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120-517: Many Khoe-Sān peoples are the descendants of a very early dispersal of anatomically modern humans to Southern Africa before 150,000 years ago. (However, see below for recent work supporting a multi-regional hypothesis that suggests the Khoi-San may be a source population for anatomically modern humans.) Their languages show a vague typological similarity, largely confined to the prevalence of click consonants . They are not verifiably derived from

240-1045: A 2017 study to be between 350 and 260,000 years ago, compatible with the estimated age of early H. sapiens . The study states that the deep split-time estimation of 350 to 260 thousand years ago is consistent with the archaeological estimate for the onset of the Middle Stone Age across sub-Saharan Africa and coincides with archaic H. sapiens in southern Africa represented by, for example, the Florisbad skull dating to 259 (± 35) thousand years ago. H. s. idaltu , found at Middle Awash in Ethiopia, lived about 160,000 years ago, and H. sapiens lived at Omo Kibish in Ethiopia about 233,000-195,000 years ago. Two fossils from Guomde, Kenya, dated to at least (and likely more than) 180,000 years ago and (more precisely) to 300–270,000 years ago, have been tentatively assigned to H. sapiens and similarities have been noted between them and

360-555: A common proto-language, but are today split into at least three separate and unrelated language families ( Khoe-Kwadi , Tuu and Kxʼa ). It has been suggested that the Khoekhoe may represent Late Stone Age arrivals to Southern Africa, displaced by Bantu expansion reaching the area roughly around 1,500AD . Sān are popularly thought of as foragers in the entire southern Africa and regions of Botswana, Namibia, Angola, Zambia, Zimbabwe, Lesotho and Northern South Africa. The word sān

480-463: A comparatively homogeneous single species of H. sapiens from more diverse varieties of archaic humans (all of which were descended from the early dispersal of H. erectus some 1.8 million years ago) was debated in terms of two competing models during the 1980s: " recent African origin " postulated the emergence of H. sapiens from a single source population in Africa, which expanded and led to

600-468: A convention popular in the 1990s, Neanderthals were classified as a subspecies of H. sapiens , as H. s. neanderthalensis , while AMH (or European early modern humans , EEMH) was taken to refer to " Cro-Magnon " or H. s. sapiens . Under this nomenclature (Neanderthals considered H. sapiens ), the term "anatomically modern Homo sapiens " (AMHS) has also been used to refer to EEMH ("Cro-Magnons"). It has since become more common to designate Neanderthals as

720-406: A higher forehead, and reduced brow ridge . Early modern people and some living people do however have quite pronounced brow ridges, but they differ from those of archaic forms by having both a supraorbital foramen or notch, forming a groove through the ridge above each eye. This splits the ridge into a central part and two distal parts. In current humans, often only the central section of the ridge

840-489: A huge group of languages spread throughout Western, Central and Southern Africa. The Benue–Congo branch includes the Bantu languages, which are found throughout Central, Southern, and Eastern Africa. A characteristic feature of most Atlantic–Congo languages, including almost all the Bantu languages except Swahili, Sotho-Tswana and Nguni languages, is their use of tone. They generally lack case inflection , but grammatical gender

960-419: A linguistic phylum by Joseph Greenberg in 1955. Their genetic relationship was questioned later in the 20th century, and the term now serves mostly as a convenience term without implying genetic unity, much like " Papuan " and " Australian " are. Their most notable uniting feature is their click consonants . They are categorized in two families, and a number of possible language isolates. The Kxʼa family

1080-564: A major new population center near the Great Lakes of East Africa, where a rich environment supported a dense population. The Urewe culture dominated the Great Lakes region between 650BC and 550BC. It was one of Africa's oldest iron-smelting centres. By the first century BC, Bantu speaking communities in the great lakes region developed iron forging techniques that enabled them to produce carbon steel . Movements by small groups to

1200-475: A more complex intermixing could have taken place. Further east, Bantu-speaking communities had reached the great Central African rainforest, and by 500   BC, pioneering groups had emerged into the savannas to the south, in what are now the Democratic Republic of Congo , Angola , and Zambia . Another stream of migration, having moved east by 3,000 years ago (1000   BC), was creating

1320-470: A number of other species (by some authors considered subspecies of either H. sapiens or H. erectus ). The divergence of the lineage leading to H. sapiens out of ancestral H. erectus (or an intermediate species such as Homo antecessor ) is estimated to have occurred in Africa roughly 500,000 years ago. The earliest fossil evidence of early modern humans appears in Africa around 300,000 years ago, with

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1440-503: A number of physiological details which can be taken as reliably differentiating the physiology of Neanderthals vs. anatomically modern humans. The term "anatomically modern humans" (AMH) is used with varying scope depending on context, to distinguish "anatomically modern" Homo sapiens from archaic humans such as Neanderthals and Middle and Lower Paleolithic hominins with transitional features intermediate between H. erectus , Neanderthals and early AMH called archaic Homo sapiens . In

1560-465: A self-designation after the end of apartheid in the late 1990s. Since the 2010s, there has been a "Khoisan activist" movement, demanding recognition and land rights from the government and white minority which owns large parts of the country's private land. It is suggested that the ancestors of the modern Khoisan expanded to southern Africa (from East or Central Africa ) before 150,000 years ago, possibly as early as before 260,000 years ago, so that by

1680-486: A separate species, H. neanderthalensis , so that AMH in the European context refers to H. sapiens , but the question is by no means resolved. In this more narrow definition of H. sapiens , the subspecies Homo sapiens idaltu , discovered in 2003, also falls under the umbrella of "anatomically modern". The recognition of H. sapiens idaltu as a valid subspecies of the anatomically modern human lineage would justify

1800-508: A smaller, more receded dentary, making the rest of the jaw-line stand out, giving an often quite prominent chin. The central part of the mandible forming the chin carries a triangularly shaped area forming the apex of the chin called the mental trigon , not found in archaic humans. Particularly in living populations, the use of fire and tools requires fewer jaw muscles, giving slender, more gracile jaws. Compared to archaic people, modern humans have smaller, lower faces. The body skeletons of even

