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51-642: A fen is a type of wetland. Fen , Fenn, Fens , Fenns , may also refer to: Fen Fens can be found around the world, but the vast majority are located at the mid to high latitudes of the Northern Hemisphere. They are dominated by sedges and mosses , particularly graminoids that may be rarely found elsewhere, such as the sedge species Carex exilis . Fens are highly biodiverse ecosystems and often serve as habitats for endangered or rare species, with species composition changing with water chemistry. They also play important roles in

102-447: A distinct type of wetland, shares many biogeochemical characteristics with other wetlands. Like all wetlands, they play an important role in nutrient cycling because they are located at the interface of aerobic (oxic) and anaerobic (anoxic) environments. Most wetlands have a thin top layer of oxygenated soil in contact with the atmosphere or oxygenated surface waters. Nutrients and minerals may cycle between this oxidized top layer and

153-458: A fen directly is particularly damaging because it lowers the water table. A lower water table can increase aeration and dry out peat, allowing for aerobic decomposition or burning of the organic matter in peat. Draining a fen indirectly by decreasing its water supply can be just as damaging. Disrupting groundwater flow into the fen with nearby human activities such as quarrying or residential development changes how much water and nutrients enter

204-592: A fen, especially its pH, are directly influenced by the type of rocks its groundwater supply contacts. pH is a major factor in determining fen species composition and richness, with more basic fens called "rich" and more acidic fens called "poor." Rich fens tend to be highly biodiverse and harbor a number of rare or endangered species, and biodiversity tends to decrease as the richness of fen decreases. Fens tend to be found above rocks that are rich in calcium, such as limestone . When groundwater flows past calcareous (calcium-rich) rocks like limestone ( calcium carbonate ),

255-464: A fuel. Pollutants can alter the chemistry of fens and facilitate invasion by invasive species . Common pollutants of fens include road salts, nutrients from septic tanks , and runoff of agricultural fertilizers and pesticides. Shakespeare used the term "fen-sucked" to describe the fog (literally: rising from marshes) in King Lear , when Lear says, "Infect her beauty, You fen-sucked fogs drawn by

306-671: A gradient from poor to rich, with bogs at the poor end, extremely rich fens at the rich end, and poor fens in between. In this context, "rich" and "poor" refer to the species richness, or how biodiverse a fen or bog is. The richness of these species is strongly influenced by pH and concentrations of calcium and bicarbonate. These factors assist in identifying where along the gradient a particular fen falls. In general, rich fens are minerotrophic , or dependent on mineral-rich groundwater, while bogs are ombrotrophic , or dependent on precipitation for water and nutrients. Poor fens fall between these two. Rich fens are strongly minerotrophic; that is,

357-611: A highly varied environment might provide as a haven for species during times of stress like flooding or drought. Freshwater swamp forests contain soft, unstable, and anoxic soil due to their waterlogged condition, which may have influenced the evolution of unique root adaptations in these trees that resemble those seen in a real mangrove forest. The vegetation in freshwater swamp forests is adapted to survive in waterlogged conditions. Common tree species include various types of mangroves , palms, and hardwood trees like mahogany and ebony . The tree canopy tends to be dense and can create

408-412: A large proportion of their water comes from mineral-rich ground or surface water. Fens that are more distant from surface waters such as rivers and lakes, however, are more rich than fens that are connected. This water is dominated by calcium and bicarbonate, resulting in a slightly acidic to slightly basic pH characteristic of rich fens. These conditions promote high biodiversity. Within rich fens, there

459-536: A major determinant of fen biota and biogeochemistry . Fen soils are constantly inundated because the water table is at or near the surface. The result is anaerobic (oxygen-free) soils due to the slow rate at which oxygen diffuses into waterlogged soil. Anaerobic soils are ecologically unique because Earth's atmosphere is oxygenated, while most terrestrial ecosystems and surface waters are aerobic. The anaerobic conditions found in wetland soils result in reduced , rather than oxidized , soil chemistry. A hallmark of fens

510-437: A number of environmental traits with other tropical rainforest formations. Beyond these shared characteristics, however, the environment in freshwater swamp forests and other tropical rain forest formations can vary greatly. In inland, freshwater swamp forests, flooding is typically sporadic, irregular, or seasonal. The depth of the water also varies greatly, from a few centimeters to several meters. These physical factors affect

