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Hardy Plant Society

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The Hardy Plant Society is a British charity that promotes the cultivation of hardy herbaceous plants . The society was founded in 1957 by a group of gardeners and nurserymen . It has approximately 7,500 members and provides information about familiar and less well known perennials , how to grow them and where they may be obtained. The society also works towards ensuring that garden-worthy perennial plants remain in cultivation and have the widest distribution. Its president is Roy Lancaster .

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73-428: The society aims to inform and encourage the novice gardener, stimulate and enlighten the more knowledgeable, and entertain and enthuse all gardeners bonded by a love for, and an interest in, hardy perennial plants. Throughout the year there is a wide range of events hosted by local group members, from plant study days to residential weekends to garden visits. The society exhibits at local and national shows throughout

146-516: A decline in basic activity level and information usage at 1000 ppm, when compared to 500 ppm. However a review of the literature found that a reliable subset of studies on the phenomenon of carbon dioxide induced cognitive impairment to only show a small effect on high-level decision making (for concentrations below 5000 ppm). Most of the studies were confounded by inadequate study designs, environmental comfort, uncertainties in exposure doses and differing cognitive assessments used. Similarly

219-446: A fixed structure. However, in a Coulomb explosion imaging experiment, an instantaneous image of the molecular structure can be deduced. Such an experiment has been performed for carbon dioxide. The result of this experiment, and the conclusion of theoretical calculations based on an ab initio potential energy surface of the molecule, is that none of the molecules in the gas phase are ever exactly linear. This counter-intuitive result

292-611: A glass state similar to other members of its elemental family, like silicon dioxide (silica glass) and germanium dioxide . Unlike silica and germania glasses, however, carbonia glass is not stable at normal pressures and reverts to gas when pressure is released. At temperatures and pressures above the critical point, carbon dioxide behaves as a supercritical fluid known as supercritical carbon dioxide . Table of thermal and physical properties of saturated liquid carbon dioxide: Table of thermal and physical properties of carbon dioxide (CO 2 ) at atmospheric pressure: Carbon dioxide

365-491: A mature forest will produce as much CO 2 from respiration and decomposition of dead specimens (e.g., fallen branches) as is used in photosynthesis in growing plants. Contrary to the long-standing view that they are carbon neutral, mature forests can continue to accumulate carbon and remain valuable carbon sinks , helping to maintain the carbon balance of Earth's atmosphere. Additionally, and crucially to life on earth, photosynthesis by phytoplankton consumes dissolved CO 2 in

438-428: A much larger denominator and a much smaller value than the true K a1 . The bicarbonate ion is an amphoteric species that can act as an acid or as a base, depending on pH of the solution. At high pH, it dissociates significantly into the carbonate ion ( CO 2− 3 ): In organisms, carbonic acid production is catalysed by the enzyme known as carbonic anhydrase . In addition to altering its acidity,

511-551: A protective response. The first such plant receptors were identified in rice and in Arabidopsis thaliana . Plants have some of the largest genomes of all organisms. The largest plant genome (in terms of gene number) is that of wheat ( Triticum aestivum ), predicted to encode ≈94,000 genes and thus almost 5 times as many as the human genome . The first plant genome sequenced was that of Arabidopsis thaliana which encodes about 25,500 genes. In terms of sheer DNA sequence,

584-420: A range of physical and biotic stresses which cause DNA damage , but they can tolerate and repair much of this damage. Plants reproduce to generate offspring, whether sexually , involving gametes , or asexually , involving ordinary growth. Many plants use both mechanisms. When reproducing sexually, plants have complex lifecycles involving alternation of generations . One generation, the sporophyte , which

657-406: A result of a generous legacy to the society from the estate of Kenneth Black. Plant See text Plants are the eukaryotes that form the kingdom Plantae ; they are predominantly photosynthetic . This means that they obtain their energy from sunlight , using chloroplasts derived from endosymbiosis with cyanobacteria to produce sugars from carbon dioxide and water, using

730-416: A study on the effects of the concentration of CO 2 in motorcycle helmets has been criticized for having dubious methodology in not noting the self-reports of motorcycle riders and taking measurements using mannequins. Further when normal motorcycle conditions were achieved (such as highway or city speeds) or the visor was raised the concentration of CO 2 declined to safe levels (0.2%). Poor ventilation

