117-587: The African houbara ( Chlamydotis undulata ), also known as the houbara bustard ( houbara from Arabic : حُبَارَى , romanized : ḥubārā for bustards in general), is a relatively small bustard native to North Africa , where it lives in arid habitats. The global population is listed as Vulnerable on the IUCN Red List since 2014. There is a population in the Canary Islands which has been assessed as Near Threatened in 2015. It
234-489: A coleoptile that forms the first leaf while the radicle is covered with a coleorhiza that connects to the primary root and adventitious roots form the sides. Here the hypocotyl is a rudimentary axis between radicle and plumule. The seeds of corn are constructed with these structures; pericarp, scutellum (single large cotyledon) that absorbs nutrients from the endosperm, plumule, radicle, coleoptile, and coleorhiza – these last two structures are sheath-like and enclose
351-681: A common ancestor . The divergence between the African and Canarian houbara was estimated at 20,000 to 25,000 years ago. The African houbara is found in North Africa west of the Nile , mainly in the western part of the Sahara desert region in Algeria , Egypt , Libya , Mauritania , Morocco , Tunisia and Western Sahara . Some old records exist from Sudan as well. A small population is found in
468-401: A seed is a plant embryo and nutrient reserve enclosed in a seed coat, a protective outer covering called a testa. More generally, the term "seed" means anything that can be sown , which may include seed and husk or tuber . Seeds are the product of the ripened ovule , after the embryo sac is fertilized by sperm from pollen , forming a zygote . The embryo within a seed develops from
585-575: A collection of related dialects that constitute the precursor of Arabic, first emerged during the Iron Age . Previously, the earliest attestation of Old Arabic was thought to be a single 1st century CE inscription in Sabaic script at Qaryat al-Faw , in southern present-day Saudi Arabia. However, this inscription does not participate in several of the key innovations of the Arabic language group, such as
702-435: A corpus of poetic texts, in addition to Qur'an usage and Bedouin informants whom he considered to be reliable speakers of the ʿarabiyya . Arabic spread with the spread of Islam . Following the early Muslim conquests , Arabic gained vocabulary from Middle Persian and Turkish . In the early Abbasid period , many Classical Greek terms entered Arabic through translations carried out at Baghdad's House of Wisdom . By
819-422: A curved megagametophyte often giving the seed a tight "C" shape. The last ovule shape is called amphitropous , where the ovule is partly inverted and turned back 90 degrees on its stalk (the funicle or funiculus ). In the majority of flowering plants, the zygote's first division is transversely oriented in regards to the long axis, and this establishes the polarity of the embryo. The upper or chalazal pole becomes
936-1081: A dialect of Arabic and written in the Latin alphabet . The Balkan languages, including Albanian, Greek , Serbo-Croatian, and Bulgarian , have also acquired many words of Arabic origin, mainly through direct contact with Ottoman Turkish . Arabic has influenced languages across the globe throughout its history, especially languages where Islam is the predominant religion and in countries that were conquered by Muslims. The most markedly influenced languages are Persian , Turkish , Hindustani ( Hindi and Urdu ), Kashmiri , Kurdish , Bosnian , Kazakh , Bengali , Malay ( Indonesian and Malaysian ), Maldivian , Pashto , Punjabi , Albanian , Armenian , Azerbaijani , Sicilian, Spanish, Greek, Bulgarian, Tagalog , Sindhi , Odia , Hebrew and African languages such as Hausa , Amharic , Tigrinya , Somali , Tamazight , and Swahili . Conversely, Arabic has borrowed some words (mostly nouns) from other languages, including its sister-language Aramaic, Persian, Greek, and Latin and to
1053-416: A difference in a pre-copulation display would indicate that the two are separate species. The committee responded to this scepticism, by explaining that there are differences in both courtship and pre-copulation displays. Results of analysis of mitochondrial DNA sequences of 73 Chlamydotis samples indicates that the houbara bustard and MacQueen's bustard genetically diverged around 430,000 years ago from
1170-411: A few will end in a favorable place for growth. Herbaceous perennials and woody plants often have larger seeds; they can produce seeds over many years, and larger seeds have more energy reserves for germination and seedling growth and produce larger, more established seedlings after germination. Seeds serve several functions for the plants that produce them. Key among these functions are nourishment of
1287-413: A flamboyant display raising the white feathers of the head and neck and withdrawing the head. Females lay two to four eggs on the ground. It rarely vocalizes, but males make 3-5 low booming notes during breeding displays. It is omnivorous, eating seeds , insects and other small creatures. In North Africa, the houbara bustard is hunted by falconers and by hunters with guns. The populations declined in
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#17327726223891404-434: A houbara brought from Tripoli to Vienna 's Tiergarten Schönbrunn . Otis macqueenii was proposed by John Edward Gray in 1832 for a bustard from India drawn by Thomas Hardwicke . The African houbara was subordinated to the genus Chlamydotis by René Lesson in 1839. Houbara fuertaventurae was proposed by Walter Rothschild and Ernst Hartert in 1894 for a houbara from Fuerteventura island. MacQueen's bustard
1521-487: A lesser extent and more recently from Turkish, English, French, and Italian. Arabic is spoken by as many as 380 million speakers, both native and non-native, in the Arab world, making it the fifth most spoken language in the world, and the fourth most used language on the internet in terms of users. It also serves as the liturgical language of more than 2 billion Muslims . In 2011, Bloomberg Businessweek ranked Arabic
1638-415: A location and be there at a time favorable for germination and growth. When the fruits open and release their seeds in a regular way, it is called dehiscent , which is often distinctive for related groups of plants; these fruits include capsules , follicles , legumes , silicles and siliques . When fruits do not open and release their seeds in a regular fashion, they are called indehiscent, which include
