The Floian is the second stage of the Ordovician Period . It succeeds the Tremadocian with which it forms the Lower Ordovician series. It precedes the Dapingian Stage of the Middle Ordovician . The Floian extended from 477.7 to 470 million years ago. The lower boundary is defined as the first appearance of the graptolite species Tetragraptus approximatus .
21-797: The base of this stage was ratified by the International Commission on Stratigraphy (ICS) in 2002. The Floian Stage is named after Flo , a village in Västergötland , southern Sweden . The name "Floan" was proposed in 2004, but the ICS adapted Floian as the official name of the stage. The GSSP of the Floian is the lower Tøyen Shale in Diabasbrottet Quarry ( 58°21′32″N 12°30′09″E / 58.3589°N 12.5024°E / 58.3589; 12.5024 ) which
42-436: A sensory function. In all the extant genera , the ambulacral plates are fused in pairs and form joints in the arms, known as vertebrae. In some extinct genera, they are not fused in this way. The ambulacral grooves function as internal epineural canals. All members of the order Somasteroidea (or Stomasteroidea) are extinct. Their first appearance in the fossil record was in the early Ordovician and they had probably died out by
63-540: A subphylum in the phylum Echinodermata , within the Eleutherozoa . Characteristics include a star-shaped body and radially divergent axes of symmetry. The subphylum includes the class Asteroidea (the starfish ), the class Ophiuroidea (the brittle stars and basket stars ), and the extinct taxa Somasteroidea and Stenuroidea . Originally proposed in the late 1800s, Asterozoa was not used in F.A. Bather 's two-subphylum echinoderm taxonomy in 1900. However, it
84-480: A symmetry based on a number other than five. The arms have very broad bases and their skeletal support is provided by the calcareous plates called ossicles or ambulacral plates in the body wall. These are joined with muscular and connective tissue giving flexibility. The mouth is on the lower or oral surface, with a corresponding anus on the upper or aboral surface; though the anus is absent in Paxillosida , wherein
105-574: Is an outcrop of a shale-dominated stratigraphic succession. The lower boundary of the Floian is defined as the first appearance of Tetragraptus approximatus which is about 2.1 above the Cambrian strata. Radiometric dating has set the Tremadocian -Floian boundary at 477.7 million years ago. The upper boundary which is also the base of the Dapingian stage is defined as the first appearance of
126-646: The International Stratigraphic Chart , a combined working proposal and guideline-to-date released after the last ICS deliberations prior to the upcoming (next) meeting of the IUGS. Until the IUGS accepts the recommendations, they are unofficial since the IUGS parent approves or dismisses the individual deliberation reports of the ICS, which are presented as recommendations, and span dating and strata selection criteria, and related issues including nomenclatures. In de facto everyday matters,
147-817: The Neogene System and Neogene Period. Despite the strong debate, the Quaternary saw official ratification as a geological unit from the IUGS in June 2009, placing its lower boundary to the Gelasian Stage/Age at Monte San Nicola, Sicily, Italy (until then uppermost part of the Pliocene Series/Epoch, and thus of the Neogene System/Period), 2.58 Ma BP . In addition to publishing paper and document (PDF) versions of
168-859: The conodont species Baltoniodus triangularis at the GSSP in the Huanghuachang Section, Hubei Province, China . Partial analogues of Floian stage in Baltoscandia are Hunneberg stage (lower) and Billingen stage (upper). On the Siberian Platform , Ugorian stage corresponds to Floian. The global Billingen Transgressive Event occurred in the Early Floian age. Black graptolitic argillites of Gorny Altai as well as conglomerates and gritstones of Salair , Russia , possibly correlates with this event. Discovered in
189-556: The paleontology , geology , geobiology and chronostratigraphy fields, among others. The International Commission on Stratigraphy has spawned numerous subcommittee level organizations organized and mobilized on a local country-wide or regional basis that are the true working committees of the IUGS, and these do the field work, basis comparisons in conference or co-ordination research committee meetings of local or wide-scale scope. The ICS publishes various reports and findings as well as revised references periodically, summarized in
210-496: The " International Stratigraphic Commission ", is a daughter or major subcommittee grade scientific daughter organization that concerns itself with stratigraphical , geological , and geochronological matters on a global scale. It is the largest subordinate body of the International Union of Geological Sciences (IUGS). The ICS is essentially a permanent working subcommittee , which meets far more regularly than