561-418: A pH of approximately 5.5 to 4. Peat in poor fens tends to be thicker than that of rich fens, which cuts off vegetation access to the mineral-rich soil underneath. In addition, the thicker peat reduces the influence of mineral-rich groundwater that buffers the pH. This makes the fen more ombrotrophic, or dependent on nutrient-poor precipitation for its water and nutrients. Poor fens may also form in areas where

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612-539: A relatively understudied forest type in Southeast Asia, primarily because they are difficult to access and can harbor diseases spread by insects , such as mosquitoes . In the Amazon Basin of Brazil , a seasonally flooded forest is known as a várzea , and refers to a whitewater -inundated forest. Igapó refers to blackwater -inundated forest. Mangrove and peat swamp forests , for example, have

663-445: A shaded environment beneath. Freshwater swamp forests are rich in biodiversity and provide habitat for a wide range of plant and animal species. They support numerous aquatic species, amphibians , reptiles, and mammals. Birds are also abundant in these ecosystems. The flooded conditions can make these areas particularly important for fish breeding and spawning . Fires of various intensities were used in southern Sumatra to burn down

714-601: A small amount dissolves and is carried to the fen supplied by the groundwater. When calcium carbonate dissolves, it produces bicarbonate and a calcium cation according to the following equilibrium: CaCO 3 + H 2 CO 3 ↽ − − ⇀ Ca 2 + + 2 HCO 3 − {\displaystyle {\ce {CaCO3 + H2CO3 <=> Ca^2+ + 2HCO3^-}}} where carbonic acid (H 2 CO 3 )

765-407: A small number of taxa. Similar impacts have been seen in tropical ecosystems where flooding during the wet or monsoon season has led to an increase in migratory species proportions and changes in community assemblages. On a smaller scale, modifications to the flow regime that result in changes to the physical environment can boost habitat variety and, as a result, boost species diversity. Additionally,

816-696: A type of wetland, but its use is generally limited to the southern United States. Third, different languages use different terms to describe types of wetlands. For instance, in Russian, there is no equivalent word for the term swamp as it is typically used in North America. The result is a large number of wetland classification systems that each define wetlands and wetland types in their own way. However, many classification systems include four broad categories that most wetlands fall into: marsh , swamp, bog , and fen. While classification systems differ on

867-523: A variety of ecosystem services . They act as natural buffers against flooding by absorbing excess water during heavy rains. They also filter water, helping to improve water quality by trapping sediments and pollutants. These forests are crucial for carbon sequestration , playing a role in mitigating climate change . Local communities often rely on freshwater swamp forests for resources such as timber , non-timber forest products (e.g., fruits, honey), and fish. Traditional agriculture may take place in

918-551: A wetland is. Almost all of the phosphorus that arrives in a wetland does so through sediments or plant litter from other ecosystems. Along with nitrogen, phosphorus limits wetland fertility. Under basic conditions like those found in extremely rich fens, calcium will bind to phosphate anions to make calcium phosphates , which are unavailable for uptake by plants. Mosses also play a considerable role in aiding plants in phosphorus uptake by decreasing soil phosphorus stress and stimulating phosphatase activity in organisms found below

969-519: Is a large amount of variability. The richest fens are the extreme rich (marl) fens, where marl deposits are often build up. These are often pH 7 or greater. Rich and intermediate rich fens are generally neutral to slightly acidic, with a pH of approximately 7 to 5. Rich fens are not always very productive; at high calcium concentrations, calcium ions bind to phosphate anions, reducing the availability of phosphorus and decreasing primary production. Rich bogs with limited primary production can stabilize with

1020-470: Is a more potent greenhouse gas than carbon dioxide. Methanogenic archaea that reside in the anaerobic layers of peat combine carbon dioxide and hydrogen gas to form methane and water. This methane can then escape into the atmosphere and exert its warming effects. Peatlands dominated by brown mosses and sedges such as fens have been found to emit a greater amount of methane than Sphagnum -dominated peatlands such as bogs. Fens play an important role in

1071-442: Is a type of wetland ecosystem characterized by its unique hydrology and vegetation. These forests are typically found in low-lying areas, riverbanks, and floodplains where there is a consistent supply of freshwater . A forest that is frequently flooded with relatively fresh water rich in minerals is referred to as a freshwater swamp forest. Since tropical freshwater swamp forests are a subset of tropical rainforests, they share