803-489: A twice-yearly journal, The Hardy Plant , with contributions from leading UK and international plant enthusiasts. It contains articles on hardy perennials and gardens as well as contributions from members. A newsletter is circulated three times a year that provides information on all the society's events, activities, interests and group contacts. The society publishes a series of booklets on a range of hardy plants including Geraniums, Peonies and Hostas. These can be ordered via

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876-467: A typical single C–O bond, and shorter than most other C–O multiply bonded functional groups such as carbonyls . Since it is centrosymmetric, the molecule has no electric dipole moment . As a linear triatomic molecule, CO 2 has four vibrational modes as shown in the diagram. In the symmetric and the antisymmetric stretching modes, the atoms move along the axis of the molecule. There are two bending modes, which are degenerate , meaning that they have

949-485: A waste product. In turn, oxygen is consumed and CO 2 is released as waste by all aerobic organisms when they metabolize organic compounds to produce energy by respiration . CO 2 is released from organic materials when they decay or combust, such as in forest fires. When carbon dioxide dissolves in water, it forms carbonate and mainly bicarbonate ( HCO − 3 ), which causes ocean acidification as atmospheric CO 2 levels increase. Carbon dioxide

1022-469: Is Emiliania huxleyi whose calcite scales have formed the basis of many sedimentary rocks such as limestone , where what was previously atmospheric carbon can remain fixed for geological timescales. Plants can grow as much as 50% faster in concentrations of 1,000 ppm CO 2 when compared with ambient conditions, though this assumes no change in climate and no limitation on other nutrients. Elevated CO 2 levels cause increased growth reflected in

1095-404: Is diploid (with 2 sets of chromosomes ), gives rise to the next generation, the gametophyte , which is haploid (with one set of chromosomes). Some plants also reproduce asexually via spores . In some non-flowering plants such as mosses, the sexual gametophyte forms most of the visible plant. In seed plants (gymnosperms and flowering plants), the sporophyte forms most of the visible plant, and

1168-419: Is 304.128(15) K (30.978(15) °C) at 7.3773(30) MPa (72.808(30) atm). Another form of solid carbon dioxide observed at high pressure is an amorphous glass-like solid. This form of glass, called carbonia , is produced by supercooling heated CO 2 at extreme pressures (40–48  GPa , or about 400,000 atmospheres) in a diamond anvil . This discovery confirmed the theory that carbon dioxide could exist in

1241-569: Is 53% more dense than dry air, but is long lived and thoroughly mixes in the atmosphere. About half of excess CO 2 emissions to the atmosphere are absorbed by land and ocean carbon sinks . These sinks can become saturated and are volatile, as decay and wildfires result in the CO 2 being released back into the atmosphere. CO 2 is eventually sequestered (stored for the long term) in rocks and organic deposits like coal , petroleum and natural gas . Nearly all CO2 produced by humans goes into

1314-406: Is a chemical compound with the chemical formula CO 2 . It is made up of molecules that each have one carbon atom covalently double bonded to two oxygen atoms. It is found in the gas state at room temperature and at normally-encountered concentrations it is odorless. As the source of carbon in the carbon cycle , atmospheric CO 2 is the primary carbon source for life on Earth. In

1387-547: Is a similar process. Structures such as runners enable plants to grow to cover an area, forming a clone . Many plants grow food storage structures such as tubers or bulbs which may each develop into a new plant. Some non-flowering plants, such as many liverworts, mosses and some clubmosses, along with a few flowering plants, grow small clumps of cells called gemmae which can detach and grow. Plants use pattern-recognition receptors to recognize pathogens such as bacteria that cause plant diseases. This recognition triggers

1460-415: Is an end product of cellular respiration in organisms that obtain energy by breaking down sugars, fats and amino acids with oxygen as part of their metabolism . This includes all plants, algae and animals and aerobic fungi and bacteria. In vertebrates , the carbon dioxide travels in the blood from the body's tissues to the skin (e.g., amphibians ) or the gills (e.g., fish ), from where it dissolves in

1533-402: Is commercially used in its solid form, commonly known as " dry ice ". The solid-to-gas phase transition occurs at 194.7 Kelvin and is called sublimation . The symmetry of a carbon dioxide molecule is linear and centrosymmetric at its equilibrium geometry. The length of the carbon–oxygen bond in carbon dioxide is 116.3  pm , noticeably shorter than the roughly 140 pm length of