1755-690: A millennium before the modern period . Early lexicographers ( لُغَوِيُّون lughawiyyūn ) sought to explain words in the Quran that were unfamiliar or had a particular contextual meaning, and to identify words of non-Arabic origin that appear in the Quran. They gathered shawāhid ( شَوَاهِد 'instances of attested usage') from poetry and the speech of the Arabs—particularly the Bedouin ʾaʿrāb [ ar ] ( أَعْراب ) who were perceived to speak
1872-594: A result, many European languages have borrowed words from it. Arabic influence, mainly in vocabulary, is seen in European languages (mainly Spanish and to a lesser extent Portuguese , Catalan , and Sicilian ) owing to the proximity of Europe and the long-lasting Arabic cultural and linguistic presence, mainly in Southern Iberia, during the Al-Andalus era. Maltese is a Semitic language developed from
1989-462: A script derived from ASA attest to a language known as Hasaitic . On the northwestern frontier of Arabia, various languages known to scholars as Thamudic B , Thamudic D, Safaitic , and Hismaic are attested. The last two share important isoglosses with later forms of Arabic, leading scholars to theorize that Safaitic and Hismaic are early forms of Arabic and that they should be considered Old Arabic . Linguists generally believe that "Old Arabic",
2106-470: A single language, despite mutual incomprehensibility among differing spoken versions. From a linguistic standpoint, it is often said that the various spoken varieties of Arabic differ among each other collectively about as much as the Romance languages . This is an apt comparison in a number of ways. The period of divergence from a single spoken form is similar—perhaps 1500 years for Arabic, 2000 years for
2223-435: A supply of nutrients for the embryo in most monocotyledons and the endospermic dicotyledons. Seeds have been considered to occur in many structurally different types (Martin 1946). These are based on a number of criteria, of which the dominant one is the embryo-to-seed size ratio. This reflects the degree to which the developing cotyledons absorb the nutrients of the endosperm, and thus obliterate it. Six types occur amongst
2340-550: A tegmen from the inner integument while unitegmic seeds have only one integument. Usually, parts of the testa or tegmen form a hard protective mechanical layer. The mechanical layer may prevent water penetration and germination. Amongst the barriers may be the presence of lignified sclereids . The outer integument has a number of layers, generally between four and eight organised into three layers: (a) outer epidermis, (b) outer pigmented zone of two to five layers containing tannin and starch, and (c) inner epidermis. The endotegmen
2457-507: A type of Arabic. Cypriot Arabic is recognized as a minority language in Cyprus. The sociolinguistic situation of Arabic in modern times provides a prime example of the linguistic phenomenon of diglossia , which is the normal use of two separate varieties of the same language, usually in different social situations. Tawleed is the process of giving a new shade of meaning to an old classical word. For example, al-hatif lexicographically means
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#17327726223892574-507: A variety of regional vernacular Arabic dialects , which are not necessarily mutually intelligible. Classical Arabic is the language found in the Quran , used from the period of Pre-Islamic Arabia to that of the Abbasid Caliphate . Classical Arabic is prescriptive, according to the syntactic and grammatical norms laid down by classical grammarians (such as Sibawayh ) and the vocabulary defined in classical dictionaries (such as
2691-405: A way for the species to survive dry or cold seasons. Ephemeral plants are usually annuals that can go from seed to seed in as few as six weeks. Seed germination is a process by which a seed embryo develops into a seedling. It involves the reactivation of the metabolic pathways that lead to growth and the emergence of the radicle or seed root and plumule or shoot. The emergence of the seedling above
2808-476: A wider audience." In the wake of the industrial revolution and European hegemony and colonialism , pioneering Arabic presses, such as the Amiri Press established by Muhammad Ali (1819), dramatically changed the diffusion and consumption of Arabic literature and publications. Rifa'a al-Tahtawi proposed the establishment of Madrasat al-Alsun in 1836 and led a translation campaign that highlighted
2925-590: Is a minimum level of comprehension between all Arabic dialects, this level can increase or decrease based on geographic proximity: for example, Levantine and Gulf speakers understand each other much better than they do speakers from the Maghreb. The issue of diglossia between spoken and written language is a complicating factor: A single written form, differing sharply from any of the spoken varieties learned natively, unites several sometimes divergent spoken forms. For political reasons, Arabs mostly assert that they all speak
3042-559: Is a sister language rather than their direct ancestor. Arabia had a wide variety of Semitic languages in antiquity. The term "Arab" was initially used to describe those living in the Arabian Peninsula , as perceived by geographers from ancient Greece . In the southwest, various Central Semitic languages both belonging to and outside the Ancient South Arabian family (e.g. Southern Thamudic) were spoken. It
3159-509: Is at a suitable temperature with proper soil moisture. This true dormancy or innate dormancy is therefore caused by conditions within the seed that prevent germination. Thus dormancy is a state of the seed, not of the environment. Induced dormancy, enforced dormancy or seed quiescence occurs when a seed fails to germinate because the external environmental conditions are inappropriate for germination, mostly in response to conditions being too dark or light, too cold or hot, or too dry. Seed dormancy
3276-478: Is believed that the ancestors of the Modern South Arabian languages (non-Central Semitic languages) were spoken in southern Arabia at this time. To the north, in the oases of northern Hejaz , Dadanitic and Taymanitic held some prestige as inscriptional languages. In Najd and parts of western Arabia, a language known to scholars as Thamudic C is attested. In eastern Arabia, inscriptions in
3393-481: Is caused by conditions outside the embryo, including: Endogenous dormancy is caused by conditions within the embryo itself, including: The following types of seed dormancy do not involve seed dormancy, strictly speaking, as lack of germination is prevented by the environment, not by characteristics of the seed itself (see Germination ): Not all seeds undergo a period of dormancy. Seeds of some mangroves are viviparous; they begin to germinate while still attached to
