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Life

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174-402: Life on Earth: Life is a quality that distinguishes matter that has biological processes , such as signaling and self-sustaining processes, from matter that does not. It is defined descriptively by the capacity for homeostasis , organisation , metabolism , growth , adaptation , response to stimuli , and reproduction . All life over time eventually reaches a state of death , and none

348-419: A double helix . The two DNA strands are known as polynucleotides since they are composed of simpler units called nucleotides . Each nucleotide is composed of a nitrogen-containing nucleobase —either cytosine (C), guanine (G), adenine (A), or thymine (T)—as well as a sugar called deoxyribose and a phosphate group . The nucleotides are joined to one another in a chain by covalent bonds between

522-419: A living systems theory viewpoint that does not necessarily depend on molecular chemistry. One systemic definition of life is that living things are self-organizing and autopoietic (self-producing). Variations of this include Stuart Kauffman 's definition as an autonomous agent or a multi-agent system capable of reproducing itself, and of completing at least one thermodynamic work cycle . This definition

696-428: A nucleus and other membrane-bound organelles , although they have circular DNA and ribosomes . Bacteria and Archaea are two domains of prokaryotes. The other primary type is the eukaryote cell, which has a distinct nucleus bound by a nuclear membrane and membrane-bound organelles, including mitochondria , chloroplasts , lysosomes , rough and smooth endoplasmic reticulum , and vacuoles . In addition, their DNA

870-412: A nucleus of protons and neutrons , and a surrounding "cloud" of orbiting electrons which "take up space". However, this is only somewhat correct because subatomic particles and their properties are governed by their quantum nature , which means they do not act as everyday objects appear to act – they can act like waves as well as particles , and they do not have well-defined sizes or positions. In

1044-491: A quantity of matter . As such, there is no single universally agreed scientific meaning of the word "matter". Scientifically, the term "mass" is well-defined, but "matter" can be defined in several ways. Sometimes in the field of physics "matter" is simply equated with particles that exhibit rest mass (i.e., that cannot travel at the speed of light), such as quarks and leptons. However, in both physics and chemistry , matter exhibits both wave -like and particle -like properties,

1218-401: A system and its surroundings. impermeable to matter impermeable to matter A system is enclosed by walls that bound it and connect it to its surroundings. Often a wall restricts passage across it by some form of matter or energy, making the connection indirect. Sometimes a wall is no more than an imaginary two-dimensional closed surface through which the connection to the surroundings

1392-430: A baryon, is given a baryon number of 1/3. So the net amount of matter, as measured by the number of quarks (minus the number of antiquarks, which each have a baryon number of −1/3), which is proportional to baryon number, and number of leptons (minus antileptons), which is called the lepton number, is practically impossible to change in any process. Even in a nuclear bomb, none of the baryons (protons and neutrons of which

1566-424: A body of steam or air in a steam engine , such as Sadi Carnot defined in 1824. It could also be just one nuclide (i.e. a system of quarks ) as hypothesized in quantum thermodynamics . The system is the part of the universe being studied, while the surroundings is the remainder of the universe that lies outside the boundaries of the system. It is also known as the environment or the reservoir . Depending on

1740-635: A charge of −1  e . They also carry colour charge , which is the equivalent of the electric charge for the strong interaction . Quarks also undergo radioactive decay , meaning that they are subject to the weak interaction . Baryons are strongly interacting fermions, and so are subject to Fermi–Dirac statistics. Amongst the baryons are the protons and neutrons, which occur in atomic nuclei, but many other unstable baryons exist as well. The term baryon usually refers to triquarks—particles made of three quarks. Also, "exotic" baryons made of four quarks and one antiquark are known as pentaquarks , but their existence

1914-410: A classification of thermodynamic systems according to internal processes consisting in energy redistribution (passive systems) and energy conversion (active systems). If there is a temperature difference inside the thermodynamic system, for example in a rod, one end of which is warmer than the other, then thermal energy transfer processes occur in it, in which the temperature of the colder part rises and

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2088-404: A common origin. All life forms require certain core chemical elements for their biochemical functioning. These include carbon , hydrogen , nitrogen , oxygen , phosphorus , and sulfur —the elemental macronutrients for all organisms. Together these make up nucleic acids , proteins and lipids , the bulk of living matter. Five of these six elements comprise the chemical components of DNA,

2262-527: A consequence of these microbial activities, the physical-chemical environment on Earth has been changing on a geologic time scale , thereby affecting the path of evolution of subsequent life. For example, the release of molecular oxygen by cyanobacteria as a by-product of photosynthesis induced global changes in the Earth's environment. Because oxygen was toxic to most life on Earth at the time, this posed novel evolutionary challenges, and ultimately resulted in

2436-436: A contact equilibrium with respect to that substance is possible. By suitable thermodynamic operations , the pure substance reservoir can be dealt with as a closed system. Its internal energy and its entropy can be determined as functions of its temperature, pressure, and mole number. A thermodynamic operation can render impermeable to matter all system walls other than the contact equilibrium wall for that substance. This allows

2610-623: A desired degree, the resulting substance is said to be chemically pure . Chemical substances can exist in several different physical states or phases (e.g. solids , liquids , gases , or plasma ) without changing their chemical composition. Substances transition between these phases of matter in response to changes in temperature or pressure . Some chemical substances can be combined or converted into new substances by means of chemical reactions . Chemicals that do not possess this ability are said to be inert . A definition of "matter" based on its physical and chemical structure is: matter

