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120-600: Botswana , officially the Republic of Botswana , is a landlocked country in Southern Africa . Botswana is topographically flat, with approximately 70 per cent of its territory part of the Kalahari Desert . It is bordered by South Africa to the south and southeast, Namibia to the west and north, Zambia to the north and Zimbabwe to the northeast. With a population of slightly over 2.4 million people and

240-413: A 'grey zone' between electoral democracy and electoral autocracy . Furthermore, they show that Botswana lost its status as a "liberal democracy" in 2021, with its liberal, participatory and deliberative components decreasing "at a statistically significant level", with the latter component being noted as becoming "significantly worse". The 2023 Transparency International Corruption Index ranks Botswana

360-509: A GDP per capita of about US$ 70 per year in the late 1960s—Botswana has transformed itself into an upper middle-income country. GDP per capita grew from $ 439 in 1950 to $ 15,842 in 2018. Although Botswana was resource-abundant, a good institutional framework allowed the country to reinvest resource-income to generate stable future income. By one estimate, it has the fourth-highest gross national income at purchasing power parity in Africa, giving it

480-418: A blood meal equal in weight to themselves. The dipteran crop is heavily understudied, with Glossina being one of the few genera having relatively reliable information available: Moloo and Kutuza 1970 for G. brevipalpis (including its innervation) and Langley 1965 for G. morsitans . The reproductive tract of adult females includes a uterus , which can become large enough to hold the third-instar larva at

600-638: A border with a coastal country but any path to a coast would still lead across at least two borders. The Free City of Frankfurt which was independent between 1815 and 1866 was doubly landlocked as it bordered the Electorate of Hesse , the Grand Duchy of Hesse , Hesse-Homburg , and Nassau . In the German Confederation there were several other landlocked states that only bordered landlocked states and landlocked exclaves of coastal states:

720-456: A comparable land area to France , Botswana is one of the most sparsely populated countries in the world. It is essentially the nation-state of the Tswana people , who constitute nearly 80 per cent of the population. The Tswana ethnic group are descended mainly from Bantu-speaking peoples who migrated into southern Africa , including modern Botswana, in several waves before AD 600. In 1885,

840-720: A consensus estimate. In that era, the ancestors of the modern-day Kalanga moved into what is now the north-eastern area of the country. These proto-Kalanga were closely connected to states in Zimbabwe as well as to the Mapungubwe state. One notable remanant of this period is Domboshaba ruins, a cultural and heritage site in Botswana initially occupied towards the end of the Great Zimbabwe period (1250–1450), with stone walls that have an average height of 1.8 metres. The site

960-537: A country's development. It cuts a nation off from important sea resources such as fishing , and impedes or prevents direct access to maritime trade , a crucial component of economic and social advance. As such, coastal regions, or inland regions that have access to the World Ocean, tended to be wealthier and more heavily populated than inland regions that have no access to the World Ocean. Paul Collier in his book The Bottom Billion argues that being landlocked in

1080-602: A democratic self-government in Botswana. An independence conference was held in London in February 1966. The seat of government was moved in 1965 from Mahikeng in South Africa, to the newly established Gaborone , located near Botswana's border with South Africa. Based on the 1965 constitution, the country held its first general elections under universal suffrage and gained independence on 30 September 1966. Seretse Khama ,

1200-410: A few hours, and is almost never observed outside of the laboratory. Tsetse next develops a hard external case, the puparium, and become pupae - small, hard-shelled oblongs with two distinctively small, dark lobes at the tail (breathing) end. Tsetse pupae are under 1 centimetre ( 1 ⁄ 2  in) long. Within the puparial shell, tsetse complete the last two larval instars and the pupal stage. At

1320-413: A half years of current imports. The constitution provides for an independent judiciary, and the government respects this in practice. The legal system is sufficient to conduct secure commercial dealings, although a growing backlog of cases prevents timely trials. Botswana is ranked second only to South Africa among sub-Saharan African countries in the 2014 International Property Rights Index. In Botswana,

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1440-404: A large bulb. The thorax is large, made of three fused segments. Three pairs of legs are attached to the thorax, as are two wings and two halteres . The abdomen is short but wide and changes dramatically in volume during feeding. The internal anatomy of the tsetse is fairly typical of the insects ; the crop is large enough to accommodate a huge increase in size during feeding, as tsetse can take

1560-457: A leader in the independence movement, was elected as the first president, and subsequently re-elected twice. Khama died in office in 1980. The presidency passed to the sitting vice-president, Quett Masire , who was elected in his own right in 1984 and re-elected in 1989 and 1994. Masire retired from office in 1998. He was succeeded by Festus Mogae , who was elected in 1999 and re-elected in 2004. The presidency passed in 2008 to Ian Khama (son of

1680-577: A leader named Kgabo II, made their way into the southern Kalahari by AD 1500, at the latest, and his people drove the Bakgalagadi inhabitants west into the desert. Over the years, several offshoots of the Bakwena moved into adjoining territories. The Bangwaketse occupied areas to the west, while the Bangwato moved northeast into former Kalanga areas. Not long afterwards, a Bangwato offshoot known as

1800-443: A poor geographical neighbourhood is one of four major development "traps" by which a country can be held back. In general, he found that when a neighbouring country experiences better growth, it tends to spill over into favorable development for the country itself. For landlocked countries, the effect is particularly strong, as they are limited in their trading activity with the rest of the world. He states, "If you are coastal, you serve

1920-413: A pregnant tsetse and her offspring, so all newly emerged tsetse adults are free of infection. An uninfected fly that feeds on an infected vertebrate animal may acquire trypanosomes in its proboscis or gut. These trypanosomes, depending on the species, may remain in place, move to a different part of the digestive tract, or migrate through the tsetse body into the salivary glands. When an infected tsetse bites