3510-408: Is credited with establishing the rules of Arabic prosody . Al-Jahiz (776–868) proposed to Al-Akhfash al-Akbar an overhaul of the grammar of Arabic, but it would not come to pass for two centuries. The standardization of Arabic reached completion around the end of the 8th century. The first comprehensive description of the ʿarabiyya "Arabic", Sībawayhi's al - Kitāb , is based first of all upon
3627-472: Is credited with standardizing Arabic grammar , or an-naḥw ( النَّحو "the way" ), and pioneering a system of diacritics to differentiate consonants ( نقط الإعجام nuqaṭu‿l-i'jām "pointing for non-Arabs") and indicate vocalization ( التشكيل at-tashkīl ). Al-Khalil ibn Ahmad al-Farahidi (718–786) compiled the first Arabic dictionary, Kitāb al-'Ayn ( كتاب العين "The Book of the Letter ع "), and
African houbara - Misplaced Pages Continue
3744-402: Is derived from the inner epidermis of the inner integument, the exotegmen from the outer surface of the inner integument. The endotesta is derived from the inner epidermis of the outer integument, and the outer layer of the testa from the outer surface of the outer integument is referred to as the exotesta . If the exotesta is also the mechanical layer, this is called an exotestal seed, but if
3861-411: Is dull brown with black markings on the wings, a greyish neck and a black ruff along the side of the neck. Males are larger and heavier than females. The African houbara is a small to mid-sized bustard. It measures 55–75 cm (22–30 in) in length and spans 135–170 cm (53–67 in) across the wings. It is brown above and white below, with a black stripe down the sides of its neck. In flight,
3978-574: Is not present in the spoken varieties, but deletes Classical words that sound obsolete in MSA. In addition, MSA has borrowed or coined many terms for concepts that did not exist in Quranic times, and MSA continues to evolve. Some words have been borrowed from other languages—notice that transliteration mainly indicates spelling and not real pronunciation (e.g., فِلْم film 'film' or ديمقراطية dīmuqrāṭiyyah 'democracy'). The current preference
4095-444: Is not the same as seed persistence in the soil or on the plant, though even in scientific publications dormancy and persistence are often confused or used as synonyms. Often, seed dormancy is divided into four major categories: exogenous; endogenous; combinational; and secondary. A more recent system distinguishes five classes: morphological, physiological, morphophysiological, physical, and combinational dormancy. Exogenous dormancy
4212-855: Is official in Mali and recognized as a minority language in Morocco, while the Senegalese government adopted the Latin script to write it. Maltese is official in (predominantly Catholic ) Malta and written with the Latin script . Linguists agree that it is a variety of spoken Arabic, descended from Siculo-Arabic , though it has experienced extensive changes as a result of sustained and intensive contact with Italo-Romance varieties, and more recently also with English. Due to "a mix of social, cultural, historical, political, and indeed linguistic factors", many Maltese people today consider their language Semitic but not
4329-535: Is protection against disease. Seeds protect and nourish the embryo or young plant. They usually give a seedling a faster start than a sporeling from a spore, because of the larger food reserves in the seed and the multicellularity of the enclosed embryo. Unlike animals, plants are limited in their ability to seek out favorable conditions for life and growth. As a result, plants have evolved many ways to disperse their offspring by dispersing their seeds (see also vegetative reproduction ). A seed must somehow "arrive" at
4446-572: Is the third most widespread official language after English and French, one of six official languages of the United Nations , and the liturgical language of Islam . Arabic is widely taught in schools and universities around the world and is used to varying degrees in workplaces, governments and the media. During the Middle Ages , Arabic was a major vehicle of culture and learning, especially in science, mathematics and philosophy. As
4563-446: Is the ability of the embryo to germinate and is affected by a number of different conditions. Some plants do not produce seeds that have functional complete embryos, or the seed may have no embryo at all, often called empty seeds. Predators and pathogens can damage or kill the seed while it is still in the fruit or after it is dispersed. Environmental conditions like flooding or heat can kill the seed before or during germination. The age of
4680-590: Is the variety used in most current, printed Arabic publications, spoken by some of the Arabic media across North Africa and the Middle East, and understood by most educated Arabic speakers. "Literary Arabic" and "Standard Arabic" ( فُصْحَى fuṣḥá ) are less strictly defined terms that may refer to Modern Standard Arabic or Classical Arabic. Some of the differences between Classical Arabic (CA) and Modern Standard Arabic (MSA) are as follows: MSA uses much Classical vocabulary (e.g., dhahaba 'to go') that
4797-411: Is then aborted or absorbed during early development. The seed is composed of the embryo (the result of fertilization) and tissue from the mother plant, which also form a cone around the seed in coniferous plants such as pine and spruce . Seeds are very diverse, and as such there are many terms are used to describe them. A typical seed includes two basic parts: In addition, the endosperm forms
African houbara - Misplaced Pages Continue
4914-413: Is to avoid direct borrowings, preferring to either use loan translations (e.g., فرع farʻ 'branch', also used for the branch of a company or organization; جناح janāḥ 'wing', is also used for the wing of an airplane, building, air force, etc.), or to coin new words using forms within existing roots ( استماتة istimātah ' apoptosis ', using the root موت m/w/t 'death' put into
5031-524: Is used to denote concepts that have arisen in the industrial and post-industrial era , especially in modern times. Due to its grounding in Classical Arabic, Modern Standard Arabic is removed over a millennium from everyday speech, which is construed as a multitude of dialects of this language. These dialects and Modern Standard Arabic are described by some scholars as not mutually comprehensible. The former are usually acquired in families, while
5148-419: Is usually triploid , and is rich in oil or starch , and protein . In gymnosperms, such as conifers , the food storage tissue (also called endosperm) is part of the female gametophyte , a haploid tissue. The endosperm is surrounded by the aleurone layer (peripheral endosperm), filled with proteinaceous aleurone grains. Originally, by analogy with the animal ovum , the outer nucellus layer ( perisperm )