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1606-502: Is concerned about the conservation of garden plants . Countless fine plants have totally disappeared from cultivation and remain but a memory. The society is working towards ensuring that older, rarer and lesser-known perennial plants are conserved and made available to gardeners generally. Members suggest plants which they believe are garden-worthy but hard to source and they are grown in members' gardens so their hardiness and tolerance to different growing conditions can be compared across

1679-496: Is known as botany , a branch of biology . All living things were traditionally placed into one of two groups, plants and animals . This classification dates from Aristotle (384–322 BC), who distinguished different levels of beings in his biology , based on whether living things had a "sensitive soul" or like plants only a "vegetative soul". Theophrastus , Aristotle's student, continued his work in plant taxonomy and classification. Much later, Linnaeus (1707–1778) created

1752-632: Is one of the main causes of excessive CO 2 concentrations in closed spaces, leading to poor indoor air quality . Carbon dioxide differential above outdoor concentrations at steady state conditions (when the occupancy and ventilation system operation are sufficiently long that CO 2 concentration has stabilized) are sometimes used to estimate ventilation rates per person. Higher CO 2 concentrations are associated with occupant health, comfort and performance degradation. ASHRAE Standard 62.1–2007 ventilation rates may result in indoor concentrations up to 2,100 ppm above ambient outdoor conditions. Thus if

1825-410: Is ordinarily a difficult and slow reaction: The redox potential for this reaction near pH 7 is about −0.53 V versus the standard hydrogen electrode . The nickel-containing enzyme carbon monoxide dehydrogenase catalyses this process. Photoautotrophs (i.e. plants and cyanobacteria ) use the energy contained in sunlight to photosynthesize simple sugars from CO 2 absorbed from

1898-405: Is produced as a by-product. Ribulose-1,5-bisphosphate carboxylase oxygenase , commonly abbreviated to RuBisCO, is the enzyme involved in the first major step of carbon fixation, the production of two molecules of 3-phosphoglycerate from CO 2 and ribulose bisphosphate , as shown in the diagram at left. RuBisCO is thought to be the single most abundant protein on Earth. Phototrophs use

1971-472: Is the true first acid dissociation constant, defined as where the denominator includes only covalently bound H 2 CO 3 and does not include hydrated CO 2 (aq). The much smaller and often-quoted value near 4.16 × 10 (or pK a1 = 6.38) is an apparent value calculated on the (incorrect) assumption that all dissolved CO 2 is present as carbonic acid, so that Since most of the dissolved CO 2 remains as CO 2 molecules, K a1 (apparent) has

2044-555: Is the main cause of these increased CO 2 concentrations, which are the primary cause of climate change . Its concentration in Earth's pre-industrial atmosphere since late in the Precambrian was regulated by organisms and geological features. Plants , algae and cyanobacteria use energy from sunlight to synthesize carbohydrates from carbon dioxide and water in a process called photosynthesis , which produces oxygen as

2117-419: Is trivially due to the fact that the nuclear motion volume element vanishes for linear geometries. This is so for all molecules except diatomic molecules . Carbon dioxide is soluble in water, in which it reversibly forms H 2 CO 3 (carbonic acid), which is a weak acid , because its ionization in water is incomplete. The hydration equilibrium constant of carbonic acid is, at 25 °C: Hence,

2190-465: Is used in CO 2 scrubbers and has been suggested as a possible starting point for carbon capture and storage by amine gas treating . Only very strong nucleophiles, like the carbanions provided by Grignard reagents and organolithium compounds react with CO 2 to give carboxylates : In metal carbon dioxide complexes , CO 2 serves as a ligand , which can facilitate the conversion of CO 2 to other chemicals. The reduction of CO 2 to CO

2263-558: The Antarctic flora , consisting of algae, mosses, liverworts, lichens, and just two flowering plants, have adapted to the prevailing conditions on that southern continent. Plants are often the dominant physical and structural component of the habitats where they occur. Many of the Earth's biomes are named for the type of vegetation because plants are the dominant organisms in those biomes, such as grassland , savanna , and tropical rainforest . Carbon dioxide Carbon dioxide

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2336-706: The Cretaceous so rapid that Darwin called it an " abominable mystery ". Conifers diversified from the Late Triassic onwards, and became a dominant part of floras in the Jurassic . In 2019, a phylogeny based on genomes and transcriptomes from 1,153 plant species was proposed. The placing of algal groups is supported by phylogenies based on genomes from the Mesostigmatophyceae and Chlorokybophyceae that have since been sequenced. Both