2040-490: A project to reintroduce indigenous vegetation into communities in Kgalagadi South, Kweneng North and Boteti. Reintroduction of indigenous vegetation will help reduce the degradation of the land. The United States Government has also entered into an agreement with Botswana, giving them US$ 7 million to reduce Botswana's debt by US$ 8.3 million. The US stipulated that Botswana will focus on more extensive conservation of

2160-433: A relatively high standard of living in Africa, around that of Mexico. As of 2022, the unemployment rate stood at 25.4%, while youth unemployment reached 45.41% in 2023. The latest available data from 2015/2016 estimate that 17.2% of Botswana’s population is multidimensionally poor , with an additional 19.7% at risk. The Ministry of Trade and Industry of Botswana is responsible for promoting business development throughout

2280-399: A susceptible host, the fly may regurgitate part of a previous blood meal that contains trypanosomes, or may inject trypanosomes in its saliva. Inoculation must contain a minimum of 300 to 450 individual trypanosomes to be successful, and may contain up to 40,000 cells. In the case of T. b. brucei infecting G. p. gambiensis , during this time the parasite changes the proteome contents of

2400-440: A time; she will retain each egg within her uterus, the offspring developing internally (during the first three larval stages ), in an adaptation called adenotrophic viviparity . During this time, the female feeds the developing offspring with a milky substance (secreted by a modified gland) in the uterus. In the third larval stage, the tsetse larvae leave the uterus and begin an independent life. The newly-birthed larvae crawl into

2520-438: A tsetse bite leading to an inoculation in the subcutaneous tissue. The infection moves into the lymphatic system , leading to a characteristic swelling of the lymph glands called Winterbottom's sign . The infection progresses into the blood stream and eventually crosses into the central nervous system and invades the brain leading to extreme lethargy and eventually to death . The species Trypanosoma brucei , which causes

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2640-514: A vital role in the longevity of Toutswemogala Hill's extended occupation, as many grain storage structures have also been found on the site. Many different stratified layers of housing floors further signal continuous occupation over hundreds of years. The arrival of the Tswana speakers' ancestors who came to control the region has yet to be dated precisely. Members of the Bakwena , a chieftaincy under

2760-754: Is "doubly landlocked" or "double-landlocked" when it is surrounded entirely by landlocked countries (i.e. requiring the crossing of at least two national borders to reach a coastline). There are two such countries: After the dissolution of the Holy Roman Empire , the Kingdom of Württemberg became a doubly landlocked state, bordering Bavaria , Baden , Switzerland , the Grand Duchy of Hesse ( Wimpfen exclave), Hohenzollern-Sigmaringen , and Hohenzollern-Hechingen . The latter two were themselves landlocked between each other, Württemberg, and Baden. In 1866 they became an exclave of Prussia , giving Württemberg

2880-409: Is a legal penalty for murder in Botswana, and executions are carried out by hanging. Many of the indigenous San people have been forcibly relocated from their land to reservations. To make them relocate, they were denied access to water on their land and faced arrest if they hunted, which was their primary source of food. Their lands lie in the middle of the world's richest diamond field. Officially,

3000-519: Is a respected place for the people living in the region, and it is believed that the chief lived on the top of the hill with his helpers or assistants. These states, located outside of current Botswana's borders, appear to have kept massive herds of cattle—apparently at numbers approaching modern cattle density—in what is now the Central District. This massive cattle-raising complex prospered until around 1300 and seems to have regressed following

3120-476: Is also altered: Production of aromatic L-amino acid decarboxylase - involved in dopamine and serotonin synthesis - and α-methyldopa hypersensitive protein was induced. This is very similar to the alterations in other dipteran vectors' head proteomes under infection by other eukaryotic parasites of mammals, found in another study by the same team in the same year. The trypanosomes are injected into vertebrate muscle tissue, but make their way, first into

3240-465: Is called sleeping sickness . In animals, tsetse-vectored trypanosomiases include nagana , souma (a French term which may not be a distinct condition ), and surra according to the animal infected and the trypanosome species involved. The usage is not strict and while nagana generally refers to the disease in cattle and horses it is commonly used for any of the animal trypanosomiases. Trypanosomes are animal parasites , specifically protozoans of

3360-402: Is not yet well understood. However, since the sexual phase of the trypanosome life cycle occurs within the tsetse host, biological transmission is a required step in the life cycle of the tsetse-vectored trypanosomes. The cycle of biological transmission of trypanosomiasis involves two phases, one inside the tsetse host and the other inside the vertebrate host. Trypanosomes are not passed between

3480-514: Is predominantly flat, tending towards gently rolling tableland . Botswana is dominated by the Kalahari Desert , which covers up to 70% of its land surface. The Limpopo River Basin, the major landform of all of southern Africa, lies partly in Botswana, with the basins of its tributaries, the Notwane , Bonwapitse , Mahalapye , Lotsane , Motloutse and the Shashe , located in the eastern part of

3600-492: Is the third-least corrupt country in Africa, according to the Corruption Perceptions Index published by Transparency International . The economy is dominated by mining and tourism . Botswana has a per capita GDP ( purchasing power parity ) of about $ 20,158 as of 2024. Botswana is the world's biggest diamond-producing country. Its relatively high gross national income per capita (by some estimates

3720-721: Is the third-least corrupt country in Africa, just below Cabo Verde and the Seychelles . Botswana is also a member of the Commonwealth of Nations . At the time of independence, Botswana had no armed forces. It was only after the Rhodesian and South African armies attacked the Zimbabwe People's Revolutionary Army and Umkhonto we Sizwe bases respectively that the Botswana Defence Force (BDF)

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3840-408: Is to draw from "indigenous knowledge and traditional land management systems". The leaders of this movement are supposed to be the people in the community to draw indigenous people in, which in turn increases their opportunities to earn an income, thus decreasing poverty. The UNDP also stated that the government has to effectively implement policies to allow people to manage their own local resources and