5265-445: The Lisān al-ʻArab ). Modern Standard Arabic (MSA) largely follows the grammatical standards of Classical Arabic and uses much of the same vocabulary. However, it has discarded some grammatical constructions and vocabulary that no longer have any counterpart in the spoken varieties and has adopted certain new constructions and vocabulary from the spoken varieties. Much of the new vocabulary
5382-589: The Arab world . The ISO assigns language codes to 32 varieties of Arabic , including its standard form of Literary Arabic, known as Modern Standard Arabic , which is derived from Classical Arabic . This distinction exists primarily among Western linguists; Arabic speakers themselves generally do not distinguish between Modern Standard Arabic and Classical Arabic, but rather refer to both as al-ʿarabiyyatu l-fuṣḥā ( اَلعَرَبِيَّةُ ٱلْفُصْحَىٰ "the eloquent Arabic") or simply al-fuṣḥā ( اَلْفُصْحَىٰ ). Arabic
5499-490: The Canary Islands . The Asian houbara or MacQueen's bustard which was earlier included in this species occurs east of the Sinai Peninsula. The North African species is sedentary unlike the migratory northern populations of MacQueen's bustards. The subspecies fuertaventurae of the Canary Islands is highly restricted and endangered. A 1997 survey found a total population of about 500 birds. The African houbara has
5616-568: The Xth form , or جامعة jāmiʻah 'university', based on جمع jamaʻa 'to gather, unite'; جمهورية jumhūriyyah 'republic', based on جمهور jumhūr 'multitude'). An earlier tendency was to redefine an older word although this has fallen into disuse (e.g., هاتف hātif 'telephone' < 'invisible caller (in Sufism)'; جريدة jarīdah 'newspaper' < 'palm-leaf stalk'). Colloquial or dialectal Arabic refers to
5733-417: The cone scales as they develop in some species of conifer . Angiosperm (flowering plants) seeds consist of three genetically distinct constituents: (1) the embryo formed from the zygote, (2) the endosperm, which is normally triploid, (3) the seed coat from tissue derived from the maternal tissue of the ovule. In angiosperms, the process of seed development begins with double fertilization , which involves
5850-412: The embryo , dispersal to a new location, and dormancy during unfavorable conditions. Seeds fundamentally are means of reproduction, and most seeds are the product of sexual reproduction which produces a remixing of genetic material and phenotype variability on which natural selection acts. Plant seeds hold endophytic microorganisms that can perform various functions, the most important of which
5967-421: The flowering plants , the ovary ripens into a fruit which contains the seed and serves to disseminate it. Many structures commonly referred to as "seeds" are actually dry fruits. Sunflower seeds are sometimes sold commercially while still enclosed within the hard wall of the fruit, which must be split open to reach the seed. Different groups of plants have other modifications, the so-called stone fruits (such as
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#17327726223896084-424: The legumes (such as beans and peas ), trees such as the oak and walnut , vegetables such as squash and radish , and sunflowers . According to Bewley and Black (1978), Brazil nut storage is in hypocotyl and this place of storage is uncommon among seeds. All gymnosperm seeds are albuminous. The seed coat develops from the maternal tissue, the integuments , originally surrounding the ovule. The seed coat in
6201-494: The northern Hejaz . These features are evidence of common descent from a hypothetical ancestor , Proto-Arabic . The following features of Proto-Arabic can be reconstructed with confidence: On the other hand, several Arabic varieties are closer to other Semitic languages and maintain features not found in Classical Arabic, indicating that these varieties cannot have developed from Classical Arabic. Thus, Arabic vernaculars do not descend from Classical Arabic: Classical Arabic
6318-416: The peach ) have a hardened fruit layer (the endocarp ) fused to and surrounding the actual seed. Nuts are the one-seeded, hard-shelled fruit of some plants with an indehiscent seed, such as an acorn or hazelnut . The first land plants evolved around 468 million years ago, and reproduced using spores. The earliest seed bearing plants to appear were the gymnosperms , which have no ovaries to contain
6435-419: The "learned" tradition (Classical Arabic). This variety and both its classicizing and "lay" iterations have been termed Middle Arabic in the past, but they are thought to continue an Old Higazi register. It is clear that the orthography of the Quran was not developed for the standardized form of Classical Arabic; rather, it shows the attempt on the part of writers to record an archaic form of Old Higazi. In
6552-710: The "purest," most eloquent form of Arabic—initiating a process of jamʿu‿l-luɣah ( جمع اللغة 'compiling the language') which took place over the 8th and early 9th centuries. Kitāb al-'Ayn ( c. 8th century ), attributed to Al-Khalil ibn Ahmad al-Farahidi , is considered the first lexicon to include all Arabic roots ; it sought to exhaust all possible root permutations —later called taqālīb ( تقاليب ) — calling those that are actually used mustaʿmal ( مستعمَل ) and those that are not used muhmal ( مُهمَل ). Lisān al-ʿArab (1290) by Ibn Manzur gives 9,273 roots, while Tāj al-ʿArūs (1774) by Murtada az-Zabidi gives 11,978 roots. Seeds In botany ,
6669-454: The 11th and 12th centuries in al-Andalus , the zajal and muwashah poetry forms developed in the dialectical Arabic of Cordoba and the Maghreb. The Nahda was a cultural and especially literary renaissance of the 19th century in which writers sought "to fuse Arabic and European forms of expression." According to James L. Gelvin , " Nahda writers attempted to simplify the Arabic language and script so that it might be accessible to
6786-571: The 4th to the 6th centuries, the Nabataean script evolved into the Arabic script recognizable from the early Islamic era. There are inscriptions in an undotted, 17-letter Arabic script dating to the 6th century CE, found at four locations in Syria ( Zabad , Jebel Usays , Harran , Umm el-Jimal ). The oldest surviving papyrus in Arabic dates to 643 CE, and it uses dots to produce the modern 28-letter Arabic alphabet. The language of that papyrus and of
6903-834: The 8th century, knowledge of Classical Arabic had become an essential prerequisite for rising into the higher classes throughout the Islamic world, both for Muslims and non-Muslims. For example, Maimonides , the Andalusi Jewish philosopher, authored works in Judeo-Arabic —Arabic written in Hebrew script . Ibn Jinni of Mosul , a pioneer in phonology , wrote prolifically in the 10th century on Arabic morphology and phonology in works such as Kitāb Al-Munṣif , Kitāb Al-Muḥtasab , and Kitāb Al-Khaṣāʾiṣ [ ar ] . Ibn Mada' of Cordoba (1116–1196) realized