2409-457: The carpels or ovaries , which develop into fruits that contain seeds . Fruits may be dispersed whole, or they may split open and the seeds dispersed individually. Plants reproduce asexually by growing any of a wide variety of structures capable of growing into new plants. At the simplest, plants such as mosses or liverworts may be broken into pieces, each of which may regrow into whole plants. The propagation of flowering plants by cuttings

2482-751: The glaucophytes , in the clade Archaeplastida . There are about 380,000 known species of plants, of which the majority, some 260,000, produce seeds . They range in size from single cells to the tallest trees . Green plants provide a substantial proportion of the world's molecular oxygen; the sugars they create supply the energy for most of Earth's ecosystems and other organisms , including animals, either eat plants directly or rely on organisms which do so. Grain , fruit , and vegetables are basic human foods and have been domesticated for millennia. People use plants for many purposes , such as building materials , ornaments, writing materials , and, in great variety, for medicines . The scientific study of plants

2555-1003: The "chlorophyte algae" and the "streptophyte algae" are treated as paraphyletic (vertical bars beside phylogenetic tree diagram) in this analysis, as the land plants arose from within those groups. The classification of Bryophyta is supported both by Puttick et al. 2018, and by phylogenies involving the hornwort genomes that have also since been sequenced. Rhodophyta [REDACTED] Glaucophyta [REDACTED] Chlorophyta [REDACTED] Prasinococcales   Mesostigmatophyceae Chlorokybophyceae Spirotaenia [REDACTED] Klebsormidiales [REDACTED] Chara [REDACTED] Coleochaetales [REDACTED] Hornworts [REDACTED] Liverworts [REDACTED] Mosses [REDACTED] Lycophytes [REDACTED] [REDACTED] Gymnosperms [REDACTED] Angiosperms [REDACTED] Plant cells have distinctive features that other eukaryotic cells (such as those of animals) lack. These include

2628-439: The air and water: Carbon dioxide is colorless. At low concentrations, the gas is odorless; however, at sufficiently high concentrations, it has a sharp, acidic odor. At standard temperature and pressure , the density of carbon dioxide is around 1.98 kg/m , about 1.53 times that of air . Carbon dioxide has no liquid state at pressures below 0.51795(10) MPa (5.11177(99) atm ). At a pressure of 1 atm (0.101325 MPa),

2701-444: The air, carbon dioxide is transparent to visible light but absorbs infrared radiation , acting as a greenhouse gas . Carbon dioxide is soluble in water and is found in groundwater , lakes , ice caps , and seawater . It is a trace gas in Earth's atmosphere at 421  parts per million (ppm) , or about 0.042% (as of May 2022) having risen from pre-industrial levels of 280 ppm or about 0.028%. Burning fossil fuels

2774-801: The amount of cytoplasm stays the same. Most plants are multicellular . Plant cells differentiate into multiple cell types, forming tissues such as the vascular tissue with specialized xylem and phloem of leaf veins and stems , and organs with different physiological functions such as roots to absorb water and minerals, stems for support and to transport water and synthesized molecules, leaves for photosynthesis, and flowers for reproduction. Plants photosynthesize , manufacturing food molecules ( sugars ) using energy obtained from light . Plant cells contain chlorophylls inside their chloroplasts, which are green pigments that are used to capture light energy. The end-to-end chemical equation for photosynthesis is: This causes plants to release oxygen into

2847-431: The atmosphere. Green plants provide a substantial proportion of the world's molecular oxygen, alongside the contributions from photosynthetic algae and cyanobacteria. Plants that have secondarily adopted a parasitic lifestyle may lose the genes involved in photosynthesis and the production of chlorophyll. Growth is determined by the interaction of a plant's genome with its physical and biotic environment. Factors of

2920-430: The atmosphere. Less than 1% of CO2 produced annually is put to commercial use, mostly in the fertilizer industry and in the oil and gas industry for enhanced oil recovery . Other commercial applications include food and beverage production, metal fabrication, cooling, fire suppression and stimulating plant growth in greenhouses. Carbon dioxide cannot be liquefied at atmospheric pressure. Low-temperature carbon dioxide

2993-467: The basis of the modern system of scientific classification , but retained the animal and plant kingdoms , naming the plant kingdom the Vegetabilia. When the name Plantae or plant is applied to a specific group of organisms or taxa , it usually refers to one of four concepts. From least to most inclusive, these four groupings are: There are about 382,000 accepted species of plants, of which

