek classification. The Lesbian dialect
1512-407: The antivenom . These snakes can decide how much venom to inject depending on the circumstances. The most important determinant of venom expenditure is generally the size of the snake; larger specimens can deliver much more venom. The species is also important, since some are likely to inject more venom than others, may have more venom available, strike more accurately, or deliver a number of bites in
1575-603: The epic poems , the Iliad and the Odyssey , and in later poems by other authors. Homeric Greek had significant differences in grammar and pronunciation from Classical Attic and other Classical-era dialects. The origins, early form and development of the Hellenic language family are not well understood because of a lack of contemporaneous evidence. Several theories exist about what Hellenic dialect groups may have existed between
1638-501: The present , future , and imperfect are imperfective in aspect; the aorist , present perfect , pluperfect and future perfect are perfective in aspect. Most tenses display all four moods and three voices, although there is no future subjunctive or imperative. Also, there is no imperfect subjunctive, optative or imperative. The infinitives and participles correspond to the finite combinations of tense, aspect, and voice. The indicative of past tenses adds (conceptually, at least)
1701-1031: The 5th century BC. Ancient pronunciation cannot be reconstructed with certainty, but Greek from the period is well documented, and there is little disagreement among linguists as to the general nature of the sounds that the letters represent. /oː/ raised to [uː] , probably by the 4th century BC. Greek, like all of the older Indo-European languages , is highly inflected. It is highly archaic in its preservation of Proto-Indo-European forms. In ancient Greek, nouns (including proper nouns) have five cases ( nominative , genitive , dative , accusative , and vocative ), three genders ( masculine , feminine , and neuter ), and three numbers (singular, dual , and plural ). Verbs have four moods ( indicative , imperative , subjunctive , and optative ) and three voices (active, middle, and passive ), as well as three persons (first, second, and third) and various other forms. Verbs are conjugated through seven combinations of tenses and aspect (generally simply called "tenses"):
1764-913: The Americas, Africa, Eurasia, and South Asia. In the Americas, they are native from south of 48°N . In the Old World , viperids are located everywhere except Siberia , Ireland, and north of the Arctic Circle in Norway and Sweden. Wild viperids are not found in Australia . The common adder , a viperid, is the only venomous snake found in Great Britain . Viperid venoms typically contain an abundance of protein -degrading enzymes, called proteases , that produce symptoms such as pain, strong local swelling and necrosis , blood loss from cardiovascular damage complicated by coagulopathy , and disruption of
1827-495: The Archaic period of ancient Greek (see Homeric Greek for more details): Μῆνιν ἄειδε, θεά, Πηληϊάδεω Ἀχιλῆος οὐλομένην, ἣ μυρί' Ἀχαιοῖς ἄλγε' ἔθηκε, πολλὰς δ' ἰφθίμους ψυχὰς Ἄϊδι προΐαψεν ἡρώων, αὐτοὺς δὲ ἑλώρια τεῦχε κύνεσσιν οἰωνοῖσί τε πᾶσι· Διὸς δ' ἐτελείετο βουλή· ἐξ οὗ δὴ τὰ πρῶτα διαστήτην ἐρίσαντε Ἀτρεΐδης τε ἄναξ ἀνδρῶν καὶ δῖος Ἀχιλλεύς. The beginning of Apology by Plato exemplifies Attic Greek from
1890-881: The Classical period of ancient Greek. (The second line is the IPA , the third is transliterated into the Latin alphabet using a modern version of the Erasmian scheme .) Ὅτι [hóti Hóti μὲν men mèn ὑμεῖς, hyːmêːs hūmeîs, Viperidae Vipers are snakes in the family Viperidae , found in most parts of the world, except for Antarctica , Australia , Hawaii , Madagascar , New Zealand , Ireland , and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs that permit deep penetration and injection of their venom . Three subfamilies are currently recognized. They are also known as viperids . The name "viper"
1953-545: The Dorians. The Greeks of this period believed there were three major divisions of all Greek people – Dorians, Aeolians, and Ionians (including Athenians), each with their own defining and distinctive dialects. Allowing for their oversight of Arcadian, an obscure mountain dialect, and Cypriot, far from the center of Greek scholarship, this division of people and language is quite similar to the results of modern archaeological-linguistic investigation. One standard formulation for
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2016-630: The IUCN red-list and CITES Apenndix lists. Some of the protected species are: This however does not touch the number of elapidae that are under threat, for instance 9% of elapid sea snakes are threatened with another 6% near-threatened. A rather large road block that stands in the way of more species being put under protection is lack of knowledge of the taxa; many known species have little research done on their behaviors or actual population as they live in very remote areas or live in habitats that are so vast its nearly impossible to conduct population studies, like