7020-412: The Middle East and North Africa have become a badge of sophistication and modernity and ... feigning, or asserting, weakness or lack of facility in Arabic is sometimes paraded as a sign of status, class, and perversely, even education through a mélange of code-switching practises." Arabic has been taught worldwide in many elementary and secondary schools, especially Muslim schools. Universities around
7137-690: The Qur'an is referred to by linguists as "Quranic Arabic", as distinct from its codification soon thereafter into " Classical Arabic ". In late pre-Islamic times, a transdialectal and transcommunal variety of Arabic emerged in the Hejaz , which continued living its parallel life after literary Arabic had been institutionally standardized in the 2nd and 3rd century of the Hijra , most strongly in Judeo-Christian texts, keeping alive ancient features eliminated from
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#17327726223897254-576: The Romance languages. Also, while it is comprehensible to people from the Maghreb , a linguistically innovative variety such as Moroccan Arabic is essentially incomprehensible to Arabs from the Mashriq , much as French is incomprehensible to Spanish or Italian speakers but relatively easily learned by them. This suggests that the spoken varieties may linguistically be considered separate languages. With
7371-636: The cells are filled with starch , as for instance cereal grains , or not (non-farinaceous). The endosperm may also be referred to as "fleshy" or "cartilaginous" with thicker soft cells such as coconut , but may also be oily as in Ricinus (castor oil), Croton and Poppy . The endosperm is called "horny" when the cell walls are thicker such as date and coffee , or "ruminated" if mottled, as in nutmeg , palms and Annonaceae . In most monocotyledons (such as grasses and palms ) and some ( endospermic or albuminous ) dicotyledons (such as castor beans )
7488-405: The cells of the outer epidermis enlarge radially and their walls thicken, with nucleus and cytoplasm compressed into the outer layer. these cells which are broader on their inner surface are called palisade cells. In the inner epidermis, the cells also enlarge radially with plate like thickening of the walls. The mature inner integument has a palisade layer, a pigmented zone with 15–20 layers, while
7605-574: The conversion of Semitic mimation to nunation in the singular. It is best reassessed as a separate language on the Central Semitic dialect continuum. It was also thought that Old Arabic coexisted alongside—and then gradually displaced— epigraphic Ancient North Arabian (ANA), which was theorized to have been the regional tongue for many centuries. ANA, despite its name, was considered a very distinct language, and mutually unintelligible, from "Arabic". Scholars named its variant dialects after
7722-407: The early 19th century, influential works being: Angiosperm seeds are "enclosed seeds", produced in a hard or fleshy structure called a fruit that encloses them for protection. Some fruits have layers of both hard and fleshy material. In gymnosperms, no special structure develops to enclose the seeds, which begin their development "naked" on the bracts of cones. However, the seeds do become covered by
7839-440: The elaiosomes are eaten. The remainder of the seed, which is hard and inedible to the ants, then germinates either within the nest or at a removal site where the seed has been discarded by the ants. This dispersal relationship is an example of mutualism , since the plants depend upon the ants to disperse seeds, while the ants depend upon the plants seeds for food. As a result, a drop in numbers of one partner can reduce success of
7956-429: The embryo is embedded in the endosperm (and nucellus), which the seedling will use upon germination . In the non-endospermic dicotyledons the endosperm is absorbed by the embryo as the latter grows within the developing seed, and the cotyledons of the embryo become filled with stored food. At maturity, seeds of these species have no endosperm and are also referred to as exalbuminous seeds. The exalbuminous seeds include
8073-587: The emergence of Central Semitic languages, particularly in grammar. Innovations of the Central Semitic languages—all maintained in Arabic—include: There are several features which Classical Arabic, the modern Arabic varieties, as well as the Safaitic and Hismaic inscriptions share which are unattested in any other Central Semitic language variety, including the Dadanitic and Taymanitic languages of
8190-728: The eve of the conquests: Northern and Central (Al-Jallad 2009). The modern dialects emerged from a new contact situation produced following the conquests. Instead of the emergence of a single or multiple koines, the dialects contain several sedimentary layers of borrowed and areal features, which they absorbed at different points in their linguistic histories. According to Veersteegh and Bickerton, colloquial Arabic dialects arose from pidginized Arabic formed from contact between Arabs and conquered peoples. Pidginization and subsequent creolization among Arabs and arabized peoples could explain relative morphological and phonological simplicity of vernacular Arabic compared to Classical and MSA. In around
8307-607: The fact that they participate in the innovations common to all forms of Arabic. The earliest attestation of continuous Arabic text in an ancestor of the modern Arabic script are three lines of poetry by a man named Garm(')allāhe found in En Avdat, Israel , and dated to around 125 CE. This is followed by the Namara inscription , an epitaph of the Lakhmid king Imru' al-Qays bar 'Amro, dating to 328 CE, found at Namaraa, Syria. From
8424-452: The fertilised ovule, an immature plant from which a new plant will grow under proper conditions. The embryo has one cotyledon or seed leaf in monocotyledons , two cotyledons in almost all dicotyledons and two or more in gymnosperms. In the fruit of grains (caryopses) the single monocotyledon is shield shaped and hence called a scutellum . The scutellum is pressed closely against the endosperm from which it absorbs food and passes it to
8541-645: The first few years of their lives deriving energy from the fungi and do not produce green leaves. At up to 55 pounds (25 kilograms) the largest seed is the coco de mer (Lodoicea maldivica). This indicates a 25 Billion fold difference in seed weight. Plants that produce smaller seeds can generate many more seeds per flower, while plants with larger seeds invest more resources into those seeds and normally produce fewer seeds. Small seeds are quicker to ripen and can be dispersed sooner, so autumn all blooming plants often have small seeds. Many annual plants produce great quantities of smaller seeds; this helps to ensure at least