1920-576: A study published in 2020, there are indications that 2% to 19% (or about ≃6.6 and ≃7.0%) of the DNA of four West African populations may have come from an unknown archaic hominin which split from the ancestor of humans and Neanderthals between 360 kya to 1.02 mya. Generally, modern humans are more lightly built (or more "gracile") than the more "robust" archaic humans . Nevertheless, contemporary humans exhibit high variability in many physiological traits , and may exhibit remarkable "robustness". There are still

2040-498: A virtual skull shape of the last common human ancestor to modern humans/ H. sapiens , representative of the earliest modern humans, and suggested that modern humans arose between 350,000 and 260,000 years ago through a merging of populations in East and South Africa while North-African fossils may represent a population which introgressed into Neandertals during the LMP. According to

2160-638: Is associated with the African megadroughts during MIS 5 , beginning 130,000 years ago. A 2011 study located the origin of basal population of contemporary human populations at 130,000 years ago, with the Khoi-San representing an "ancestral population cluster" located in southwestern Africa (near the coastal border of Namibia and Angola ). While early modern human expansion in Sub-Saharan Africa before 130 kya persisted, early expansion to North Africa and Asia appears to have mostly disappeared by

2280-546: Is characteristic, with some languages having two dozen genders ( noun classes ). The root of the verb tends to remain unchanged, with either particles or auxiliary verbs expressing tenses and moods. For example, in a number of languages the infinitival is the auxiliary designating the future. Before the expansion of Bantu-speaking farmers, Central, Southern, and Southeast Africa were likely populated by Pygmy foragers, Khoisan -speaking hunter-gatherers , Nilo-Saharan -speaking herders, and Cushitic -speaking pastoralists . It

2400-484: Is clear that there were human populations in the region at the time of the expansion, and pygmies are their closest living relatives. However, mtDNA genetic research from Cabinda suggests that only haplogroups that originated in West Africa are found there today, and the distinctive L 0 of the pre-Bantu population is missing, suggesting that there was a complete population replacement. In South Africa, however,

2520-484: Is estimated as having taken place over 500,000 years ago (marking the split of the H. sapiens lineage from ancestors shared with other known archaic hominins). But the oldest split among modern human populations (such as the Khoisan split from other groups) has been recently dated to between 350,000 and 260,000 years ago, and the earliest known examples of H. sapiens fossils also date to about that period, including

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2640-553: Is evidence that the characteristic human brain development, especially the prefrontal cortex, was due to "an exceptional acceleration of metabolome evolution ... paralleled by a drastic reduction in muscle strength. The observed rapid metabolic changes in brain and muscle, together with the unique human cognitive skills and low muscle performance, might reflect parallel mechanisms in human evolution." The Schöningen spears and their correlation of finds are evidence that complex technological skills already existed 300,000 years ago, and are

2760-471: Is from the Khoekhoe language and refers to foragers ("those who pick things up from the ground") who do not own livestock. As such, it was used in reference to all hunter-gatherer populations who came into contact with Khoekhoe-speaking communities, and was largely referring to the lifestyle, distinct from a pastoralist or agriculturalist one, and not to any particular ethnicity. While there are attendant cosmologies and languages associated with this way of life,

2880-468: Is less than a metre (1000 mm; 39.4 in). The contemporary San and Khoi peoples resemble those represented by the ancient Sangoan skeletal remains. Against the traditional interpretation that finds a common origin for the Khoi and San, other evidence has suggested that the ancestors of the Khoi peoples are relatively recent pre-Bantu agricultural immigrants to southern Africa who abandoned agriculture as

3000-535: Is of the order of about 1% to 4% in Europeans and East Asians, and highest among Melanesians (the last also having Denisova hominin admixture at 4% to 6% in addition to neanderthal admixture). Cumulatively, about 20% of the Neanderthal genome is estimated to remain present spread in contemporary populations. In September 2019, scientists reported the computerized determination, based on 260 CT scans , of

3120-477: Is preserved (if it is preserved at all). This contrasts with archaic humans, where the brow ridge is pronounced and unbroken. Modern humans commonly have a steep, even vertical forehead whereas their predecessors had foreheads that sloped strongly backwards. According to Desmond Morris , the vertical forehead in humans plays an important role in human communication through eyebrow movements and forehead skin wrinkling. Brain size in both Neanderthals and AMH

3240-521: Is significantly larger on average (but overlapping in range) than brain size in H. erectus . Neanderthal and AMH brain sizes are in the same range, but there are differences in the relative sizes of individual brain areas, with significantly larger visual systems in Neanderthals than in AMH. Compared to archaic people, anatomically modern humans have smaller, differently shaped teeth. This results in

3360-445: Is taken to reflect a process towards a smaller and more fine-boned skeleton beginning around 50,000–30,000 years ago. The cranium lacks a pronounced occipital bun in the neck, a bulge that anchored considerable neck muscles in Neanderthals. Modern humans, even the earlier ones, generally have a larger fore-brain than the archaic people, so that the brain sits above rather than behind the eyes. This will usually (though not always) give

3480-634: Is the remnant of an independent western Batwa ( Mbenga or "Baaka") language. Before the Bantu expansion, Khoisan -speaking peoples inhabited Southern Africa. Their descendants have largely mixed with other peoples and adopted other languages. A few still live by foraging, often supplemented by working for neighbouring farmers in the arid regions around the Kalahari desert, while a larger number of Nama continue their traditional subsistence by raising livestock in Namibia and adjacent South Africa. Prior to

3600-426: Is thought that Central African Pygmies and Bantus branched out from a common ancestral population c. 70,000 years ago. Many Batwa groups speak Bantu languages; however, a considerable portion of their vocabulary is not Bantu in origin. Much of this vocabulary is botanical, deals with honey collecting, or is otherwise specialised for the forest and is shared between western Batwa groups. It has been proposed that this

3720-602: Is used to determine when different subgroups separated from one another, and hence their last common ancestry. The authors of these studies suggested that the San may have been one of the first populations to differentiate from the most recent common paternal ancestor of all extant humans. Various Y-chromosome studies since confirmed that the Khoisan carry some of the most divergent (oldest) Y-chromosome haplogroups . These haplogroups are specific sub-groups of haplogroups A and B ,