2079-802: The Indian Ocean and the south-west Pacific. They occupy coastal waters and shallows, and are common in coral reefs. However, the range of Hydrophis platurus extends across the Pacific to the coasts of Central and South America. Venoms of species in the Elapidae are mainly neurotoxic for immobilizing prey and defense. The main group of toxins are PLA2 and three-finger toxins (3FTx). Other toxic components in some species comprise cardiotoxins and cytotoxins , which cause heart dysfunctions and cellular damage, respectively. Cobra venom also contains hemotoxins that clot or solidify blood. Most members are venomous to varying extents, and some are considered among
2142-499: The Laticaudinae (sea kraits). Currently, none are universally recognized. Molecular evidence via techniques like karyotyping, protein electrophoretic analyses, immunological distance and DNA sequencing, suggests reciprocal monophyly of two groups: African, Asian, and New World Elapinae versus Australasian and marine Hydrophiinae . The Australian terrestrial elapids are technically 'hydrophiines', although they are not sea snakes. It
2205-415: The ability to detect thermal radiation emitted by warm-blooded animals , helping them better understand their environment. Internally the organ forms a small pit lined with membranes, external and internal, attached to the trigeminal nerve . Infrared light signals the internal membranes, which in turn signal the trigeminal nerve and send the infrared signals to the brain, where they are overlaid onto
2268-550: The aorist. Following Homer 's practice, the augment is sometimes not made in poetry , especially epic poetry. The augment sometimes substitutes for reduplication; see below. Almost all forms of the perfect, pluperfect, and future perfect reduplicate the initial syllable of the verb stem. (A few irregular forms of perfect do not reduplicate, whereas a handful of irregular aorists reduplicate.) The three types of reduplication are: Irregular duplication can be understood diachronically. For example, lambanō (root lab ) has
2331-486: The assessed level of threat, although larger assailants and higher threat levels may not necessarily lead to larger amounts of venom being injected. Hemotoxic venom takes more time than neurotoxic venom to immobilize prey, so viperid snakes need to track down prey animals after they have been bitten, in a process known as "prey relocalization". Vipers are able to do this via certain proteins contained in their venom. This important adaptation allowed rattlesnakes to evolve
2394-419: The augment when it was word-initial. In verbs with a preposition as a prefix, the augment is placed not at the start of the word, but between the preposition and the original verb. For example, προσ(-)βάλλω (I attack) goes to προσ έ βαλoν in the aorist. However compound verbs consisting of a prefix that is not a preposition retain the augment at the start of the word: αὐτο(-)μολῶ goes to ηὐ τομόλησα in
2457-611: The blood-clotting system. Also being vasculotoxic in nature, viperine venom causes vascular endothelial damage and hemolysis . Death is usually caused by collapse in blood pressure. This is in contrast to elapid venoms, which generally contain neurotoxins that disable muscle contraction and cause paralysis. Death from elapid bites usually results from asphyxiation because the diaphragm can no longer contract, but this rule does not always apply; some elapid bites include proteolytic symptoms typical of viperid bites, while some viperid bites produce neurotoxic symptoms. Proteolytic venom
2520-615: The dialect of Sparta ), and Northern Peloponnesus Doric (including Corinthian ). All the groups were represented by colonies beyond Greece proper as well, and these colonies generally developed local characteristics, often under the influence of settlers or neighbors speaking different Greek dialects. After the conquests of Alexander the Great in the late 4th century BC, a new international dialect known as Koine or Common Greek developed, largely based on Attic Greek , but with influence from other dialects. This dialect slowly replaced most of
2583-530: The dialects is: West vs. non-West Greek is the strongest-marked and earliest division, with non-West in subsets of Ionic-Attic (or Attic-Ionic) and Aeolic vs. Arcadocypriot, or Aeolic and Arcado-Cypriot vs. Ionic-Attic. Often non-West is called 'East Greek'. Arcadocypriot apparently descended more closely from the Mycenaean Greek of the Bronze Age. Boeotian Greek had come under
Elapidae - Misplaced Pages Continue