8658-510: The fourth most useful language for business, after English, Mandarin Chinese , and French. Arabic is written with the Arabic alphabet , an abjad script that is written from right to left . Arabic is usually classified as a Central Semitic language . Linguists still differ as to the best classification of Semitic language sub-groups. The Semitic languages changed between Proto-Semitic and
8775-507: The fruits achenes , caryopses , nuts , samaras , and utricles . Other seeds are enclosed in fruit structures that aid wind dispersal in similar ways: Myrmecochory is the dispersal of seeds by ants . Foraging ants disperse seeds which have appendages called elaiosomes (e.g. bloodroot , trilliums , acacias , and many species of Proteaceae ). Elaiosomes are soft, fleshy structures that contain nutrients for animals that eat them. The ants carry such seeds back to their nest, where
8892-408: The fusion of two male gametes with the egg cell and the central cell to form the primary endosperm and the zygote. Right after fertilization, the zygote is mostly inactive, but the primary endosperm divides rapidly to form the endosperm tissue. This tissue becomes the food the young plant will consume until the roots have developed after germination . After fertilization, the ovules develop into
9009-446: The grasses, are not distinct structures, but are fused with the fruit wall to form a pericarp .) The testae of both monocots and dicots are often marked with patterns and textured markings, or have wings or tufts of hair. When the seed coat forms from only one layer, it is also called the testa, though not all such testae are homologous from one species to the next. The funiculus abscisses (detaches at fixed point – abscission zone),
9126-431: The growing parts. Embryo descriptors include small, straight, bent, curved, and curled. Within the seed, there usually is a store of nutrients for the seedling that will grow from the embryo. The form of the stored nutrition varies depending on the kind of plant. In angiosperms, the stored food begins as a tissue called the endosperm , which is derived from the mother plant and the pollen via double fertilization . It
9243-597: The inclusion of new words into their published standard dictionaries. They also publish old and historical Arabic manuscripts. In 1997, a bureau of Arabization standardization was added to the Educational, Cultural, and Scientific Organization of the Arab League . These academies and organizations have worked toward the Arabization of the sciences, creating terms in Arabic to describe new concepts, toward
9360-434: The inner epidermis may remain a single layer, it may also divide to produce two to three layers and accumulates starch, and is referred to as the colourless layer. By contrast, the outer epidermis becomes tanniferous . The inner integument may consist of eight to fifteen layers. As the cells enlarge, and starch is deposited in the outer layers of the pigmented zone below the outer epidermis, this zone begins to lignify, while
9477-444: The innermost layer is known as the fringe layer. In gymnosperms, which do not form ovaries, the ovules and hence the seeds are exposed. This is the basis for their nomenclature – naked seeded plants. Two sperm cells transferred from the pollen do not develop the seed by double fertilization, but one sperm nucleus unites with the egg nucleus and the other sperm is not used. Sometimes each sperm fertilizes an egg cell and one zygote
9594-613: The language. Software and books with tapes are an important part of Arabic learning, as many of Arabic learners may live in places where there are no academic or Arabic language school classes available. Radio series of Arabic language classes are also provided from some radio stations. A number of websites on the Internet provide online classes for all levels as a means of distance education; most teach Modern Standard Arabic, but some teach regional varieties from numerous countries. The tradition of Arabic lexicography extended for about
9711-604: The late 6th century AD, a relatively uniform intertribal "poetic koine" distinct from the spoken vernaculars developed based on the Bedouin dialects of Najd , probably in connection with the court of al-Ḥīra . During the first Islamic century, the majority of Arabic poets and Arabic-writing persons spoke Arabic as their mother tongue. Their texts, although mainly preserved in far later manuscripts, contain traces of non-standardized Classical Arabic elements in morphology and syntax. Abu al-Aswad al-Du'ali ( c. 603 –689)
9828-424: The latter example these hairs are the source of the textile crop cotton . Other seed appendages include the raphe (a ridge), wings, caruncles (a soft spongy outgrowth from the outer integument in the vicinity of the micropyle), spines, or tubercles. A scar also may remain on the seed coat, called the hilum , where the seed was attached to the ovary wall by the funicle. Just below it is a small pore, representing
9945-420: The latter is taught in formal education settings. However, there have been studies reporting some degree of comprehension of stories told in the standard variety among preschool-aged children. The relation between Modern Standard Arabic and these dialects is sometimes compared to that of Classical Latin and Vulgar Latin vernaculars (which became Romance languages ) in medieval and early modern Europe. MSA
10062-465: The long wings show large areas of black and brown on the flight feathers. The sexes are similar, but the female, between 55–65 cm (22–26 in) tall, is rather smaller and greyer above than the male, at 65–75 cm (26–30 in) tall. The body mass is 1.8–3.2 kg (4.0–7.1 lb) in males and 1.2–1.7 kg (2.6–3.7 lb) in females. Psophia undulata was the scientific name proposed by Joseph Franz von Jacquin in 1784 who described
10179-402: The main area of growth of the embryo, while the lower or micropylar pole produces the stalk-like suspensor that attaches to the micropyle. The suspensor absorbs and manufactures nutrients from the endosperm that are used during the embryo's growth. The main components of the embryo are: Monocotyledonous plants have two additional structures in the form of sheaths. The plumule is covered with