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3066-869: The condition. There are few studies of the health effects of long-term continuous CO 2 exposure on humans and animals at levels below 1%. Occupational CO 2 exposure limits have been set in the United States at 0.5% (5000 ppm) for an eight-hour period. At this CO 2 concentration, International Space Station crew experienced headaches, lethargy, mental slowness, emotional irritation, and sleep disruption. Studies in animals at 0.5% CO 2 have demonstrated kidney calcification and bone loss after eight weeks of exposure. A study of humans exposed in 2.5 hour sessions demonstrated significant negative effects on cognitive abilities at concentrations as low as 0.1% (1000   ppm) CO 2 likely due to CO 2 induced increases in cerebral blood flow. Another study observed

3139-400: The country to stimulate interest in hardy plants. Visitors can see hardy plants in bloom and leaf in their natural season and find out more about the work of the society. An extensive library of digital images covering a wide range of plants and garden scenes is available to members and members of the public. Images have been donated by photographers in the society. The Hardy Plant Society

3212-513: The country – information that is collected in a database and reflected on the society's webpages as a resource for HPS members and the public. http://www.hardy-plant.org.uk/about-plants/conservation Every member can join in the annual Seed Distribution Scheme by obtaining or donating hardy perennial seed . The Seed List offers over 2,000 varieties of rare, unusual and familiar seeds and is sent to every member in December. National membership of

3285-517: The degenerate pair of bending modes at 667 cm (wavelength 15.0 μm). The symmetric stretching mode does not create an electric dipole so is not observed in IR spectroscopy, but it is detected in Raman spectroscopy at 1388 cm (wavelength 7.20 μm), with a Fermi resonance doublet at 1285 cm . In the gas phase, carbon dioxide molecules undergo significant vibrational motions and do not keep

3358-488: The development of forests in swampy environments dominated by clubmosses and horsetails, including some as large as trees, and the appearance of early gymnosperms , the first seed plants . The Permo-Triassic extinction event radically changed the structures of communities. This may have set the scene for the evolution of flowering plants in the Triassic (~ 200  million years ago ), with an adaptive radiation in

3431-566: The dispersing effects of wind, it can collect in sheltered/pocketed locations below average ground level, causing animals located therein to be suffocated. Carrion feeders attracted to the carcasses are then also killed. Children have been killed in the same way near the city of Goma by CO 2 emissions from the nearby volcano Mount Nyiragongo . The Swahili term for this phenomenon is mazuku . Adaptation to increased concentrations of CO 2 occurs in humans, including modified breathing and kidney bicarbonate production, in order to balance

3504-541: The effects of blood acidification ( acidosis ). Several studies suggested that 2.0 percent inspired concentrations could be used for closed air spaces (e.g. a submarine ) since the adaptation is physiological and reversible, as deterioration in performance or in normal physical activity does not happen at this level of exposure for five days. Yet, other studies show a decrease in cognitive function even at much lower levels. Also, with ongoing respiratory acidosis , adaptation or compensatory mechanisms will be unable to reverse

3577-551: The electrical conductivity of fully deionized water without CO 2 saturation is comparably low in relation to these data. CO 2 is a potent electrophile having an electrophilic reactivity that is comparable to benzaldehyde or strongly electrophilic α,β-unsaturated carbonyl compounds . However, unlike electrophiles of similar reactivity, the reactions of nucleophiles with CO 2 are thermodynamically less favored and are often found to be highly reversible. The reversible reaction of carbon dioxide with amines to make carbamates

3650-478: The fungi and some of the algae. By the definition used in this article, plants form the clade Viridiplantae (green plants), which consists of the green algae and the embryophytes or land plants ( hornworts , liverworts , mosses , lycophytes , ferns , conifers and other gymnosperms , and flowering plants ). A definition based on genomes includes the Viridiplantae, along with the red algae and

3723-409: The gametophyte is very small. Flowering plants reproduce sexually using flowers, which contain male and female parts: these may be within the same ( hermaphrodite ) flower, on different flowers on the same plant , or on different plants . The stamens create pollen , which produces male gametes that enter the ovule to fertilize the egg cell of the female gametophyte. Fertilization takes place within

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3796-469: The gas deposits directly to a solid at temperatures below 194.6855(30) K (−78.4645(30) °C) and the solid sublimes directly to a gas above this temperature. In its solid state, carbon dioxide is commonly called dry ice . Liquid carbon dioxide forms only at pressures above 0.51795(10) MPa (5.11177(99) atm); the triple point of carbon dioxide is 216.592(3) K (−56.558(3) °C) at 0.51795(10) MPa (5.11177(99) atm) (see phase diagram). The critical point