2646-510: The divergence of early Greek-like speech from the common Proto-Indo-European language and the Classical period. They have the same general outline but differ in some of the detail. The only attested dialect from this period is Mycenaean Greek , but its relationship to the historical dialects and the historical circumstances of the times imply that the overall groups already existed in some form. Scholars assume that major Ancient Greek period dialect groups developed not later than 1120 BC, at
2709-408: The eyes. Each of the two fangs is at the front of the mouth on a short maxillary bone that can rotate back and forth. When not in use, the fangs fold back against the roof of the mouth and are enclosed in a membranous sheath. This rotating mechanism allows for very long fangs to be contained in a relatively small mouth. The left and right fangs can be rotated together or independently. During a strike,
2772-405: The ideal amount of predigestion for the lowest amount of venom. Almost all vipers have keeled scales , a stocky build with a short tail, and a triangle-shaped head distinct from the neck, owing to the location of the venom glands. The great majority have vertically elliptical, or slit-shaped, pupils that can open wide to cover most of the eye or close almost completely, which helps them to see in
2835-574: The king cobra and kraits ). Elapids may display a series of warning signs if provoked, either obviously or subtly. Cobras and mambas lift their inferior body parts, expand hoods, and hiss if threatened; kraits often curl up before hiding their heads down their bodies. In general, sea snakes are able to respire through their skin. Experiments with the yellow-bellied sea snake, Hydrophis platurus , have shown that this species can satisfy about 20% of its oxygen requirements in this manner, allowing for prolonged dives. The sea kraits ( Laticauda spp. ) are
2898-520: The language, which are the best-attested periods and considered most typical of Ancient Greek. From the Hellenistic period ( c. 300 BC ), Ancient Greek was followed by Koine Greek , which is regarded as a separate historical stage, though its earliest form closely resembles Attic Greek , and its latest form approaches Medieval Greek . There were several regional dialects of Ancient Greek; Attic Greek developed into Koine. Ancient Greek
2961-494: The meantime, Elaps was renamed Homoroselaps and moved back to the Elapidae. However, Nagy et al. (2005) regard it as a sister taxon to Atractaspis , which should have been assigned to the Atractaspididae . * Not including the nominate subspecies With the dangers the taxa presents given their venomous nature it is very difficult for activists and conservationists alike to get species on protection lists such as
3024-500: The mouth can open nearly 180° and the maxilla rotates forward, erecting the fangs as late as possible so that the fangs do not become damaged, as they are brittle. The jaws close upon impact and the muscular sheaths encapsulating the venom glands contract, injecting the venom as the fangs penetrate the target. This action is very fast; in defensive strikes, it will be more a stab than a bite. Viperids use this mechanism primarily for immobilization and digestion of prey. Pre-digestion occurs as
3087-508: The older dialects, although the Doric dialect has survived in the Tsakonian language , which is spoken in the region of modern Sparta. Doric has also passed down its aorist terminations into most verbs of Demotic Greek . By about the 6th century AD, the Koine had slowly metamorphosed into Medieval Greek . Phrygian is an extinct Indo-European language of West and Central Anatolia , which
3150-456: The only cure to treat elapidae bites. There are commercial monovalent and polyvalent antivenoms for cobras, mambas, and some other important elapids. Recently, experimental antivenoms based on recombinant toxins have shown that it is feasible to create antivenoms with a wide spectrum of coverage. The venom of spitting cobras is more cytotoxic rather than neurotoxic. It damages local cells, especially those in eyes, which are deliberately targeted by
3213-487: The perfect stem eilēpha (not * lelēpha ) because it was originally slambanō , with perfect seslēpha , becoming eilēpha through compensatory lengthening. Reduplication is also visible in the present tense stems of certain verbs. These stems add a syllable consisting of the root's initial consonant followed by i . A nasal stop appears after the reduplication in some verbs. The earliest extant examples of ancient Greek writing ( c. 1450 BC ) are in
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#17327721581433276-599: The sea snakes least adapted to aquatic life. Their bodies are less compressed laterally, and they have thicker bodies and ventral scaling. Because of this, they are capable of some land movement. They spend much of their time on land, where they lay their eggs and digest prey. Terrestrial elapids are found worldwide in tropical and subtropical regions, mostly in the Southern Hemisphere. Most prefer humid tropical environments, though there are many that can still be found in arid environments. Sea snakes occur mainly in