10296-883: The many national or regional varieties which constitute the everyday spoken language. Colloquial Arabic has many regional variants; geographically distant varieties usually differ enough to be mutually unintelligible , and some linguists consider them distinct languages. However, research indicates a high degree of mutual intelligibility between closely related Arabic variants for native speakers listening to words, sentences, and texts; and between more distantly related dialects in interactional situations. The varieties are typically unwritten. They are often used in informal spoken media, such as soap operas and talk shows , as well as occasionally in certain forms of written media such as poetry and printed advertising. Hassaniya Arabic , Maltese , and Cypriot Arabic are only varieties of modern Arabic to have acquired official recognition. Hassaniya
10413-422: The mature seed can be a paper-thin layer (e.g. peanut ) or something more substantial (e.g. thick and hard in honey locust and coconut ), or fleshy as in the sarcotesta of pomegranate . The seed coat helps protect the embryo from mechanical injury, predators, and drying out. Depending on its development, the seed coat is either bitegmic or unitegmic . Bitegmic seeds form a testa from the outer integument and
10530-514: The mechanical layer is the endotegmen, then the seed is endotestal. The exotesta may consist of one or more rows of cells that are elongated and pallisade like (e.g. Fabaceae ), hence 'palisade exotesta'. In addition to the three basic seed parts, some seeds have an appendage, an aril , a fleshy outgrowth of the funicle ( funiculus ), (as in yew and nutmeg ) or an oily appendage, an elaiosome (as in Corydalis ), or hairs (trichomes). In
10647-456: The micropyle of the ovule. Seeds are very diverse in size. The dust-like orchid seeds are the smallest, with about one million seeds per gram; they are often embryonic seeds with immature embryos and no significant energy reserves. Orchids and a few other groups of plants are mycoheterotrophs which depend on mycorrhizal fungi for nutrition during germination and the early growth of the seedling. Some terrestrial orchid seedlings, in fact, spend
10764-399: The monocotyledons, ten in the dicotyledons, and two in the gymnosperms (linear and spatulate). This classification is based on three characteristics: embryo morphology, amount of endosperm and the position of the embryo relative to the endosperm. In endospermic seeds, there are two distinct regions inside the seed coat, an upper and larger endosperm and a lower smaller embryo. The embryo is
10881-782: The need for a lexical injection in Arabic, to suit concepts of the industrial and post-industrial age (such as sayyārah سَيَّارَة 'automobile' or bākhirah باخِرة 'steamship'). In response, a number of Arabic academies modeled after the Académie française were established with the aim of developing standardized additions to the Arabic lexicon to suit these transformations, first in Damascus (1919), then in Cairo (1932), Baghdad (1948), Rabat (1960), Amman (1977), Khartum [ ar ] (1993), and Tunis (1993). They review language development, monitor new words and approve
10998-424: The one whose sound is heard but whose person remains unseen. Now the term al-hatif is used for a telephone. Therefore, the process of tawleed can express the needs of modern civilization in a manner that would appear to be originally Arabic. In the case of Arabic, educated Arabs of any nationality can be assumed to speak both their school-taught Standard Arabic as well as their native dialects, which depending on
11115-403: The optimal conditions for survival of the resulting seedling; the second is spreading germination of a batch of seeds over time so a catastrophe (e.g. late frosts, drought, herbivory ) does not result in the death of all offspring of a plant ( bet-hedging ). Seed dormancy is defined as a seed failing to germinate under environmental conditions optimal for germination, normally when the environment
11232-477: The other. In South Africa , the Argentine ant ( Linepithema humile ) has invaded and displaced native species of ants. Unlike the native ant species, Argentine ants do not collect the seeds of Mimetes cucullatus or eat the elaiosomes. In areas where these ants have invaded, the numbers of Mimetes seedlings have dropped. Seed dormancy has two main functions: the first is synchronizing germination with
11349-549: The overhaul of Arabic grammar first proposed by Al-Jahiz 200 years prior. The Maghrebi lexicographer Ibn Manzur compiled Lisān al-ʿArab ( لسان العرب , "Tongue of Arabs"), a major reference dictionary of Arabic, in 1290. Charles Ferguson 's koine theory claims that the modern Arabic dialects collectively descend from a single military koine that sprang up during the Islamic conquests; this view has been challenged in recent times. Ahmad al-Jallad proposes that there were at least two considerably distinct types of Arabic on
11466-410: The parent. The large, heavy root allows the seed to penetrate into the ground when it falls. Many garden plant seeds will germinate readily as soon as they have water and are warm enough; though their wild ancestors may have had dormancy, these cultivated plants lack it. After many generations of selective pressure by plant breeders and gardeners, dormancy has been selected out. For annuals , seeds are
11583-455: The plumule and radicle, acting as a protective covering. The maturing ovule undergoes marked changes in the integuments, generally a reduction and disorganization but occasionally a thickening. The seed coat forms from the two integuments or outer layers of cells of the ovule, which derive from tissue from the mother plant, the inner integument forms the tegmen and the outer forms the testa . (The seed coats of some monocotyledon plants, such as
11700-405: The rate of germination. This is given as a percent of germination over a certain amount of time, 90% germination in 20 days, for example. 'Dormancy' is covered above; many plants produce seeds with varying degrees of dormancy, and different seeds from the same fruit can have different degrees of dormancy. It's possible to have seeds with no dormancy if they are dispersed right away and do not dry (if
11817-410: The region may be mutually unintelligible. Some of these dialects can be considered to constitute separate languages which may have "sub-dialects" of their own. When educated Arabs of different dialects engage in conversation (for example, a Moroccan speaking with a Lebanese), many speakers code-switch back and forth between the dialectal and standard varieties of the language, sometimes even within
11934-458: The same sentence. The issue of whether Arabic is one language or many languages is politically charged, in the same way it is for the varieties of Chinese , Hindi and Urdu , Serbian and Croatian , Scots and English, etc. In contrast to speakers of Hindi and Urdu who claim they cannot understand each other even when they can, speakers of the varieties of Arabic will claim they can all understand each other even when they cannot. While there