3960-675: Is usually not considered a continent but a geographical region by the English-speaking countries ) also has no landlocked countries. All landlocked countries, except Bolivia and Paraguay , are located on the continental mainland of Afro-Eurasia . Tsetse fly Tsetse ( / ˈ s iː t s i / SEET -see , US : / ˈ t s iː t s i / TSEET -see or UK : / ˈ t s ɛ t s ə / TSET -sə ) (sometimes spelled tzetze ; also known as tik-tik flies) are large, biting flies that inhabit much of tropical Africa . Tsetse flies include all

4080-413: Is wider, rather than elongated, and shorter than the wings. Four characteristics collectively separate adult tsetse from other kinds of flies: Like all other insects , tsetse flies have an adult body comprising three visibly distinct parts: the head, the thorax, and the abdomen. The head has large eyes, distinctly separated on each side, and a distinct, forward-pointing proboscis attached underneath by

4200-771: The Baltic Sea , Black Sea , and Pacific Ocean . On the other hand, some landlocked countries can have access to the ocean along wide navigable rivers. For instance, Paraguay (and Bolivia to a lesser extent) have access to the ocean through the Paraguay and Paraná rivers. Several countries have coastlines on landlocked bodies of water, such as the Caspian Sea and the Dead Sea . Since these seas are in effect lakes without access to wider seaborne trade, countries such as Kazakhstan are still considered landlocked. Although

4320-573: The Battle of Dimawe and, after about eight years of intermittent tensions and hostilities, eventually came to a peace agreement in Potchefstroom in 1860. From that point on, the modern-day border between South Africa and Botswana was agreed on, and the Afrikaners and Batswana traded and worked together comparatively peacefully. In 1884, Batawana, a northern-based Tswana clan's cavalry under

4440-596: The British colonised the area and declared a protectorate named Bechuanaland . As part of the decolonisation of Africa , Bechuanaland became an independent Commonwealth republic under its current name on 30 September 1966. Since then, it has been a parliamentary republic with a consistent record of uninterrupted democratic elections , although the Botswana Democratic Party was the only ruling party from independence until 2024. As of 2024, Botswana

4560-485: The Chobe District has the world's largest concentration of African elephants . The park covers about 11,000 km (4,247 sq mi) and supports about 350 species of birds. In Botswana forest cover is around 27% of the total land area, equivalent to 15,254,700 hectares (ha) of forest in 2020, down from 18,803,700 hectares (ha) in 1990. In 2020, naturally regenerating forest covered 15,254,700 hectares, of

4680-845: The Florissant Formation in North America and the Enspel Lagerstätte of Germany, dating to the late Eocene and late Oligocene respectively. Glossina is almost entirely restricted to wooded grasslands and forested areas of the Afrotropics . As of 1990, tsetse flies were reported from a maximum latitude of approximately 15° north in Senegal (Niayes Region), to a minimum of 28.5° south in South Africa (KwaZulu-Natal Province). Only two subspecies - G. f. fuscipes and G. m. submorsitans - are present in

4800-971: The German Empire and Britain coveted the territory of Botswana. During the Berlin Conference , Britain decided to annex Botswana to safeguard the Road to the North and thus connect the Cape Colony to its territories further north. It unilaterally annexed Tswana territories in January 1885 and then sent the Warren Expedition north to consolidate control over the area and convince the chiefs to accept British overrule. Despite their misgivings, they eventually acquiesced to this fait accompli . In 1890, areas north of 22 degrees were added to

4920-545: The Grand Duchy of Hesse , Hesse-Homburg , Nassau (all until 1866), Saxe-Coburg-Saalfeld , Saxe-Hildburghausen (both until 1826), and Reuss, elder line (until 1871). All of these bordered Prussia but not the main territory with sea access. There were no doubly landlocked countries from the unification of Germany in 1871 until the end of World War I . Liechtenstein bordered the Austro-Hungarian Empire , which had an Adriatic coastline, and Uzbekistan

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5040-486: The Kgabo Commission in 1991, factionalism and political rivalries have dominated Botswana politics. The Barata-Phathi faction was led by Peter Mmusi , Daniel Kwelagobe and Ponatshego Kedikilwe , while the A Team faction was led by Mompati Merafhe and Jacob Nkate. When Festus Mogae and Ian Khama became president and vice-president, respectively, they aligned with the A Team. Khama effectively expelled

5160-710: The Reduviidae , members of the Hemiptera . Animal trypanosomiasis , also called nagana when it occurs in bovine cattle or horses or sura when it occurs in domestic pigs , is caused by several trypanosome species. These diseases reduce the growth rate, milk productivity, and strength of farm animals, generally leading to the eventual death of the infected animals. Certain species of cattle are called trypanotolerant because they can survive and grow even when infected with trypanosomes although they also have lower productivity rates when infected. The course of

5280-529: The Russian Empire and then the Soviet Union , gained its independence with the dissolution of the latter in 1991 and became the second doubly landlocked country. However, Uzbekistan's doubly landlocked status depends on whether the Caspian Sea is considered a lake or a sea. In the latter case, Uzbekistan is not doubly landlocked, since its neighbors Turkmenistan and Kazakhstan have access to

5400-740: The Sound Dues , a toll needed to be paid to reach Western European waters. Sweden bypassed it by conquering Scania in 1658. Landlocked countries may be bordered by a single country having direct access to the high seas , two or more such countries, or be surrounded by other landlocked countries, making a country doubly landlocked. Three countries are landlocked by a single country ( enclaved countries): Seven landlocked countries are surrounded by only two mutually bordering neighbours (semi-enclaved countries): To this group could be added three landlocked territories, two of them de facto states with limited or no international recognition : A country