3339-513: The sea snakes. Ancient Greek language Ancient Greek ( Ἑλληνῐκή , Hellēnikḗ ; [hellɛːnikɛ́ː] ) includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek ( c. 1400–1200 BC ), Dark Ages ( c. 1200–800 BC ), the Archaic or Epic period ( c. 800–500 BC ), and
3402-531: The snakes. The venom may cause intense pain on contact with the eye and may lead to blindness. It is not lethal on skin if no wound provides any chance for the toxins to enter the bloodstream . The table below lists out all of the elapid genera and no subfamilies. In the past, many subfamilies were recognized, or have been suggested for the Elapidae, including the Elapinae, Hydrophiinae (sea snakes), Micrurinae (coral snakes), Acanthophiinae (Australian elapids), and
3465-423: The strike-and-release bite mechanism, which provided a huge benefit to snakes by minimizing contact with potentially dangerous prey animals. This adaptation, then, requires the snake to track down the bitten animal to eat it, in an environment full of other animals of the same species. Western diamondback rattlesnakes respond more actively to mouse carcasses that have been injected with crude rattlesnake venom. When
3528-517: The syllabic script Linear B . Beginning in the 8th century BC, however, the Greek alphabet became standard, albeit with some variation among dialects. Early texts are written in boustrophedon style, but left-to-right became standard during the classic period. Modern editions of ancient Greek texts are usually written with accents and breathing marks , interword spacing , modern punctuation , and sometimes mixed case , but these were all introduced later. The beginning of Homer 's Iliad exemplifies
3591-467: The time of the Dorian invasions —and that their first appearances as precise alphabetic writing began in the 8th century BC. The invasion would not be "Dorian" unless the invaders had some cultural relationship to the historical Dorians . The invasion is known to have displaced population to the later Attic-Ionic regions, who regarded themselves as descendants of the population displaced by or contending with
3654-408: The various components of the venom were separated out, the snakes responded to mice injected with two kinds of disintegrins , which are responsible for allowing the snakes to track down their prey. Type genus = Vipera Laurenti, 1768 Pit vipers have specialized sensory organs near the nostrils called heat-sensing pits. The location of this organ is unique to pit vipers. These pits have
3717-529: The venom contains proteases , which degrade tissues. Secondarily, it is used for self defense, though in cases with nonprey, such as humans, they may give a dry bite (not inject any venom). A dry bite allows the snake to conserve its precious reserve of venom, because once it has been depleted, time is needed to replenish it, leaving the snake vulnerable. In addition to being able to deliver dry bites, vipers can inject larger quantities of venom into larger prey targets, and smaller amounts into small prey. This causes
3780-443: The visual image created by the eyes. Whether family Viperidae is attributed to Oppel (1811), as opposed to Laurenti (1768) or Gray (1825), is subject to some interpretation. The consensus among leading experts, though, is that Laurenti used viperae as the plural of vipera (Latin for "viper", "adder", or "snake") and did not intend for it to indicate a family group taxon. Rather, it is attributed to Oppel, based on his Viperini as
3843-447: The world's most venomous snakes based upon their murine LD 50 values, such as the taipans. Large species, mambas and cobras included, are dangerous for their ability to inject large quantities of venom upon a single envenomation and/or striking at a high position proximal to the victim's brain , which is vulnerable to neurotoxicity. Antivenom is promptly required to be administered if bitten by any elapids. Specific antivenoms are
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#17327721581433906-480: Was Aeolic. For example, fragments of the works of the poet Sappho from the island of Lesbos are in Aeolian. Most of the dialect sub-groups listed above had further subdivisions, generally equivalent to a city-state and its surrounding territory, or to an island. Doric notably had several intermediate divisions as well, into Island Doric (including Cretan Doric ), Southern Peloponnesus Doric (including Laconian ,
3969-452: Was a pluricentric language , divided into many dialects. The main dialect groups are Attic and Ionic , Aeolic , Arcadocypriot , and Doric , many of them with several subdivisions. Some dialects are found in standardized literary forms in literature , while others are attested only in inscriptions. There are also several historical forms. Homeric Greek is a literary form of Archaic Greek (derived primarily from Ionic and Aeolic) used in
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