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3840-604: The Arctic . Both Neanderthal and EEMH had somewhat larger cranial volumes on average than modern Europeans, suggesting the relaxation of selection pressures for larger brain volume after the end of the LGM. Examples for still later adaptations related to agriculture and animal domestication including East Asian types of ADH1B associated with rice domestication , or lactase persistence , are due to recent selection pressures. Bantu expansion The Bantu expansion

3960-456: The Bantu languages are treated as synonymous with the geographic location of ceramic remnants; the popular approach of attempting to correlate linguistic reconstructions with archaeological data has resulted in propagation of the faulty presumption and circular reasoning that the earliest ceramic manufacturing in a given area is evidence for the earliest presence of Bantu-speakers . Within

4080-634: The Cape Flats . Anatomically modern humans Early modern human ( EMH ), or anatomically modern human ( AMH ), are terms used to distinguish Homo sapiens (the only extant Hominina species) that are anatomically consistent with the range of phenotypes seen in contemporary humans, from extinct archaic human species. This distinction is useful especially for times and regions where anatomically modern and archaic humans co-existed, for example, in Paleolithic Europe . Among

4200-459: The Congo River system. The expansion reached South Africa, probably as early as AD 300. Bantuists believe that the Bantu expansion most probably began on the highlands between Cameroon and Nigeria . The 60,000-km Mambilla region straddling the borderlands here has been identified as containing remnants of "the Bantu who stayed home" as the bulk of Bantu-speakers moved away from

4320-649: The German Colonial Empire between 1904 and 1908. In South Africa the Khoi-San suffered genocide at the hands of the Bantu Nguni tribes in the provinces of KwaZulu Natal and the Eastern Cape. The Khoi-San of both provinces were subjected to violent campaigns of territorial dominance by the invading Bantu settlers of the Zulu and Xhosa tribes. The Bantu genocide killed or drove off the men, while

4440-669: The Jebel Irhoud remains from Morocco (ca. 300,000 or 350–280,000 years ago), the Florisbad Skull from South Africa (ca. 259,000 years ago), and the Omo remains from Ethiopia (ca. 195,000, or, as more recently dated, ca. 233,000 years ago). An mtDNA study in 2019 proposed an origin of modern humans in Botswana (and a Khoisan split) of around 200,000 years. However, this proposal has been widely criticized by scholars, with

4560-575: The Mesolithic and the Neolithic , due to increased selection pressures and due to founder effects associated with migration . Alleles predictive of light skin have been found in Neanderthals , but the alleles for light skin in Europeans and East Asians, associated with KITLG and ASIP , are (as of 2012 ) thought to have not been acquired by archaic admixture but recent mutations since

4680-797: The Monomatapa kings built the Great Zimbabwe complex. The Swahili city-states were also established early in this period. These include sultanates based at Lamu , Mombasa , Kilwa , Pate and Malindi . The Swahili traded with the inland kingdoms, including Great Zimbabwe. Such processes of state-formation occurred with increasing frequency from the 16th century onward. They likely resulted from denser population, which led to more specialised divisions of labour, including military power, while making outmigration more effortful. Other factors promoting state-formation were increased trade among African communities and with European and Arab traders on

4800-499: The Sandawe language of Tanzania ("Khoe–Sandawe"). The Hadza language of Tanzania has been associated with the Khoisan group due to the presence of click consonants. The Khoisan are one of the only populations with epicanthic folds outside of East Asia. They typically have hair texture of the tightest possible curl, a form of kinky hair sometimes referred to as "peppercorn" because of how it can roll into separate rounds on

4920-743: The University of the Free State discovered 8,000-year-old carvings made by the Khoisan people. The carvings depicted a hippopotamus, horse, and antelope in the 'Rain Snake' Dyke of the Vredefort impact structure , which may have spiritual significance regarding the rain-making mythology of the Khoisan. In the Herero and Namaqua genocide , about 10,000 Nama , a Khoekhoe group, and an unknown number of San people were killed in an extermination campaign by

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5040-685: The Xhosa people . Georg Schmidt, a Moravian Brother from Herrnhut , Saxony, now Germany, founded Genadendal in 1738, which was the first mission station in southern Africa, among the Khoi people in Baviaanskloof in the Riviersonderend Mountains . Early European settlers sometimes intermarried with Khoikhoi women, resulting in a sizeable mixed-race population now known as the Griqua . The Griqua people too would migrate to what

5160-459: The brain case is quite rounded and distinct from that of the Neanderthals and is similar to the brain case of modern humans. It is uncertain whether the robust traits of some of the early modern humans like Skhul V reflects mixed ancestry or retention of older traits. The "gracile" or lightly built skeleton of anatomically modern humans has been connected to a change in behavior, including increased cooperation and "resource transport". There

5280-603: The lipid catabolic process . A 2017 study found correlation of Neanderthal admixture in phenotypic traits in modern European populations. Physiological or phenotypical changes have been traced to Upper Paleolithic mutations, such as the East Asian variant of the EDAR gene, dated to c. 35,000 years ago. Recent divergence of Eurasian lineages was sped up significantly during the Last Glacial Maximum (LGM),

5400-850: The recent "out of Africa" migration , likely between 60,000 and 40,000 years ago. Recent admixture analyses have added to the complexity, finding that Eastern Neanderthals derive up to 2% of their ancestry from anatomically modern humans who left Africa some 100 kya . The extent of Neanderthal admixture (and introgression of genes acquired by admixture) varies significantly between contemporary racial groups, being absent in Africans, intermediate in Europeans and highest in East Asians. Certain genes related to UV-light adaptation introgressed from Neanderthals have been found to have been selected for in East Asians specifically from 45,000 years ago until around 5,000 years ago. The extent of archaic admixture

5520-462: The recent peopling of Eurasia some 70 kya. They were much more widespread than today, their modern distribution being due to their decimation in the course of the Bantu expansion . They were dispersed throughout much of southern and southeastern Africa. There was also a significant back-migration of bearers of L0 towards eastern Africa between 120 and 75 kya. Rito et al. (2013) speculate that pressure from such back-migration may even have contributed to