5520-594: The Umbrella for Democratic Change has operated as a coalition of opposition parties. The most recent election was held in 2024 , with the Botswana Democratic Party losing its majority for the first time in history, ending its 58 year rule over the country. The election saw Duma Boko being elected as president. In Botswana's early years, its politics were managed by President Seretse Khama and vice-president (later president) Quett Masire . Since

5640-592: The United Nations . The country's name means "Land of the Tswana ", referring to the dominant ethnic group in Botswana . The Constitution of Botswana recognizes a homogeneous Tswana state. The demonym Batswana was originally applied to the Tswana, and has also come to be used generally as a term for all citizens of Botswana. It is estimated that hominids lived in Botswana during the Pleistocene . Stone tools and animal remains indicate that all areas of

5760-670: The lymphatic system , then into the bloodstream, and eventually into the brain. The disease causes the swelling of the lymph glands, emaciation of the body, and eventually leads to death. Uninfected tsetse may bite the infected animal prior to its death and acquire the disease, thereby closing the transmission cycle. The tsetse-vectored trypanosomiases affect various vertebrate species including humans, antelopes, bovine cattle, camels, horses, sheep, goats, and pigs. These diseases are caused by several different trypanosome species that may also survive in wild animals such as crocodiles and monitor lizards. The diseases have different distributions across

5880-492: The 1920 establishment of two advisory councils to represent both Africans and Europeans. The African Council consisted of the eight heads of the Tswana tribes and some elected members. Proclamations in 1934 regulated tribal rule and powers. A European-African advisory council was formed in 1951, and the 1961 constitution established a consultative legislative council. In June 1964, the United Kingdom accepted proposals for

6000-700: The 45 landlocked countries, including those in Africa , Asia , and South America , have been classified as Landlocked Developing Countries (LLDCs) by the United Nations . Nine of the twelve countries with the lowest Human Development Index rankings are landlocked. International initiatives are aimed at reducing inequalities resulting from issues such as these, such as the United Nations Sustainable Development Goal 10 , which aims to reduce inequality substantially by 2030. In 1990, there were only 30 landlocked countries in

6120-486: The 7th to late 19th century, indicating it was occupied for more than 1,000 years. The hill was part of the formation of early states in southern Africa, with cattle as a major source of economy. The Toutswe settlement includes house-floors, large heaps of vitrified cow dung, and burials while the outstanding structure is the stone wall. Around 1000 AD, the Toutswe people moved into Botswana. However, agriculture also played

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6240-400: The A Team from the party in 2010 after he became president. A new rivalry formed in 2018 when Khama's chosen successor, Mokgweetsi Masisi, became president. He opposed Khama, and the two formed a political rivalry that continues to loom over Batswana politics in the 2020s. Botswana was ranked as a "flawed democracy" and 33rd out of 167 states in the 2023 Democracy Index (The Economist) , which

6360-469: The African continent, so are transmitted by different species. This table summarizes this information: Human African trypanosomiasis, also called sleeping sickness , is caused by trypanosomes of the species Trypanosoma brucei . This disease is invariably fatal if left untreated, but can almost always be cured with current medicines if the disease is diagnosed early enough. Sleeping sickness begins with

6480-594: The Batawana migrated into the Okavango Delta , probably in the 1790s. The first written records relating to modern-day Botswana appear in 1824. These records show that the Bangwaketse had become the predominant power in the region. Under the rule of Makaba II, the Bangwaketse kept vast herds of cattle in well-protected desert areas, and used their military prowess to raid their neighbours. Other chiefdoms in

6600-598: The Batswana—most of whom were driven into the desert or sanctuary areas such as hilltops and caves. Only after 1843, when the Amandebele moved into western Zimbabwe, did this threat subside. During the 1840s and 1850s, trade with Cape Colony -based merchants opened up and enabled the Batswana chiefdoms to rebuild. The Bakwena , Bangwaketse, Bangwato and Batawana cooperated to control the lucrative ivory trade and used

6720-583: The Caspian Sea is connected to the Black Sea via the man-made Volga–Don Canal , large oceangoing ships are unable to traverse it. Some countries or important parts of countries have coastlines or river ports reachable by oceangoing ships, but only through a strait or river part of the territory of another country. The other country can put restriction on passage. Between 1429 and 1857 Poland , Sweden , Russia and more Baltic countries were suffering from

6840-597: The Caspian Sea. Notes: The landlocked countries and de facto states can be grouped in contiguous groups as follows: Notes: There are the following 12 "single" landlocked countries (each of them borders no other landlocked country or de facto state): According to the United Nations geoscheme (excluding the de facto states ), Africa has the most landlocked countries, at 16, followed by Europe (14), Asia (12), and South America (2). However, if Armenia , Azerbaijan , Kazakhstan , and South Ossetia (de facto state) are counted as parts of Europe, then Europe has

6960-1232: The Department of Mines and Mineral Resources, Green Technology and Energy Security maintains data regarding mining throughout the country. Debswana , the largest diamond mining company operating in Botswana, is a joint venture, 50% owned by the government. Landlocked country Generally, being landlocked creates political and economic disadvantages that having access to international waters would avoid. For this reason, nations large and small throughout history have fought to gain access to open waters, even at great expense in wealth, bloodshed, and political capital . The economic disadvantages of being landlocked can be alleviated or aggravated depending on degree of development, surrounding trade routes and freedom of trade , commonality of language, and other considerations. Some landlocked countries in Europe are affluent, such as Andorra , Austria , Liechtenstein , Luxembourg , San Marino , Switzerland , and Vatican City , all of which, excluding Luxembourg (a founding member of NATO ), frequently employ neutrality in global political issues. However, 32 out of

7080-821: The Okavango Delta) are major tourist destinations. Other reserves include the Central Kalahari Game Reserve located in the Kalahari Desert in Ghanzi District ; Makgadikgadi Pans National Park and Nxai Pan National Park are in the Central District in the Makgadikgadi Pan . Botswana faces two major environmental problems, drought and desertification , which are heavily linked. Three-quarters of