231-472: The Floian age. Two species of Paroistodus are known from the Floian deposits of Baltoscandia and South China. Several thousand chemically isolated graptolite specimens including genera Baltograptus and Pseudophyllograptus were collected from the upper Floian sediments of Skattungbyn , Dalarna , central Sweden . Presented mostly by juveniles and isolated siculae , these graptolites inhabited primarily in shallow water environment. Trilobites of
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#1732772220980252-556: The Floian strata of Newfoundland , coral-like fossils of Reptamsassia divergens and Reptamsassia minuta are the oldest example of symbiotic intergrowth of modular species. This allows to judge the level of development of reef ecosystems of the Early Ordovician. Conodonts Serratognathus , Prioniodus and Oepikodus were distributed in Kazakhstan , Korea , China , Indochina and Australasia during
273-718: The International Stratigraphic Chart, the ICS also provides a machine-readable version of the chart formulated using the Web Ontology Language (OWL) and, in particular, Time Ontology in OWL . The ICS' chart web page also provides an interactive version of the chart, based on the OWL data. The logo of International Commission on Stratigraphy was designed after the Chinese character of "mountain", 山. Asterozoa The Asterozoa are
294-502: The deliberative results reported out of any meetings of the ICS are widely accepted and immediately enter everyday use, except in the rare cases where they result in a strong body of dissenting opinion, which matters are resolved before the full IUGS. One such controversy arose in 2009 when the ICS deliberated and decided that the Pliocene Series of the current but unofficially named Quaternary Period should be shifted into
315-598: The fossil record. (i.e. section of the rock record as of a core sample section or accessible exposed strata, which when a core sample are usually "trayed" in long pieces, also called "sections" about a meter in length.) Additionally the ICS defines an alternative type of benchmark and criteria called Global Standard Stratigraphic Ages (GSSAs) where the characteristics and dating criteria set solely by physical sciences methods (such as magnetic alignment sequences, radiological criteria, etcetera.) as well as encouraging an international and open debate amongst Earth scientists in
336-586: The genera Tsaidamaspis , Zhiyia and Liexiaspis were found in the Floian part of the Duoquanshan Formation , northwest China. Falloaster anquiroisitus , an asterozoan of problematic classification, is known from the Floian Garden City Formation of Idaho , USA . International Commission on Stratigraphy The International Commission on Stratigraphy ( ICS ), sometimes unofficially referred to as
357-430: The late Devonian. They are similar to the asteroids in that their bodies are flattened dorsoventrally and they have five petaloid arms with broad bases. The ambulacral plates in somasteroids are simple and unspecialized, and the arms were thought to be not flexible and were unable to assist in feeding, but the oral mouth parts were more complex. An example for Somasteroidea is Villebrunaster fezouataensis . Stenuroidea
378-529: The mouth serves also as an anus, otherwise leading to a blind gut, similar to their Ophiuroid relatives. Their large coelomic cavity can extend from the disc into the arms. The paired gonads are also located in the arms and release gametes via the gonoducts. The lateral flexion of the arms is limited by how the ambulacral plates are arranged. On the oral surface of the disc and radiating arms are four rows of ambulacral plates which are associated with two or four rows of tube feet, which are used in locomotion. On
399-442: The quadrennial meetings scheduled by the IUGS, when it meets as a congress or membership of the whole . One of its main aims, a project begun in 1974, is to establish a multidisciplinary standard and global geologic time scale that will ease paleontological and geobiological comparisons region to region by benchmarks with stringent and rigorous strata criteria called Global Boundary Stratotype Section and Points (GSSPs) within
420-429: The upper or aboral surface of the disc is an opening, the madreporite . This is connected to the water vascular system which assists in respiration and provides hydraulic pressure for the tube feet . Members of the class Ophiuroidea are characterised by having the gut and internal organs confined to the central disc. The arms are clearly separated from the disc. The tube feet do not have adhesive properties, but serve
441-455: Was adopted as part of a four-subphylum taxonomy by the Treatise on Invertebrate Paleontology in 1966. Asterozoa is a monophyletic clade ; its sister group is Echinozoa . Members of the class Asteroidea are characterised by a star-shaped body plan consisting of a central disc and multiple radiating arms. They usually exhibit pentamerous radial symmetry, but some species typically have
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