5640-698: The 11th and 16th centuries, powerful Bantu-speaking states on a scale larger than local chiefdoms began to emerge. Notable early kingdoms include the Kingdom of the Kongo in present-day Angola and the Democratic Republic of the Congo , the Bunyoro Kitara Kingdom in the Great Lakes region, the Kingdom of Mapungubwe (c.1075–c.1220) in present-day South Africa , and the Zambezi River, where

5760-474: The 1980s all extant groups have tended to be subsumed into a single species, H. sapiens , avoiding division into subspecies altogether. Some sources show Neanderthals ( H. neanderthalensis ) as a subspecies ( H. sapiens neanderthalensis ). Similarly, the discovered specimens of the H. rhodesiensis species have been classified by some as a subspecies ( H. sapiens rhodesiensis ), although it remains more common to treat these last two as separate species within

5880-689: The 2010s with the discovery of admixture events ( introgression ) of populations of H. sapiens with populations of archaic humans over the period of between roughly 100,000 and 30,000 years ago, both in Eurasia and in Sub-Saharan Africa. Neanderthal admixture , in the range of 1–4%, is found in all modern populations outside of Africa, including in Europeans, Asians, Papua New Guineans, Australian Aboriginals, Native Americans, and other non-Africans. This suggests that interbreeding between Neanderthals and anatomically modern humans took place after

6000-498: The Americas by about 40,000–25,000 years ago. Evidence for the overwhelming contribution of this "recent" ( L3 -derived) expansion to all non-African populations was established based on mitochondrial DNA , combined with evidence based on physical anthropology of archaic specimens , during the 1990s and 2000s, and has also been supported by Y DNA and autosomal DNA . The assumption of complete replacement has been revised in

6120-434: The Bantu expansion. Nilo-Saharan -speaking herder populations comprised a third group of the area's pre-Bantu expansion inhabitants. Linguistic, archeological and genetic evidence indicates that during the course of the Bantu expansion, "independent waves of migration of western African and East African Bantu-speakers into southern Africa occurred." In some places, genetic evidence suggests that Bantu language expansion

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6240-650: The Bantu. Local population dropped after the Khoi were exposed to smallpox from Europeans. The Khoi waged more frequent attacks against Europeans when the Dutch East India Company enclosed traditional grazing land for farms. Khoikhoi social organisation was profoundly damaged and, in the end, destroyed by colonial expansion and land seizure from the late 17th century onwards. As social structures broke down, some Khoikhoi people settled on farms and became bondsmen (bondservants) or farm workers; many were incorporated into existing Khoi clan and family groups of

6360-502: The Central African rainforest is extremely spotty and consequently far from convincing so as to be taken as a reflection of a steady influx of Bantu speakers into the forest, let alone movement on a larger scale." Seidensticker (2024) indicates that the prevalent paradigm for the Bantu expansion has a forced connection between Central African ceramics and Central African languages , where the geographic location of speakers of

6480-716: The Colonial/Apartheid era, Afrikaans-speaking persons with partial Khoesān ancestry were historically also grouped as Cape Blacks ( Afrikaans : Kaap Swartes ) or Western Cape Blacks ( Afrikaans : Wes-Kaap Swartes ) to rather inaccurately distinguish them from the Bantu-speaking peoples , the other indigenous African population of South Africa who also had significant Khoe-San ancestry. The term Khoisan (also spelled KhoiSan , Khoi-San , Khoe-San ) has also been introduced in South African usage as

6600-621: The Dutch in 1812) to retain a racist based system of governance because on average only white people owned property adequate to meet the test. In the Herero and Namaqua genocide in German South-West Africa , over 10,000 Nama are estimated to have been killed during 1904–1907. The San of the Kalahari were described in Specimens of Bushman Folklore by Wilhelm H. I. Bleek and Lucy C. Lloyd (1911). They were brought to

6720-430: The Khoisan in both matrilineal and patrilineal groupings is a further indicator that they represent a population historically distinct from other Africans. Some genomic studies have further revealed that Khoisan groups have been influenced by 9 to 30% genetic admixture in the last few thousand years from an East African population who carried a Eurasian admixture component. Furthermore, they place an East African origin for

6840-633: The Khoisan. The San populations ancestral to the Khoisan were spread throughout much of southern and eastern Africa throughout the Late Stone Age after about 75 ka. A further expansion dated to about 20 ka has been proposed based on the distribution of the L0d haplogroup. Rosti et al. suggest a connection of this recent expansion with the spread of click consonants to eastern African languages ( Hadza language ). The Late Stone Age Sangoan industry occupied southern Africa in areas where annual rainfall

6960-399: The LGM. Phenotypes associated with the " white " or " Caucasian " populations of Western Eurasian stock emerge during the LGM, from about 19,000 years ago. Average cranial capacity in modern human populations varies in the range of 1,200 to 1,450 cm for adult males. Larger cranial volume is associated with climatic region, the largest averages being found in populations of Siberia and

7080-539: The Mambilla region. Initially, archaeologists believed that they could find archaeological similarities in the region's ancient cultures that the Bantu-speakers were held to have traversed. Linguists, classifying the languages and creating a genealogical table of relationships, believed they could reconstruct material culture elements. They believed that the expansion was caused by the development of agriculture,

7200-438: The Neanderthal lineage, as "pre-Neanderthal" or "early Neanderthal", while the divergence time between the Neanderthal and modern lineages has been pushed back to before the emergence of H. heidelbergensis , to close to 800,000 years ago, the approximate time of disappearance of H. antecessor . The term Middle Paleolithic is intended to cover the time between the first emergence of H. sapiens (roughly 300,000 years ago) and

7320-784: The Omo Kibbish remains. Fossil evidence for modern human presence in West Asia is ascertained for 177,000 years ago, and disputed fossil evidence suggests expansion as far as East Asia by 120,000 years ago. In July 2019, anthropologists reported the discovery of 210,000 year old remains of a H. sapiens and 170,000 year old remains of a H. neanderthalensis in Apidima Cave , Peloponnese , Greece , more than 150,000 years older than previous H. sapiens finds in Europe. A significant dispersal event, within Africa and to West Asia,