7200-548: The United Kingdom's Westminster system and the Tswana people's tribal governments. Botswana has a centralised government in which national law supersedes local law. Local laws are developed by local councils and district councils. They are heavily influenced by tribal governments, which are led by the tribe's chief. The Parliament of Botswana consists of the President and the National Assembly , which serves as

7320-553: The area, by this time, had capitals of 10,000 or so and were fairly prosperous. This equilibrium came to end during the Mfecane period, 1823–1843, when a succession of invading peoples from South Africa entered the country. Although the Bangwaketse were able to defeat the invading Bakololo in 1826, over time, all the major chiefdoms in Botswana were attacked, weakened and impoverished. The Bakololo and AmaNdebele raided repeatedly and took large numbers of cattle, women and children from

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7440-697: The collapse of Mapungubwe. During this era, the first Tswana-speaking groups, the Bakgalagadi , moved into the southern areas of the Kalahari . These various peoples were connected to trade routes that ran via the Limpopo River to the Indian Ocean ; trade goods from Asia such as beads made their way to Botswana, most likely in exchange for ivory, gold and rhinoceros horn . The Toutswemogala Hill Iron Age settlement's radio-carbon dates range from

7560-652: The command of Kgosi Moremi, fought and defeated the Ndebele's invasion of northern Botswana at the Battle of Khutiyabasadi . This is the start of the collapse of the Ndebele Kingdom in Zimbabwe and it helped the Tswana speaking authority. Due to newly peaceful conditions, trade thrived between 1860 and 1880. Christian missionaries were able to take advantage of this. The Lutherans and the London Missionary Society both became established in

7680-463: The country by 1856. By 1880, every major village had a resident missionary, and their influence slowly grew. Khama III (reigned 1875–1923) was the first of the Tswana chiefs to make Christianity a state religion, and a great deal of Tswana customary law changed as a result. Christianity became the de facto official religion in all the chiefdoms by World War I . During the Scramble for Africa , both

7800-419: The country were inhabited at least 400,000 years ago. It was claimed to have been the birthplace of all modern humans from around 200,000 years ago. Evidence left by modern humans, such as cave paintings, is about 73,000 years old. The earliest known inhabitants of southern Africa are thought to have been the forebears of present-day San ("Bushmen") and Khoi peoples. Both groups speak click languages from

7920-403: The country's human and animal populations depend on groundwater due to drought. Groundwater use through deep borehole drilling has somewhat eased the effects of drought. Surface water is scarce in Botswana, and less than 5% of the agriculture in the country is sustainable by rainfall. In the remaining 95% of the country, raising livestock is the primary source of rural income. Approximately 71% of

8040-414: The country's land is used for communal grazing, which has been a major cause of the desertification and the accelerating soil erosion in the country. Since raising livestock has been profitable for the people of Botswana, they continue to exploit the land with dramatically increasing numbers of animals. From 1966 to 1991, the livestock population grew from 1.7   million to 5.5   million. Similarly,

8160-601: The country. According to the International Monetary Fund , economic growth averaged over 9% per year from 1966 to 1999. Botswana has a high level of economic freedom compared to other African countries. The government has maintained a sound fiscal policy , despite consecutive budget deficits in 2002 and 2003, and a negligible level of foreign debt . It earned the highest sovereign credit rating in Africa and has stockpiled foreign exchange reserves (over $ 7 billion in 2005/2006) amounting to almost two and

8280-422: The country. The Notwane provides water to the capital through the Gaborone Dam . The Chobe River meets with the Zambezi River at a place called Kazungula . Botswana has diverse areas of wildlife habitat. In addition to the delta and desert areas, there are grasslands and savannas . Northern Botswana has one of the few remaining large populations of the endangered African wild dog . Chobe National Park in

8400-403: The current responsible Undersecretary-General is Anwarul Karim Chowdhury . Some countries have a long coastline, but much of it may not be readily usable for trade and commerce. For instance, in its early history, Russia 's only ports were on the Arctic Ocean and frozen shut for much of the year. The wish to gain control of a warm-water port was a major motivator of Russian expansion towards

8520-420: The disease, has often been subdivided into three subspecies that were identified based either on the vertebrate hosts which the strain could infect or on the virulence of the disease in humans. The trypanosomes infectious to animals and not to humans were named Trypanosoma brucei brucei . Strains that infected humans were divided into two subspecies based on their different virulences: Trypanosoma brucei gambiense

8640-494: The egg; following egg development and birth is the three larval stages, a pupal stage, and the emergence and maturation of the adult. Overall Suidae are the most important hosts. Waterbuck ( Kobus ellipsiprymnus ) are unmolested by Glossina because they produce volatiles which act as repellents . Waterbuck odor volatiles are under testing and development as repellents to protect livestock . By species, bloodmeals are derived from: The genome of Glossina morsitans

8760-538: The elucidation of the strains of T. brucei infective to humans requires a more complex explanation. Procyclins are proteins developed in the surface coating of trypanosomes whilst in their tsetse fly vector. Other forms of human trypanosomiasis also exist but are not transmitted by tsetse. The most notable is American trypanosomiasis, known as Chagas disease , which occurs in South America , caused by Trypanosoma cruzi , and transmitted by certain insects of

8880-454: The end of each pregnancy . The article " Parasitic flies of domestic animals " has a diagram of the anatomy of dipteran flies. Most tsetse flies are, physically, very tough. Houseflies, and even horseflies, are easily killed with a flyswatter, for example; a great deal of effort is needed to crush a tsetse fly. Tsetse has an unusual life cycle , which may be due to the richness of their blood food source. A female fertilizes only one egg at