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1122-419: Is abundant in peat. When the organic matter in peat is decomposed in the absence of oxygen, ammonium is produced via ammonification . In the oxidized surface layer of the wetland, this ammonium is oxidized to nitrite and nitrate by nitrification . The production of ammonium in the reduced layer and its consumption in the top oxidized layer drives upward diffusion of ammonium. Likewise, nitrate production in

1173-458: Is difficult for a number of reasons. First, wetlands are diverse and varied ecosystems that are not easily categorized according to inflexible definitions. They are often described as a transition between terrestrial and aquatic ecosystems with characteristics of both. This makes it difficult to delineate the exact extent of a wetland. Second, terms used to describe wetland types vary greatly by region. The term bayou , for example, describes

1224-478: Is estimated that there are approximately 1.1 million square kilometers of fens worldwide, but quantifying the extent of fens is difficult. Because wetland definitions vary regionally, not all countries define fens the same way. In addition, wetland data is not always available or of high quality. Fens are also difficult to rigidly delineate and measure, as they are located between terrestrial and aquatic ecosystems. Rigidly defining types of wetlands, including fens,

1275-420: Is greater than decomposition, which results in the accumulation of organic matter as peat. Resident mosses usually carry out decomposition within the fen, and temperate fens are often driven by plant roots' decomposition. These peat stores sequester an enormous amount of carbon. Nevertheless, it is difficult to determine whether fens net take up or emit greenhouse gases . This is because fens emit methane, which

1326-576: Is produced by the dissolution of carbon dioxide in water. In fens, the bicarbonate anion produced in this equilibrium acts as a pH buffer, which keeps the pH of the fen relatively stable. Fens supplied by groundwater that doesn't flow through minerals and act as a buffer when dissolved tend to be more acidic. The same effect is observed when groundwater flows through minerals with low solubility, such as sand. In extreme rich fens, calcium carbonate can precipitate out of solution to form marl deposits. Calcium carbonate precipitates out of solution when

1377-404: Is that a significant portion of their water supply is derived from groundwater (minerotrophy). Because hydrology is the dominant factor in wetlands, the chemistry of the groundwater has an enormous effect on the characteristics of the fen it supplies. Groundwater chemistry, in turn, is largely determined by the geology of the rocks that the groundwater flows through. Thus, the characteristics of

1428-719: The Okavango Delta in Botswana and the highland slopes in Lesotho . Fens can also be found at the colder latitudes of the Southern Hemisphere. They are found in New Zealand and southwest Argentina, but the extent is much less than that of the northern latitudes. Locally, fens are most often found at the intersection of terrestrial and aquatic ecosystems, such as the headwaters of streams and rivers. It

1479-430: The cycling of nutrients such as carbon, nitrogen, and phosphorus due to the lack of oxygen (anaerobic conditions) in waterlogged organic fen soils. Fens have historically been converted to agricultural land. Aside from such conversion, fens face a number of other threats, including peat cutting, pollution, invasive species, and nearby disturbances that lower the water table in the fen, such as quarrying. Interrupting

1530-432: The partial pressure of carbon dioxide in the solution falls. The decrease in carbon dioxide partial pressure is caused by uptake by plants for photosynthesis or direct loss to the atmosphere. This reduces the availability of carbonic acid in solution, shifting the above equilibrium back towards the formation of calcium carbonate. The result is the precipitation of calcium carbonate and the formation of marl. Fen, being

1581-811: The accumulation of mosses and mycorrhiza , which promote phosphorus cycling and can support the growth of new vegetation and bacteria. Brown mosses (family Amblystegiaceae ) and sedges (genus Carex ) are the dominant vegetation. However, an accumulation of mosses such as Sphagnum can lead to the acidification of the rich fen, potentially converting it into a poor fen. Compared to poor fens, rich fens have higher concentrations of bicarbonate, base cations (Na , Ca , K , Mg ), and sulfate . Poor fens are, in many ways, an intermediate between rich fens and bogs. Hydrologically, they are more similar to rich fens than to bogs, but regarding vegetation composition and chemistry, they are more similar to bogs than rich fens. They are much more acidic than their rich counterparts, with