3869-583: The great majority, some 283,000, produce seeds . The table below shows some species count estimates of different green plant (Viridiplantae) divisions . About 85–90% of all plants are flowering plants. Several projects are currently attempting to collect records on all plant species in online databases, e.g. the World Flora Online . Plants range in scale from single-celled organisms such as desmids (from 10  micrometres   (μm) across) and picozoa (less than 3 μm across), to

3942-515: The green pigment chlorophyll . Exceptions are parasitic plants that have lost the genes for chlorophyll and photosynthesis, and obtain their energy from other plants or fungi. Most plants are multicellular , except for some green algae. Historically, as in Aristotle's biology , the plant kingdom encompassed all living things that were not animals , and included algae and fungi . Definitions have narrowed since then; current definitions exclude

4015-601: The harvestable yield of crops, with wheat, rice and soybean all showing increases in yield of 12–14% under elevated CO 2 in FACE experiments. Increased atmospheric CO 2 concentrations result in fewer stomata developing on plants which leads to reduced water usage and increased water-use efficiency . Studies using FACE have shown that CO 2 enrichment leads to decreased concentrations of micronutrients in crop plants. This may have knock-on effects on other parts of ecosystems as herbivores will need to eat more food to gain

4088-470: The land 1,200  million years ago , but it was not until the Ordovician , around 450  million years ago , that the first land plants appeared, with a level of organisation like that of bryophytes. However, fossils of organisms with a flattened thallus in Precambrian rocks suggest that multicellular freshwater eukaryotes existed over 1000 mya. Primitive land plants began to diversify in

4161-412: The large water-filled central vacuole , chloroplasts , and the strong flexible cell wall , which is outside the cell membrane . Chloroplasts are derived from what was once a symbiosis of a non-photosynthetic cell and photosynthetic cyanobacteria . The cell wall, made mostly of cellulose , allows plant cells to swell up with water without bursting. The vacuole allows the cell to change in size while

4234-515: The largest trees ( megaflora ) such as the conifer Sequoia sempervirens (up to 120 metres (380 ft) tall) and the angiosperm Eucalyptus regnans (up to 100 m (325 ft) tall). The naming of plants is governed by the International Code of Nomenclature for algae, fungi, and plants and the International Code of Nomenclature for Cultivated Plants . The ancestors of land plants evolved in water. An algal scum formed on

4307-665: The late Silurian , around 420  million years ago . Bryophytes, club mosses, and ferns then appear in the fossil record. Early plant anatomy is preserved in cellular detail in an early Devonian fossil assemblage from the Rhynie chert . These early plants were preserved by being petrified in chert formed in silica-rich volcanic hot springs. By the end of the Devonian, most of the basic features of plants today were present, including roots, leaves and secondary wood in trees such as Archaeopteris . The Carboniferous period saw

4380-444: The majority of the carbon dioxide is not converted into carbonic acid, but remains as CO 2 molecules, not affecting the pH. The relative concentrations of CO 2 , H 2 CO 3 , and the deprotonated forms HCO − 3 ( bicarbonate ) and CO 2− 3 ( carbonate ) depend on the pH . As shown in a Bjerrum plot , in neutral or slightly alkaline water (pH > 6.5), the bicarbonate form predominates (>50%) becoming

4453-443: The most prevalent (>95%) at the pH of seawater. In very alkaline water (pH > 10.4), the predominant (>50%) form is carbonate. The oceans, being mildly alkaline with typical pH = 8.2–8.5, contain about 120 mg of bicarbonate per liter. Being diprotic , carbonic acid has two acid dissociation constants , the first one for the dissociation into the bicarbonate (also called hydrogen carbonate) ion ( HCO − 3 ): This

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4526-501: The physical or abiotic environment include temperature , water , light, carbon dioxide , and nutrients in the soil. Biotic factors that affect plant growth include crowding, grazing, beneficial symbiotic bacteria and fungi, and attacks by insects or plant diseases . Frost and dehydration can damage or kill plants. Some plants have antifreeze proteins , heat-shock proteins and sugars in their cytoplasm that enable them to tolerate these stresses . Plants are continuously exposed to