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7440-487: The Tanzanian Luxmanda . Geneticists in 2024 sampled ancient 10,000 year old remains from South Africa, Oakhurst Rockshelter. The examined population had a strong genetic continuity with the San and Khoe. The later advent of pastoralism and farming groups in the last 2,000 years would transform the genepool of most parts of Southern Africa, but many Khoisan preserve, and are identical to the genetic signature of

7560-525: The age of Y-chromosomal Adam have been pushed back significantly with the discovery of an ancient Y-chromosomal lineage in 2013, to likely beyond 300,000 years ago. There have, however, been no reports of the survival of Y-chromosomal or mitochondrial DNA clearly deriving from archaic humans (which would push back the age of the most recent patrilinear or matrilinear ancestor beyond 500,000 years). Fossil teeth found at Qesem Cave (Israel) and dated to between 400,000 and 200,000 years ago have been compared to

7680-641: The arrival of Bantus in Southeast Africa, Cushitic -speaking peoples had migrated into the region from the Ethiopian Highlands and other more northerly areas. The first waves consisted of Southern Cushitic speakers, who settled around Lake Turkana and parts of Tanzania beginning around 5,000 years ago. Many centuries later, around AD 1000, some Eastern Cushitic speakers also settled in northern and coastal Kenya . Khoisan -speaking hunter-gatherers also inhabited Southeast Africa before

7800-552: The beginning of the Common Era , gave rise to the Khoe people, who were pastoralists . This group carried DNA from Eurasian as well as some Neanderthal groups. Due to their early expansion and separation, the populations ancestral to the Khoisan have been estimated as having represented the "largest human population" during the majority of the anatomically modern human timeline, from their early separation before 150 kya until

7920-499: The beginning of the MIS 5 " megadrought " 130,000 years ago, there were two ancestral population clusters in Africa, bearers of mt-DNA haplogroup L0 in southern Africa ancestral to the Khoi-San, and bearers of haplogroup L1-6 in central/eastern Africa ancestral to everyone else. This group gave rise to the San population of hunter gatherers . A much later wave of migration, around or before

8040-514: The case in the Bantu language-speaking Lemba of Southern Africa . Where Bantu was adopted via language shift of existing populations, prior African languages were spoken, probably from African language families that are now lost, except as substrate influences of local Bantu languages (such as click sounds in local Bantu languages). It seems likely that the expansion of the Bantu-speaking people from their core region in West Africa began around 4000–3500   BC. Although early models posited that

8160-447: The climate dried and either joined the San as hunter-gatherers or retained pastoralism. With the hypothesized arrival of pastoralists & bantoid agro-pastoralists in southern Africa starting around 2,300 years ago, linguistic development is later seen in the click consonants and loan words from ancient Khoe-san languages into the evolution of blended agro-pastoralist & hunter-gatherer communities that would eventually evolve into

8280-647: The coasts, technological innovations in economic activity, and new techniques in the political-spiritual ritualisation of royalty as the source of national strength and health. Other inland centres established during this phase of expansion include Bigo bya Mugenyi in Uganda , Thimlich Ohinga in Kenya and the Kweneng' Ruins in South Africa . Manfred K. H. Eggert stated that "the current archaeological record in

8400-982: The common ancestors of West African and Proto-Bantu peoples may have originated in the western region of the Sahara , amid the Kiffian period at Gobero , and may have migrated southward, from the Sahara into various parts of West Africa (e.g., Benin , Cameroon , Ghana , Nigeria , Togo ), as a result of desertification of the Green Sahara in 7000 BC. From Nigeria and Cameroon, agricultural Proto-Bantu peoples began to migrate , and amid migration, diverged into East Bantu peoples (e.g., Democratic Republic of Congo ) and West Bantu peoples (e.g., Congo, Gabon ) between 2500 BC and 1200 BC. He suggests that Igbo people and Yoruba people may have admixture from back-migrated Bantu peoples. The Atlantic-Congo family comprises

8520-548: The common cultural origin of their original speakers. The linguistic core of the Bantu languages, which comprise a branch of the Atlantic-Congo language family , was located in the southern regions of Cameroon . Genetic evidence also indicates that there was a large human migration from central Africa, with varying levels of admixture with local population. The expansion is believed to have taken place in at least two waves, between about 4,000 and 2,000 years ago (approximately 2,000 BC to AD 1). Linguistic analysis suggests that

8640-568: The communities already present at the coast. Between 300 AD-1000 AD, through participation in the long-existing Indian Ocean trade route , these communities established links with Arabian and Indian traders, leading to the development of the Swahili culture . Other pioneering groups had reached modern KwaZulu-Natal in South Africa by AD 300 along the coast, and the modern Limpopo Province (formerly Northern Transvaal ) by AD 500. Between

8760-491: The contemporary height distribution measured among Malay people , for one. Following the peopling of Africa some 130,000 years ago, and the recent Out-of-Africa expansion some 70,000 to 50,000 years ago, some sub-populations of H. sapiens had been essentially isolated for tens of thousands of years prior to the early modern Age of Discovery . Combined with archaic admixture this has resulted in significant genetic variation , which in some instances has been shown to be

8880-500: The continent and led to extensive admixture between migrants and local populations. A 2023 genetic study of 1,487 Bantu speakers sampled from 143 populations across 14 African countries revealed that the expansion occurred ~4,000 years ago in Western Africa. The results showed that Bantu speakers received significant gene-flow from local groups in regions they expanded into. Based on dental evidence, Irish (2016) concluded that

9000-673: The defeat of the Xhosa rebellion in 1853, the new Cape Government endeavoured to grant the Khoi political rights to avert future racial discontent. The government enacted the Cape franchise in 1853, which decreed that all male citizens meeting a low property test, regardless of colour, had the right to vote and to seek election in Parliament. The property test was an indirect way by the British Cape Government (who took over from