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1632-556: The continuous input of groundwater stimulates production. Bogs , which lack this input of groundwater , have much lower primary production. Carbon from all types of wetlands, including fens, arrives mostly as organic carbon from either adjacent upland ecosystems or by photosynthesis in the wetland itself. Once in the wetland, organic carbon generally has three main fates: oxidation to CO 2 by aerobic respiration , burial as organic matter in peat, or decomposition to methane . In peatlands, including fens, primary production by plants

1683-879: The drier, elevated areas around the swamp forest . However, unsustainable logging and land conversion can threaten these ecosystems . Freshwater swamp forests are globally important and often designated as protected areas or Ramsar wetlands due to their ecological significance. Conservation efforts aim to protect these ecosystems from degradation and promote sustainable land use practices. Freshwater swamp forests are vulnerable to habitat destruction and degradation due to urbanization , agriculture , logging , and drainage for development. Climate change , including rising sea levels, can further threaten these ecosystems by altering water levels and salinity . Globally, freshwater swamp forests are found in Southeast Asia , tropical Africa , and South America , with

1734-564: The ecology of freshwater swamp forests, either singly or in combination. Rain, rivers, and groundwater are all sources of water for freshwater swamp forests, whereas rain is the only source of water for peat swamp forests. The hue of the water in the freshwater swamp forest is typically an indication of the quantities of plant materials present in the water and soil. Contrary to ombrotrophic swamp forests, which only absorb nutrients from rain, freshwater swamp forest soils are relatively nutrient-rich. In freshwater swamp forests, rain and changes in

1785-406: The exact criteria that define a fen, there are common characteristics that describe fens generally and imprecisely. A general definition provided by the textbook Wetlands describes a fen as "a peat-accumulating wetland that receives some drainage from surrounding mineral soil and usually supports marsh like vegetation." Three examples are presented below to illustrate more specific definitions for

1836-756: The fen. This can make the fen more ombrotrophic (dependent on precipitation), which results in acidification and a change in water chemistry. This directly impacts the habitat of these species, and many signature fen species disappear. Fens are also threatened by invasive species , fragmentation , peat cutting, and pollution. Non-native invasive species, such as the common buckthorn in North America, can invade fens and outcompete rare fen species, reducing biodiversity. Habitat fragmentation threatens fen species, especially rare or endangered species that are unable to move to nearby fens due to fragmentation. Peat cutting, while much more common in bogs, does happen in fens. Peat cut from fens has many uses, including burning as

1887-478: The flow of mineral-rich water into a fen changes the water chemistry, which can alter species richness and dry out the peat. Drier peat is more easily decomposed and can even burn. Fens are distributed around the world, but are most frequently found at the mid-high latitudes of the Northern Hemisphere. They are found throughout the temperate zone and boreal regions, but are also present in tundra and in specific environmental conditions in other regions around

1938-595: The geomorphology, affecting the spatial and temporal benthic community structure. Freshwater swamp forests are permanently or seasonally inundated by freshwater , with water levels that fluctuate throughout the year. They are often associated with slow-moving or meandering rivers , oxbow lakes , and other water bodies. Macroinvertebrate assemblages were significantly influenced by hydrological connectivity, with highly connected water bodies exhibiting more macroinvertebrate diversity than isolated water bodies, which tended to have less diverse assemblages and were predominated by

1989-459: The global nitrogen cycle due to the anaerobic conditions found in their soils, which facilitate the oxidation or reduction of one form of nitrogen to another. Most nitrogen arrives in wetlands as nitrate from runoff , in organic matter from other areas, or by nitrogen fixation in the wetland. There are three main forms of nitrogen found in wetlands: nitrogen in organic matter, oxidized nitrogen ( nitrate or nitrite ), and ammonium . Nitrogen

2040-554: The groundwater supplying the fen flows through sediments that don't dissolve well or have low buffering capacity when dissolved. Species richness tends to be lower than that of rich fens but higher than that of bogs. Poor fens, like bogs, are dominated by Sphagnum mosses, which acidify the fen and decrease nutrient availability. One of the many threats that fens face is conversion to agricultural lands. Where climates are suitable, fens have been drained for agricultural use alongside crop production, grazing , and hay making . Draining

2091-760: The largest areas being part of the Amazon basin . In Southeast Asia, they are found all over the region, frequently close to major rivers like the Mekong , Chao Phraya , and Irrawaddy in Thailand and Myanmar , as well as numerous smaller systems like the Sedili rivers in Johor . Although freshwater swamp forests are frequently found in wet climates, they can also be found in areas that are seasonally drier, such as west New Guinea and east Java . Freshwater swamp forests are