4599-495: The presence of carbon dioxide in water also affects its electrical properties. When carbon dioxide dissolves in desalinated water, the electrical conductivity increases significantly from below 1 μS/cm to nearly 30 μS/cm. When heated, the water begins to gradually lose the conductivity induced by the presence of C O 2 {\displaystyle \mathrm {CO_{2}} } , especially noticeable as temperatures exceed 30 °C. The temperature dependence of

4672-499: The presence of sufficient oxygen, manifesting as dizziness, headache, visual and hearing dysfunction, and unconsciousness within a few minutes to an hour. Concentrations of more than 10% may cause convulsions, coma, and death. CO 2 levels of more than 30% act rapidly leading to loss of consciousness in seconds. Because it is heavier than air, in locations where the gas seeps from the ground (due to sub-surface volcanic or geothermal activity) in relatively high concentrations, without

4745-496: The products of their photosynthesis as internal food sources and as raw material for the biosynthesis of more complex organic molecules, such as polysaccharides , nucleic acids , and proteins. These are used for their own growth, and also as the basis of the food chains and webs that feed other organisms, including animals such as ourselves. Some important phototrophs, the coccolithophores synthesise hard calcium carbonate scales. A globally significant species of coccolithophore

4818-420: The same amount of protein. The concentration of secondary metabolites such as phenylpropanoids and flavonoids can also be altered in plants exposed to high concentrations of CO 2 . Plants also emit CO 2 during respiration, and so the majority of plants and algae, which use C3 photosynthesis , are only net absorbers during the day. Though a growing forest will absorb many tons of CO 2 each year,

4891-411: The same frequency and same energy, because of the symmetry of the molecule. When a molecule touches a surface or touches another molecule, the two bending modes can differ in frequency because the interaction is different for the two modes. Some of the vibrational modes are observed in the infrared (IR) spectrum : the antisymmetric stretching mode at wavenumber 2349 cm (wavelength 4.25 μm) and

4964-469: The smallest published genome is that of the carnivorous bladderwort ( Utricularia gibba) at 82 Mb (although it still encodes 28,500 genes) while the largest, from the Norway spruce ( Picea abies ), extends over 19.6 Gb (encoding about 28,300 genes). Plants are distributed almost worldwide. While they inhabit several biomes which can be divided into a multitude of ecoregions , only the hardy plants of

5037-496: The society gives members the opportunity to join a local group and take part in a range of activities. There are around 40 local groups across the UK and national members are invited to join the group nearest to them. Each group offers a range of gardening activities including lectures from gardening speakers, garden visits, plant sales and educational and social events throughout the year. Most groups produce their own newsletters . Within

5110-611: The society there are groups for members with a special interest in particular groups of plants or those which require particular growing conditions. There are special interest groups covering Galanthus, Half Hardy Plants, Hardy Geraniums, Peonies, Pulmonarias, Ranunculaceae, Shade and Woodland Plants and Variegated Plants. They produce their own newsletters and organise meetings and events for fellow enthusiasts. The Correspondents Group enables members who are unable to attend meetings to remain in touch with one another and exchange gardening ideas and information. The Hardy Plant Society produces

5183-535: The upper ocean and thereby promotes the absorption of CO 2 from the atmosphere. Carbon dioxide content in fresh air (averaged between sea-level and 10 kPa level, i.e., about 30 km (19 mi) altitude) varies between 0.036% (360 ppm) and 0.041% (412 ppm), depending on the location. In humans, exposure to CO 2 at concentrations greater than 5% causes the development of hypercapnia and respiratory acidosis . Concentrations of 7% to 10% (70,000 to 100,000 ppm) may cause suffocation, even in

5256-546: The water, or to the lungs from where it is exhaled. During active photosynthesis, plants can absorb more carbon dioxide from the atmosphere than they release in respiration. Carbon fixation is a biochemical process by which atmospheric carbon dioxide is incorporated by plants, algae and cyanobacteria into energy-rich organic molecules such as glucose , thus creating their own food by photosynthesis. Photosynthesis uses carbon dioxide and water to produce sugars from which other organic compounds can be constructed, and oxygen

5329-452: The website. http://www.hardy-plant.org.uk/publications/booklets The Hardy Plant Society offers a number of small bursaries, for students and for people who are employed in horticulture, to enable them to develop their knowledge and understanding. Grants are usually made to support travel and subsistence costs for project work or study modules on topics broadly related to the society's aims and objectives. The bursaries are made available as

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