9120-658: The dental material from the younger (120,000–80,000 years ago) Skhul and Qafzeh hominins . Dispersal of early H. sapiens begins soon after its emergence, as evidenced by the North African Jebel Irhoud finds (dated to around 315,000 years ago). There is indirect evidence for H. sapiens presence in West Asia around 270,000 years ago. The Florisbad Skull from Florisbad, South Africa, dated to about 259,000 years ago, has also been classified as representing early H. sapiens . Scerri (2018) , pp. 582–594 In September 2019, scientists proposed that

9240-412: The description of contemporary humans with the subspecies name Homo sapiens sapiens . However, biological anthropologist Chris Stringer does not consider idaltu distinct enough within H. sapiens to warrant its own subspecies designation. A further division of AMH into "early" or "robust" vs. "post-glacial" or " gracile " subtypes has since been used for convenience. The emergence of "gracile AMH"

9360-414: The dispersal of East African populations out of Africa at about 70 kya. Recent work has suggested that the multi-regional hypothesis may be supported by current human population genetic data. A 2023 study published in the journal Nature suggests that current genetic data may be best understood as reflecting internal admixtures of multiple population sources across Africa, including ancestral populations of

9480-623: The distal bones were shorter, usually thought to be an adaptation to cold climate. The same adaptation is found in some modern people living in the polar regions. Height ranges overlap between Neanderthals and AMH, with Neanderthal averages cited as 164 to 168 cm (65 to 66 in) and 152 to 156 cm (60 to 61 in) for males and females, respectively, which is largely identical to pre-industrial average heights for AMH. Contemporary national averages range between 158 to 184 cm (62 to 72 in) in males and 147 to 172 cm (58 to 68 in) in females. Neanderthal ranges approximate

9600-493: The earliest H. sapiens (and last common human ancestor to modern humans) arose between 350,000 and 260,000 years ago through a merging of populations in East and South Africa . Among extant populations, the Khoi-San (or " Capoid ") hunters-gatherers of Southern Africa may represent the human population with the earliest possible divergence within the group Homo sapiens sapiens . Their separation time has been estimated in

9720-437: The earliest and most robustly built modern humans were less robust than those of Neanderthals (and from what little we know from Denisovans), having essentially modern proportions. Particularly regarding the long bones of the limbs, the distal bones (the radius / ulna and tibia / fibula ) are nearly the same size or slightly shorter than the proximal bones (the humerus and femur ). In ancient people, particularly Neanderthals,

9840-467: The earliest genetic splits among modern people, according to some evidence, dating to around the same time. Sustained archaic human admixture with modern humans is known to have taken place both in Africa and (following the recent Out-Of-Africa expansion ) in Eurasia, between about 100,000 and 30,000 years ago. The binomial name Homo sapiens was coined by Linnaeus , 1758 . The Latin noun homō (genitive hominis ) means "human being", while

9960-414: The early speakers were both iron-using and agricultural, definitive archaeological evidence that they used iron does not appear until as late as 400   BC, though they were agricultural. The western branch, not necessarily linguistically distinct, according to Christopher Ehret , followed the coast and the major rivers of the Congo system southward, reaching central Angola by around 500   BC. It

10080-575: The emergence of "anatomically modern humans". Since the 2000s, the discovery of older remains with comparable characteristics, and the discovery of ongoing hybridization between "modern" and "archaic" populations after the time of the Omo remains, have opened up a renewed debate on the age of H. sapiens in journalistic publications. H. s. idaltu , dated to 160,000 years ago, has been postulated as an extinct subspecies of H. sapiens in 2003. H. neanderthalensis , which became extinct about 40,000 years ago,

10200-433: The emergence of a much more detailed picture, intermediate between the two competing scenarios outlined above: The recent Out-of-Africa expansion accounts for the predominant part of modern human ancestry, while there were also significant admixture events with regional archaic humans. Since the 1970s, the Omo remains, originally dated to some 195,000 years ago, have often been taken as the conventional cut-off point for

10320-891: The end of Apartheid in 1994, the term "Khoisan" has gradually come to be used as a self-designation by South African Khoikhoi as representing the "first nations" of South Africa vis-a-vis the ruling Bantu majority. A conference on "Khoisan Identities and Cultural Heritage" was organised by the University of the Western Cape in 1997. and "Khoisan activism" has been reported in the South African media beginning in 2015. The South African government allowed Khoisan families (up until 1998) to pursue land claims which existed prior to 1913. The South African Deputy Chief Land Claims Commissioner, Thami Mdontswa, has said that constitutional reform would be required to enable Khoisan people to pursue further claims to land from which their direct ancestors were removed prior to 9 June 1913. In 2019, scientists from

10440-802: The end of MIS5 (75,000 years ago), and is known only from fossil evidence and from archaic admixture . Eurasia was re-populated by early modern humans in the so-called "recent out-of-Africa migration" post-dating MIS5, beginning around 70,000–50,000 years ago. In this expansion, bearers of mt-DNA haplogroup L3 left East Africa, likely reaching Arabia via the Bab-el-Mandeb , and in the Great Coastal Migration spread to South Asia, Maritime South Asia and Oceania between 65,000 and 50,000 years ago, while Europe , East and North Asia were reached by about 45,000 years ago. Some evidence suggests that an early wave of humans may have reached

10560-510: The expansion proceeded in two directions: the first went across or along the Northern border of the Congo forest region (towards East Africa), and the second – and possibly others – went south along Africa's Atlantic coast into what is now the Republic of the Congo , Gabon , Cameroon , Democratic Republic of the Congo , and Angola , or inland along the many south-to-north flowing rivers of

10680-486: The extinction of all other human varieties, while the " multiregional evolution " model postulated the survival of regional forms of archaic humans, gradually converging into the modern human varieties by the mechanism of clinal variation , via genetic drift , gene flow and selection throughout the Pleistocene. Since the 2000s, the availability of data from archaeogenetics and population genetics has led to