9000-399: The end of the pupal stage, tsetse emerges as adult flies. The adults are relatively large flies, with lengths of 0.5–1.5 centimetres ( 1 ⁄ 4 – 5 ⁄ 8  in), and have a recognizable shape, or bauplan , which makes them easy to distinguish from other flies. Tsetse have large heads, distinctly separated eyes, and unusual antennae. The thorax is quite large, while the abdomen

9120-410: The feeding time, promoting trypanosome transmission to the vertebrate host. These trypanosomes are highly evolved and have developed a life cycle that requires periods in both the vertebrate and tsetse hosts. Tsetse transmit trypanosomes in two ways, mechanical and biological transmission. The relative importance of these two modes of transmission for the propagation of tsetse-vectored trypanosomiases

9240-437: The first president), who had been serving as Mogae's vice-president since resigning his position as Commander of the Botswana Defence Force in 1998 to take up this civilian role. On 1 April 2018, Mokgweetsi Eric Keabetswe Masisi was sworn in as the fifth president of Botswana, succeeding Ian Khama. He represents the Botswana Democratic Party , which has also won a majority in every parliamentary election since independence. All

9360-417: The flies can be raised in a laboratory, and because they are relatively large, facilitating their analysis. Tsetse flies can be seen as independent individuals in three forms: as third- instar larvae, pupae, and adults. Tsetse first becomes separate from their mothers during the third larval instar, during which they have the typical appearance of maggots . However, this life stage is short, lasting at most

9480-458: The fly's head. This may be the reason/a reason for the behavioral changes seen, especially the unnecessarily increased feeding frequency, which increases transmission opportunities. This may be due in part to the altered glucose metabolism observed, causing a perceived need for more calories. (The metabolic change, in turn, being due to complete absence of glucose-6-phosphate 1-dehydrogenase in infected flies.) Monoamine neurotransmitter synthesis

9600-472: The former landlocked country of Czechoslovakia ceased to exist on 1 January 1993. On 30 September 2022, the Luhansk People's Republic (de facto state) was annexed by Russia and ceased to exist as a landlocked de facto state. On 19 September 2023, Azerbaijan launched a new offensive against the Republic of Artsakh (de facto state) and achieved a decisive victory. The Government of Artsakh

9720-641: The fourth-largest in Africa) gives the country a relatively high standard of living and the second-highest Human Development Index of continental Sub-Saharan Africa (after South Africa). Despite this, Botswana continues to grapple with high unemployment rates. Botswana is a member of the Southern African Customs Union , the Southern African Development Community , the Commonwealth of Nations and

9840-479: The genus Trypanosoma . These organisms are about the size of red blood cells. Different species of trypanosomes infect different hosts. They range widely in their effects on the vertebrate hosts. Some species, such as T. theileri , do not seem to cause any health problems except perhaps in animals that are already sick. Some strains are much more virulent . Infected flies have an altered salivary composition which lowers feeding efficiency and consequently increases

9960-573: The government denies that there is any link to mining, claiming the relocation is to preserve the wildlife and ecosystem, even though the San people have lived sustainably on the land for millennia. On the reservations, they struggle to find employment, and alcoholism is rampant. On 24 August 2018, the UN Special Rapporteur on Minorities , Fernand de Varennes, issued a statement calling on Botswana "to step up efforts to recognise and protect

10080-415: The ground and develop a hard outer shell (called the puparial case ), within which they complete their morphological transformations into adult flies. The larval life stage has a variable duration, generally 20 to 30 days, and the larvae must rely on stored resources during this time. The importance of the richness and quality of blood to this stage can be seen; all tsetse development (prior to emerging from

10200-547: The human population has increased from 574,000 in 1971 to 1.5   million in 1995, a 161% increase in 24   years. Environmentalists report that the Okavango Delta is drying up due to increased livestock grazing. The Okavango Delta is one of the major semi-forested wetlands in Botswana and one of the largest inland deltas in the world; the ecosystem is crucial to the survival of many animals. The Department of Forestry and Range Resources has already begun to implement

10320-418: The land. The country had a 2018 Forest Landscape Integrity Index mean score of 9.13/10, ranking it 8th globally out of 172 countries. The United Nations Development Programme (UNDP) claims that poverty is a major problem behind the overexploitation of resources, including land, in Botswana. The UNDP joined in with a project started in the southern community of Struizendam in Botswana. The project's purpose

10440-562: The largest single foreign contributor to the development of the BDF , and a large segment of its officer corps have received U.S. training. The Botswana government gave the United States permission to explore the possibility of establishing an Africa Command ( AFRICOM ) base in the country. Botswana is the 50th most peaceful country in the world, according to the 2024 Global Peace Index . The Botswana Centre for Human Rights , Ditshwanelo,

10560-656: The legislature has little power to check the president once appointed. The judiciary includes the High Court of Botswana , the Court of Appeal and Magistrates' Courts. Cases are often settled by customary courts with tribal chiefs presiding. Elections in Botswana are held every five years and overseen by the Independent Electoral Commission (IEC). Botswana operates a multi-party system in which many political parties compete in elections. It

10680-495: The lobes of the crop of G. m. centralis and G. m. morsitans . Tsetse are in the order Diptera , the true flies. They belong to the superfamily Hippoboscoidea, in which the tsetse's family, the Glossinidae, is one of four families of blood-feeding obligate parasites. Up to 34 species and subspecies of tsetse flies are recognized, depending on the particular classification used. All current classifications place all

10800-708: The most landlocked countries, at 20 (including all three landlocked de facto states). If these transcontinental or culturally European countries are included in Asia, then both Africa and Europe (including Kosovo and Transnistria ) have the most, at 16. Depending on the status of Kazakhstan and the South Caucasian countries, Asia has between 9 and 13 (including South Ossetia ). South America only has two landlocked countries: Bolivia and Paraguay . Australia and North America have no landlocked countries, while Antarctica has no countries at all. Oceania (which