12051-415: The scar forming an oval depression, the hilum . Anatropous ovules have a portion of the funiculus that is adnate (fused to the seed coat), and which forms a longitudinal ridge, or raphe , just above the hilum. In bitegmic ovules (e.g. Gossypium described here) both inner and outer integuments contribute to the seed coat formation. With continuing maturation the cells enlarge in the outer integument. While
12168-413: The seed affects its health and germination ability: since the seed has a living embryo, over time cells die and cannot be replaced. Some seeds can live for a long time before germination, while others can only survive for a short period after dispersal before they die. Seed vigor is a measure of the quality of seed, and involves the viability of the seed, the germination percentage, germination rate, and
12285-405: The seeds. The ovule consists of a number of components: The shape of the ovules as they develop often affects the final shape of the seeds. Plants generally produce ovules of four shapes: the most common shape is called anatropous , with a curved shape. Orthotropous ovules are straight with all the parts of the ovule lined up in a long row producing an uncurved seed. Campylotropous ovules have
12402-507: The seeds. They arose during the late Devonian period (416 million to 358 million years ago). From these early gymnosperms, seed ferns evolved during the Carboniferous period (359 to 299 million years ago); they had ovules that were borne in a cupule, which consisted of groups of enclosing branches likely used to protect the developing seed. Published literature about seed storage, viability and its hygrometric dependence began in
12519-420: The soil surface is the next phase of the plant's growth and is called seedling establishment. Three fundamental conditions must exist before germination can occur. (1) The embryo must be alive, called seed viability. (2) Any dormancy requirements that prevent germination must be overcome. (3) The proper environmental conditions must exist for germination. Far red light can prevent germination. Seed viability
12636-458: The sole example of Medieval linguist Abu Hayyan al-Gharnati – who, while a scholar of the Arabic language, was not ethnically Arab – Medieval scholars of the Arabic language made no efforts at studying comparative linguistics, considering all other languages inferior. In modern times, the educated upper classes in the Arab world have taken a nearly opposite view. Yasir Suleiman wrote in 2011 that "studying and knowing English or French in most of
12753-563: The standardization of these new terms throughout the Arabic-speaking world, and toward the development of Arabic as a world language . This gave rise to what Western scholars call Modern Standard Arabic. From the 1950s, Arabization became a postcolonial nationalist policy in countries such as Tunisia, Algeria, Morocco, and Sudan. Arabic usually refers to Standard Arabic, which Western linguists divide into Classical Arabic and Modern Standard Arabic. It could also refer to any of
12870-507: The strength of the seedlings produced. The germination percentage is simply the proportion of seeds that germinate from all seeds subject to the right conditions for growth. The germination rate is the length of time it takes for the seeds to germinate. Germination percentages and rates are affected by seed viability, dormancy and environmental effects that impact on the seed and seedling. In agriculture and horticulture quality seeds have high viability, measured by germination percentage plus
12987-501: The towns where the inscriptions were discovered (Dadanitic, Taymanitic, Hismaic, Safaitic). However, most arguments for a single ANA language or language family were based on the shape of the definite article, a prefixed h-. It has been argued that the h- is an archaism and not a shared innovation, and thus unsuitable for language classification, rendering the hypothesis of an ANA language family untenable. Safaitic and Hismaic, previously considered ANA, should be considered Old Arabic due to
13104-404: The two decades before 2004, but have been increasing since. The International Fund for Houbara Conservation developed and implemented a global conservation strategy over the past forty years with the objective of ensuring a sustainable future in the wild through conservation programmes and management plans. This strategy consists of an integrated approach combining ecology, protection measures in
13221-478: The wild, conservation breeding, and reinforcement programmes. Arabic language Arabic (endonym: اَلْعَرَبِيَّةُ , romanized : al-ʿarabiyyah , pronounced [al ʕaraˈbijːa] , or عَرَبِيّ , ʿarabīy , pronounced [ˈʕarabiː] or [ʕaraˈbij] ) is a Central Semitic language of the Afroasiatic language family spoken primarily in
13338-451: The world have classes that teach Arabic as part of their foreign languages , Middle Eastern studies , and religious studies courses. Arabic language schools exist to assist students to learn Arabic outside the academic world. There are many Arabic language schools in the Arab world and other Muslim countries. Because the Quran is written in Arabic and all Islamic terms are in Arabic, millions of Muslims (both Arab and non-Arab) study
13455-457: The zygote and grows within the mother plant to a certain size before growth is halted. The formation of the seed is the defining part of the process of reproduction in seed plants ( spermatophytes ). Other plants such as ferns , mosses and liverworts , do not have seeds and use water-dependent means to propagate themselves. Seed plants now dominate biological niches on land, from forests to grasslands both in hot and cold climates . In
13572-437: Was long regarded a subspecies of the African houbara. It was proposed as a distinct species in 2003 because of differences in plumage, vocalizations and courtship behaviour. The British Ornithologists' Union's Taxonomic Records Committee 's decision to accept this split has been questioned on the grounds that the differences in the male courtship displays may be functionally trivial, and would not prevent interbreeding, whereas
13689-426: Was referred to as albumen , and the inner endosperm layer as vitellus. Although misleading, the term began to be applied to all the nutrient matter. This terminology persists in referring to endospermic seeds as "albuminous". The nature of this material is used in both describing and classifying seeds, in addition to the embryo to endosperm size ratio. The endosperm may be considered to be farinaceous (or mealy) in which
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