10800-475: The far south several centuries before Bantu-speaking migrants did. Archaeological , linguistic , genetic , and environmental evidence all support the conclusion that the Bantu expansion was a significant human migration. Generally, the movements of Bantu language-speaking peoples from the Cameroon/Nigeria border region throughout much of sub-Saharan Africa radically reshaped the genetic structure of

10920-486: The first obvious proof of an active (big game) hunt . H. heidelbergensis already had intellectual and cognitive skills like anticipatory planning, thinking and acting that so far have only been attributed to modern man. The ongoing admixture events within anatomically modern human populations make it difficult to estimate the age of the matrilinear and patrilinear most recent common ancestors of modern populations ( Mitochondrial Eve and Y-chromosomal Adam ). Estimates of

11040-608: The further away the labourer was from Cape Town, the more difficult it became to transport agricultural produce to the markets. The issuing of grazing licences north of the Berg River in what was then the Tulbagh Basin propelled colonial expansion in the area. This system of land relocation led to the Khoijhou losing their land and livestock as well as dramatic change in the social, economic and political development. After

11160-438: The genus Homo rather than as subspecies within H. sapiens . All humans are considered to be a part of the subspecies H. sapiens sapiens , a designation which has been a matter of debate since a species is usually not given a subspecies category unless there is evidence of multiple distinct subspecies. The divergence of the lineage that would lead to H. sapiens out of archaic human varieties derived from H. erectus ,

11280-511: The globalised world's attention in the 1950s by South African author Laurens van der Post in a six-part television documentary. The Ancestral land conflict in Botswana concerns the Central Kalahari Game Reserve (CKGR), established in 1961 for wildlife, while the San were permitted to continue their hunter-gatherer lifestyle. In the 1990s, the government of Botswana began a policy of "relocating" CKGR residents outside

11400-500: The lineage of modern humans since the split from the lineage of Neanderthals , roughly 500,000 to 800,000 years ago. The time of divergence between archaic H. sapiens and ancestors of Neanderthals and Denisovans caused by a genetic bottleneck of the latter was dated at 744,000 years ago, combined with repeated early admixture events and Denisovans diverging from Neanderthals 300 generations after their split from H. sapiens , as calculated by Rogers et al. (2017). The derivation of

11520-539: The making of ceramics, and the use of iron, which permitted new ecological zones to be exploited. In 1966, Roland Oliver published an article presenting these correlations as a reasonable hypothesis. The hypothesized Bantu expansion pushed out or assimilated the hunter-forager proto- Khoisan , who had formerly inhabited Southern Africa. In Eastern and Southern Africa , Bantu speakers may have adopted livestock husbandry from other unrelated Cushitic -and Nilotic -speaking peoples they encountered. Herding practices reached

11640-579: The modern human lineage following the split from the Neanderthal lineage. Such a cladistic definition would extend the age of H. sapiens to over 500,000 years. Estimates for the split between the Homo sapiens line and combined Neanderthal / Denisovan line range from between 503,000 and 565,000 years ago; between 550,000 and 765,000 years ago; and (based on rates of dental evolution) possibly more than 800,000 years ago. Extant human populations have historically been divided into subspecies , but since around

11760-512: The now extant, amalgamated modern native linguistic communities found in South Africa, Botswana & Namibia (e.g. in South African Xhosa , Sotho , Tswana , Zulu people.) Today these groups represent the quantitative majority of extant admixed ancient Khoe-San descendants by the millions. The Khoikhoi enter the historical record with their first contact with Portuguese explorers, about 1,000 years after their displacement by

11880-427: The older hunter-gatherers. On 21 September 2020, the University of Cape Town launched its new Khoi and San Centre, with an undergraduate degree programme planned to be rolled out in the following years. The centre will support and consolidate this collaborative work on research commissions on language (including Khoekhoegowab ), sacred human remains, land and gender. Many descendants of Khoisan people still live on

12000-615: The oldest known remains of Homo sapiens are those found at the Omo-Kibish I archaeological site in south-western Ethiopia , dating to about 233,000 to 196,000 years ago, the Florisbad site in South Africa, dating to about 259,000 years ago, and the Jebel Irhoud site in Morocco, dated about 315,000 years ago. Extinct species of the genus Homo include Homo erectus (extant from roughly 2 to 0.1 million years ago) and

12120-477: The participle sapiēns means "discerning, wise, sensible". The species was initially thought to have emerged from a predecessor within the genus Homo around 300,000 to 200,000 years ago. A problem with the morphological classification of "anatomically modern" was that it would not have included certain extant populations. For this reason, a lineage-based ( cladistic ) definition of H. sapiens has been suggested, in which H. sapiens would by definition refer to

12240-472: The paternal haplogroup E1b1b found in these Southern African populations, as well as the introduction of pastoralism into the region. The paper also noted that the Bantu expansion had a notable genetic impact in a number of Khoisan groups. On the basis of PCA projections, the East African ancestry identified in the genomes of Khoe-Kwadi speakers and other southern Africans is related to an individual from

12360-486: The period held by some to mark the emergence of full behavioral modernity (roughly by 50,000 years ago, corresponding to the start of the Upper Paleolithic ). Many of the early modern human finds, like those of Jebel Irhoud , Omo , Herto , Florisbad , Skhul , and Peștera cu Oase exhibit a mix of archaic and modern traits. Skhul V, for example, has prominent brow ridges and a projecting face. However,

12480-508: The recent evidence overall (genetic, fossil, and archaeological) supporting an origin for H. sapiens approximately 100,000 years earlier and in a broader region of Africa than the study proposes. In September 2019, scientists proposed that the earliest H. sapiens (and last common human ancestor to modern humans) arose between 350,000 and 260,000 years ago through a merging of populations in East and South Africa . An alternative suggestion defines H. sapiens cladistically as including

12600-468: The region. Archaeological evidence from the separate works of Jean Hurault (1979, 1986 and 1988) and Rigobert Tueché (2000) in the region indicates cultural continuity from 3000 BC until today. The majority of the groups of the Bamenda highlands (occupied for 2000 years until today), somewhat south and contiguous with the Mambilla region, have an ancient history of descent from the north in the direction of