10920-563: The nation's formal and sole legislature, while the Ntlo ya Dikgosi serves an advisory body made up of tribal chiefs and other appointed members. Botswana's executive branch is led by the President of Botswana , who serves as both the head of state and head of government . The members of parliament choose the president, and the president then appoints the vice-president and cabinet members. The president has significant power in Botswana, and

11040-401: The naturally regenerating forest 0% was reported to be primary forest (consisting of native tree species with no clearly visible indications of human activity) and around 11% of the forest area was found within protected areas. For the year 2015, 24% of the forest area was reported to be under public ownership and 76% private ownership . The Chobe National Park and Moremi Game Reserve (in

11160-666: The new Bechuanaland Protectorate. During the 1890s, the new territory was divided into eight different reserves, with fairly small amounts of land being left as freehold for white settlers . During the early 1890s, the British government decided to hand over the Bechuanaland Protectorate to the British South Africa Company . This plan, which was well on its way to fruition despite the entreaties of Tswana leaders who toured England in protest,

11280-575: The other); Secondly, tsetse also have a long proboscis , extending directly forward, which is attached by a distinct bulb to the bottom of their heads. Fossilized tsetse has been recovered from Paleogene -aged rocks in the United States and Germany. Twenty-three extant species of tsetse flies are known from the African continent as well as the Arabian Peninsula. Tsetse without the "fly" has become more common in English , particularly in

11400-553: The previous presidents have also represented the same party. A long-running dispute over the northern border with Namibia 's Caprivi Strip was the subject of a ruling by the International Court of Justice in December 1999. It ruled that Kasikili Island belongs to Botswana. Following the independence in 1966, the president always represented the same political party until the 2024 Botswana general election , which

11520-708: The proceeds to import horses and guns, which in turn enabled them to establish control over what is now Botswana. This process was largely complete by 1880, and the Batswana subjugated thus the Bushmen, Kalanga, Bakgalagadi and other current minorities. Following the Great Trek , Afrikaners from the Cape Colony established themselves on the borders of Botswana in the Transvaal . In 1852, a coalition of Tswana chiefdoms led by Sechele I defeated Afrikaner incursions at

11640-526: The programme is giving the government information to help with policy development. Botswana is a parliamentary republic governed by the Constitution of Botswana . It is the longest uninterrupted democracy in Africa. Its seat of government is in Gaborone . Botswana's governing institutions were established after it became an independent nation in 1966. Botswana's governmental structure is based on both

11760-475: The puparial case as a full adult) occurs without feeding, with only the nutrition provided by the mother fly. She must get enough energy for her own survival (in addition to the needs of her developing offspring), as well as for the stored resources that her offspring will require until they emerge as adults. Technically, these insects undergo the standard development process of insects, beginning with oocyte formation, ovulation, fertilization, and development of

11880-473: The ratio is 3:1). Historically, traveling between a landlocked country and a country which did not border said country required the traveler to pass border controls twice or more. In recent times the advent of air travel has largely negated this impediment. Countries have acted to overcome being landlocked by acquiring land that reaches the sea: Countries can make agreements on getting free transport of goods through neighbouring countries: Losing access to

12000-586: The rights of minorities in relation to public services, land and resource use, and the use of minority languages in education and other critical areas." Botswana is divided into 10 administrative districts , 2 city districts, 4 towns, 11 sub-districts and, in total, 16 administrative divisions. They are: These are administered by 16 local authorities (district councils, city councils or town councils). In 1977, Botswana's administrative divisions were Ngamiland, Chobe, Francistown, Ngwato, Tuli, Ghanzi, Kgalagadi, Ngwaketse, Kweneng, Gaborone and Lobatse. In 2006, Chobe

12120-559: The scientific and development communities. The word is pronounced tseh-tseh in the Sotho languages and is easily rendered in other African languages. During World War II, a British de Havilland antisubmarine aircraft was known as the Tsetse Mosquito . The biology of tsetse is relatively well understood by entomologists . They have been extensively studied because of their medical, veterinary, and economic importance, because

12240-520: The sea is generally a great loss to a nation, politically, militarily, and economically. The following are examples of countries becoming landlocked. The United Nations Convention on the Law of the Sea now gives a landlocked country a right of access to and from the sea without taxation of traffic through transit states. The United Nations has a programme of action to assist landlocked developing countries, and

12360-430: The small Khoe-Kwadi , Kx’a and Tuu language families whose members hunted, gathered and traded over long distances. When cattle were first introduced into southern Africa about 2,000 years ago, pastoralism became a major feature of the economy since the region had large grasslands free of tsetse flies . It is unclear when Bantu -speaking peoples first moved into the country from the north, although AD 600 seems to be

12480-670: The species in the genus Glossina , which are placed in their own family, Glossinidae . The tsetse is an obligate parasite , which lives by feeding on the blood of vertebrate animals. Tsetse has been extensively studied because of their role in transmitting disease. They have pronounced economic and public health impacts in sub-Saharan Africa as the biological vectors of trypanosomes , causing human and animal trypanosomiasis . Tsetse can be distinguished from other large flies by two easily-observed features: Primarily, tsetse fold their wings over their abdomens completely when they are resting (so that one wing rests directly on top of

12600-550: The territories transferred to their jurisdiction, the UK kept delaying; consequently, it never occurred. The election of the Nationalist government in 1948, which instituted apartheid , and South Africa's withdrawal from the Commonwealth in 1961, ended any prospect of the UK or these territories agreeing to incorporation into South Africa. An expansion of British central authority and the evolution of native government resulted in