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2142-410: The moss cover. Helophytes have been shown to bolster phosphorus cycling within fens, especially in fen reestablishment, due to their ability to act as a phosphorus sink, which prevents residual phosphorus in the fen from being transferred away from the it. Under normal conditions, phosphorus is held within soil as dissolved inorganic phosphorus, or phosphate , which leaves trace amounts of phosphorus in

2193-429: The oxidized layer and nitrate consumption in the reduced layer by denitrification drives downward diffusion of nitrate. Denitrification in the reduced layer produces nitrogen gas and some nitrous oxide , which then exit the wetland to the atmosphere. Nitrous oxide is a potent greenhouse gas whose production is limited by nitrate and nitrite concentrations in fens. Nitrogen, along with phosphorus, controls how fertile

2244-464: The powerful sun, To fall and blister." Freshwater swamp forest Freshwater swamp forests , or flooded forests , are forests which are inundated with freshwater , either permanently or seasonally. They normally occur along the lower reaches of rivers and around freshwater lakes . Freshwater swamp forests are found in a range of climate zones , from boreal through temperate and subtropical to tropical . A freshwater swamp forest

2295-442: The presence of peat. In The Biology of Peatlands fens are defined by the following criteria: A further distinction is made between open and wooded fens, where open fens have canopy cover less than 10% and wooded fens have 10–25% canopy cover. If tall shrubs or trees dominate, the wetland is instead classified as a wooded bog or swamp forest , depending on other criteria. Hydrological conditions, as seen in other wetlands, are

2346-520: The reduced layer below, undergoing oxidation and reduction reactions by the microbial communities adapted to each layer. Many important reactions take place in the reduced layer, including denitrification , manganese reduction, iron reduction, sulfate reduction, and methanogenesis . Because wetlands are hotspots for nutrient transformations and often serve as nutrient sinks, they may be constructed to treat nutrient-rich waters created by human activities. Fens are also hotspots for primary production , as

2397-573: The rest of the ecosystem. Iron is important in phosphorus cycling within fens. Iron can bind to high levels of inorganic phosphate within the fen, leading to a toxic environment and inhibition of plant growth. In iron-rich fens, the area can become vulnerable to acidification, excess nitrogen and potassium, and low water levels. Peat soils play a role in preventing the bonding of irons to phosphate by providing high levels of organic anions for iron to bind to instead of inorganic anions such as phosphate. Bogs and fens can be thought of as two ecosystems on

2448-769: The swamp forest to make way for agriculture. There, a rich and complex environment was reduced to a habitat made up of uniform stands of fire-resistant Melaleuca L. species thickets as a result of widespread and frequent fires. Furthermore, improper management of these ecosystems through massive logging and agricultural conversion has resulted in serious deterioration and loss of ecological and biological diversity. Many plants in freshwater swamp forests have special adaptations to cope with waterlogged soils, such as pneumatophores (aerial roots) in mangroves for oxygen exchange. Some animals have adapted to life in these wet environments, like amphibians with permeable skin that can absorb oxygen from water. Freshwater swamp forests offer

2499-553: The term fen . In the Canadian Wetland Classification System, fens are defined by six characteristics: In the textbook Wetland Ecology: Principles and Conservation , Paul A. Keddy offers a somewhat simpler definition of a fen as "a wetland that is usually dominated by sedges and grasses rooted in shallow peat, often with considerable groundwater movement, and with pH greater than 6." This definition differentiates fens from swamps and marshes by

2550-420: The water table result in the following deposition of nutrients and alluvial soils. Due to the nutrient-rich soils of freshwater swamp forests, wetland rice farming and oil palm plantations have become very prevalent in agriculture. Within aquatic ecosystems, geomorphological, biological, and biogeomorphological processes and functions are significantly influenced by the flow of water. Flow significantly affects

2601-648: The world. In the United States, fens are most common in the Midwest and Northeast, but can be found across the country. In Canada, fens are most frequent in the lowlands near Hudson Bay and James Bay , but can also be found across the country. Fens are also spread across the northern latitudes of Eurasia, including Britain and Ireland, as well as Japan, but east-central Europe is especially rich in fens. Further south, fens are much rarer, but do exist under specific conditions. In Africa, fens have been found in

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