12720-529: The reserve. In 2002, the government cut off all services to CKGR residents. A legal battle began, and in 2006 the High Court of Botswana ruled that the residents had been forcibly and unconstitutionally removed. The policy of relocation continued, however, and in 2012 the San people (Basarwa) appealed to the United Nations to force the government to recognise their land and resource rights. Following

12840-774: The result of directional selection taking place over the past 15,000 years, i.e., significantly later than possible archaic admixture events. Some climatic adaptations, such as high-altitude adaptation in humans , are thought to have been acquired by archaic admixture. Introgression of genetic variants acquired by Neanderthal admixture have different distributions in European and East Asians , reflecting differences in recent selective pressures. A 2014 study reported that Neanderthal-derived variants found in East Asian populations showed clustering in functional groups related to immune and haematopoietic pathways , while European populations showed clustering in functional groups related to

12960-520: The scalp. Charles Darwin wrote about the Khoisan and sexual selection in The Descent of Man in 1882, commenting that their steatopygia , seen primarily in females, evolved through sexual selection in human evolution , and that "the posterior part of the body projects in a most wonderful manner". Historically, some females were observed by anthropologists to exhibit elongated labia minora , which sometimes projected as much as 10 cm below

13080-542: The settlement also began to attract other Khoi, Xhosa and mixed-race groups of the Cape. The so-called "Bushman wars"were to a large extent the response of the San after their dispossession. At the start of the 18th century, the Khoikhoi in the Western Cape lived in a state dominated by the Dutch. By the end of the century the majority of the Khoisan operated as 'wage labourers', not that dissimilar to slaves. Geographically,

13200-543: The southeast from the Great Lakes region were more rapid, with initial settlements widely dispersed near the coast and near rivers, due to comparatively difficult farming conditions in areas farther from water. Archaeological findings have shown that by 100 BC to 300 AD, Bantu speaking communities were present at the coastal areas of Misasa in Tanzania and Kwale in Kenya. These communities also integrated and intermarried with

13320-420: The term is an economic designator rather than a cultural or ethnic one. The compound term Khoisan / Khoesān is a modern anthropological convention in use since the early-to-mid 20th century. Khoisan is a coinage by Leonhard Schulze in the 1920s and popularised by Isaac Schapera . It entered wider usage from the 1960s based on the proposal of a " Khoisan " language family by Joseph Greenberg . During

13440-450: The two earliest branches on the human Y-chromosome tree. Similar to findings from Y-chromosome studies, mitochondrial DNA studies also showed evidence that the Khoisan people carry high frequencies of the earliest haplogroup branches in the human mitochondrial DNA tree. The most divergent (oldest) mitochondrial haplogroup, L0d , has been identified at its highest frequencies in the southern African Khoi and San groups. The distinctiveness of

13560-446: The vulva when standing. Though well documented, the motivations behind this practice and the voices of the women who perform it are rarely explored in the research. In the 1990s, genomic studies of the world's peoples found that the Y chromosome of San men share certain patterns of polymorphisms that are distinct from those of all other populations. Because the Y chromosome is highly conserved between generations, this type of DNA test

13680-644: The women and children were absorbed into the Bantu tribes. These forced assimilations led to language adaptations of clicks etc and the Khoi-San DNA also produced lighter skin tones among the Bantu. The Bantu colonisation process totally extinguished the Khoi-San independent identity,to the extent that the remnants now call themselves either Zulu or Xhosa. In Botswana , many of the indigenous San people have been forcibly relocated from their land to reservations. To make them relocate, they were denied access to water on their land and faced arrest if they hunted, which

13800-569: Was a major series of migrations of the original Proto-Bantu -speaking group , which spread from an original nucleus around West - Central Africa . In the process, the Proto-Bantu-speaking settlers displaced, eliminated or absorbed pre-existing hunter-gatherer and pastoralist groups that they encountered. There is linguistic evidence for this expansion – a great many of the languages which are spoken across sub-Equatorial Africa are remarkably similar to each other, suggesting

13920-474: Was also at one point considered to be a subspecies, H. s. neanderthalensis . H. heidelbergensis , dated 600,000 to 300,000 years ago, has long been thought to be a likely candidate for the last common ancestor of the Neanderthal and modern human lineages. However, genetic evidence from the Sima de los Huesos fossils published in 2016 seems to suggest that H. heidelbergensis in its entirety should be included in

14040-551: Was by that time the frontierlands of the Xhosa native reserves and establish Griqualand East, which contained a mostly Xhosa population. Andries Stockenström facilitated the creation of the "Kat River" Khoi settlement near the eastern frontier of the Cape Colony. The settlements thrived and expanded, and Kat River quickly became a large and successful region of the Cape that subsisted more or less autonomously. The people were predominantly Afrikaans-speaking Gonaqua Khoi, but

14160-586: Was largely a result of substantial population replacement. In other places, Bantu language expansion, like many other languages, has been documented with population genetic evidence to have occurred by means other than complete or predominant population replacement (e.g. via language shift and admixture of incoming and existing populations). For example, one study found this to be the case in Bantu language speakers who are African Pygmies or are in Mozambique , while another population genetic study found this to be

14280-596: Was proposed in 2010, combining the ǂʼAmkoe (ǂHoan) language with the ǃKung (Juu) dialect cluster. ǃKung includes about a dozen dialects, with no clear-cut delineation between them. Sands et al. (2010) propose a division into four clusters: The Khoi (Khoe) family is divided into a Khoikhoi ( Khoekhoe and Khoemana dialects) and a Kalahari (Tshu–Khwe) branch. The Kalahari branch of Khoe includes Shua and Tsoa (with dialects), and Kxoe , Naro , Gǁana and ǂHaba (with dialects). Khoe also has been tentatively aligned with Kwadi ("Kwadi–Khoe"), and more speculatively with

14400-442: Was their primary source of food. Their lands lie in the middle of the world's richest diamond field. Officially, the government denies that there is any link to mining and claims the relocation is to preserve the wildlife and ecosystem, even though the San people have lived sustainably on the land for millennia. On the reservations they struggle to find employment, and alcoholism is rampant. The "Khoisan languages" were proposed as

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