12720-777: The tsetse species in a single genus named Glossina . Most classifications place this genus as the sole member of the family Glossinidae. The Glossinidae are generally placed within the superfamily Hippoboscoidea , which contains other hematophagous families. The tsetse genus is generally split into three groups of species based on a combination of distributional, ecological, behavioral, molecular and morphological characteristics. The genus includes; savannah flies, forest flies and riverine and lacustrine flies. The 'savannah' flies: ( Morsitans group , subgenus Glossina s.s.): The 'forest' flies: ( Fusca group, subgenus Austenina ): The 'riverine' and 'lacustrine' flies: ( Palpalis group, subgenus Nemorhina ): Fossil glossinids are known from

12840-520: The very southwest of Saudi Arabia. Although Carter found G. tachiniodes in 1903 nearby, near Aden in southern Yemen, there have been no confirmations since. Tsetse are biological vectors of trypanosomes , meaning that in the process of feeding, they acquire and then transmit small, single-celled trypanosomes from infected vertebrate hosts to uninfected animals. Some tsetse-transmitted trypanosome species cause trypanosomiasis , an infectious disease. In humans, tsetse transmitted trypanosomiasis

12960-429: The world. However, the dissolutions of the Soviet Union and Czechoslovakia ; the breakup of Yugoslavia ; the independence referendums of South Ossetia (de facto state), Eritrea , Montenegro , South Sudan , and the Luhansk People's Republic (de facto state); and the unilateral declaration of independence of Kosovo (de facto state) created 15 new landlocked countries and five landlocked de facto states while

13080-431: The world; if you are landlocked, you serve your neighbors." Others have argued that being landlocked has an advantage as it creates a "natural tariff barrier" that protects the country from cheap imports. In some instances, this has led to more robust local food systems. Landlocked developing countries have significantly higher costs of international cargo transportation compared to coastal developing countries (in Asia

13200-429: Was a dominant-party state in which the Botswana Democratic Party had ruled with a majority government from independence in 1966 to 2024. The nation's elections are recognized as free and fair , but the ruling party has institutional advantages that other parties do not. Factionalism is common within Botswana's political parties, and several groups have formed new parties by splitting from established ones. Since 2019,

13320-483: Was established in 1993. Until June 2019, homosexual acts were illegal in Botswana. A Botswana High Court decision of 11 June of that year struck down provisions in the Criminal Code that punished "carnal knowledge of any person against the order of nature" and "acts of gross indecency", making Botswana one of 22 African countries that have either decriminalised or legalised homosexual acts. Capital punishment

13440-721: Was eventually foiled by the failure of the Jameson Raid in January 1896. When the Union of South Africa was formed from the main British colonies in the region in 1910, the High Commission Territories—the Bechuanaland Protectorate, Basutoland (now Lesotho ) and Swaziland (now Eswatini )—were not included, but provision was made for their later incorporation. However, the UK began to consult with their inhabitants as to their wishes. Although successive South African governments sought to have

13560-572: Was formed in 1977. The president is commander-in-chief of the armed forces and appoints a defence council. In 2019, Botswana signed the UN treaty on the Prohibition of Nuclear Weapons . Following political changes in South Africa and the region, the BDF's missions have increasingly focused on preventing poaching , preparing for disasters , and supporting foreign peacekeeping . The United States has been

13680-464: Was historic in that the opposition party won. Duma Boko became the first president who did not represent the Botswana Democratic Party (BDP). On 1 November 2024, Duma Boko, leader of opposition Umbrella for Democratic Change (UDC), was sworn in as president of Botswana. At 581,730 km (224,607 sq mi), Botswana is the world's 48th-largest country. It also has a mean altitude of roughly 1,000 metres (3,300 ft) above sea level. Botswana

13800-465: Was officially dissolved on 1 January 2024. As a result, Artsakh ceased to exist as a landlocked de facto state and the Nagorno-Karabakh region was reintegrated into Azerbaijan. As of 1 April 2024, there were 44 landlocked countries and three landlocked de facto states ( Kosovo , South Ossetia , and Transnistria ) in the world. Historically, being landlocked has been disadvantageous to

13920-431: Was removed from being an administrative division, and Ngamiland's name was changed to North West district. Chobe was readded on 31 March 2014. That same day, the administrative divisions Francistown, Gaborone, Jwaneng, Lobatse, Selibe Phikwe, and Sowa Town were also added. Since independence, Botswana has had one of the fastest growth rates in per capita income in the world. Formerly one of the world's poorest countries—with

14040-413: Was sequenced in 2014. Tsetse flies have at least three bacterial symbionts . The primary symbiont is Wigglesworthia ( Wigglesworthia glossinidia ), which live within the fly's bacteriocytes . The second symbiont is Sodalis ( Sodalis glossinidius ) intercellularly or intracellularly, and the third is some kind of Wolbachia . The salivary gland hypertrophy virus causes abnormal bleeding in

14160-414: Was the second-highest rating in Africa, and the highest ranking in continental Africa (only the offshore island nation of Mauritius bested its ranking). However, according to the 2024 V-Dem Democracy Indices , Botswana has been experiencing an episode of democratic backsliding over the past 10 years, recording its lowest ever score on the indices. The indices classify Botswana as an electoral democracy in

14280-645: Was then part of the Russian Empire , which had both ocean and sea access. With the dissolution of Austria-Hungary in 1918 and creation of an independent, landlocked Austria , Liechtenstein became the sole doubly landlocked country until 1938. In the Anschluss that year, Austria was absorbed into Nazi Germany , which possessed a border on the Baltic Sea and the North Sea . After World War II, Austria regained its independence and Liechtenstein once again became doubly landlocked. Uzbekistan, which had been part of

14400-428: Was thought to have a slower onset and Trypanosoma brucei rhodesiense refers to strains with a more rapid, virulent onset. This characterization has always been problematic but was the best that could be done given the knowledge of the time and the tools available for identification. A recent molecular study using restriction fragment length polymorphism analysis suggests that the three subspecies are polyphyletic , so

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