2784-441: A distance from other particles under everyday conditions; this creates the property of matter which appears to us as matter taking up space. For much of the history of the natural sciences , people have contemplated the exact nature of matter. The idea that matter was built of discrete building blocks, the so-called particulate theory of matter , appeared in both ancient Greece and ancient India . Early philosophers who proposed

2958-443: A few of its theoretical properties. There is considerable speculation both in science and science fiction as to why the observable universe is apparently almost entirely matter (in the sense of quarks and leptons but not antiquarks or antileptons), and whether other places are almost entirely antimatter (antiquarks and antileptons) instead. In the early universe, it is thought that matter and antimatter were equally represented, and

3132-440: A host cell to make new products. Virus self-assembly within host cells has implications for the study of the origin of life , as it may support the hypothesis that life could have started as self-assembling organic molecules . Some of the earliest theories of life were materialist, holding that all that exists is matter, and that life is merely a complex form or arrangement of matter. Empedocles (430 BC) argued that everything in

3306-415: A macroscopic scale.” Equilibrium thermodynamics, as a subject in physics, considers macroscopic bodies of matter and energy in states of internal thermodynamic equilibrium. It uses the concept of thermodynamic processes , by which bodies pass from one equilibrium state to another by transfer of matter and energy between them. The term 'thermodynamic system' is used to refer to bodies of matter and energy in

3480-471: A matter of debate; viruses lack characteristics of life such as cell membranes, metabolism and the ability to grow or respond to their environments. Viruses have been classed into "species" based on their genetics , but many aspects of such a classification remain controversial. The original Linnaean system has been modified many times, for example as follows: The attempt to organise the Eukaryotes into

3654-533: A month in a simulated Martian environment . Beyond the Solar System, the region around another main-sequence star that could support Earth-like life on an Earth-like planet is known as the habitable zone . The inner and outer radii of this zone vary with the luminosity of the star, as does the time interval during which the zone survives. Stars more massive than the Sun have a larger habitable zone, but remain on

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3828-447: A process to be reversible , each step in the process must be reversible. For a step in a process to be reversible, the system must be in equilibrium throughout the step. That ideal cannot be accomplished in practice because no step can be taken without perturbing the system from equilibrium, but the ideal can be approached by making changes slowly. The very existence of thermodynamic equilibrium, defining states of thermodynamic systems,

4002-432: A quantity is said to be permeable to it, and a thermodynamic system is classified by the permeabilities of its several walls. A transfer between system and surroundings can arise by contact, such as conduction of heat, or by long-range forces such as an electric field in the surroundings. A system with walls that prevent all transfers is said to be isolated . This is an idealized conception, because in practice some transfer

4176-451: A resurgence of interest in the work of Russell, Barge and colleagues. Hylomorphism is a theory first expressed by the Greek philosopher Aristotle (322 BC). The application of hylomorphism to biology was important to Aristotle, and biology is extensively covered in his extant writings . In this view, everything in the material universe has both matter and form, and the form of a living thing

4350-516: A sea of degenerate electrons. At a microscopic level, the constituent "particles" of matter such as protons, neutrons, and electrons obey the laws of quantum mechanics and exhibit wave–particle duality. At an even deeper level, protons and neutrons are made up of quarks and the force fields ( gluons ) that bind them together, leading to the next definition. As seen in the above discussion, many early definitions of what can be called "ordinary matter" were based upon its structure or "building blocks". On

4524-804: A set of 355 genes from the last universal common ancestor was tentatively identified. The biosphere is postulated to have developed, from the origin of life onwards, at least some 3.5 billion years ago. The earliest evidence for life on Earth includes biogenic graphite found in 3.7 billion-year-old metasedimentary rocks from Western Greenland and microbial mat fossils found in 3.48 billion-year-old sandstone from Western Australia . More recently, in 2015, "remains of biotic life " were found in 4.1 billion-year-old rocks in Western Australia. In 2017, putative fossilised microorganisms (or microfossils ) were announced to have been discovered in hydrothermal vent precipitates in

4698-618: A short period Linnaeus had classified them in the taxon Vermes in Animalia, but later placed them back in Plantae. Herbert Copeland classified the Fungi in his Protoctista , including them with single-celled organisms and thus partially avoiding the problem but acknowledging their special status. The problem was eventually solved by Whittaker , when he gave them their own kingdom in his five-kingdom system . Evolutionary history shows that

4872-465: A small number of kingdoms has been challenged. The Protozoa do not form a clade or natural grouping, and nor do the Chromista (Chromalveolata). The ability to sequence large numbers of complete genomes has allowed biologists to take a metagenomic view of the phylogeny of the whole tree of life . This has led to the realisation that the majority of living things are bacteria, and that all have

5046-403: A spark between the electrodes and initiates combustion. Heat transfer occurs across the boundary after combustion but no mass transfer takes place either way. The first law of thermodynamics for energy transfers for closed system may be stated: where U {\displaystyle U} denotes the internal energy of the system, Q {\displaystyle Q} heat added to

5220-426: A subclass of matter. A common or traditional definition of matter is "anything that has mass and volume (occupies space )". For example, a car would be said to be made of matter, as it has mass and volume (occupies space). The observation that matter occupies space goes back to antiquity. However, an explanation for why matter occupies space is recent, and is argued to be a result of the phenomenon described in

5394-490: A subject in physics, considers bodies of matter and energy that are not in states of internal thermodynamic equilibrium, but are usually participating in processes of transfer that are slow enough to allow description in terms of quantities that are closely related to thermodynamic state variables . It is characterized by presence of flows of matter and energy. For this topic, very often the bodies considered have smooth spatial inhomogeneities, so that spatial gradients, for example

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5568-406: A substance, the chemical potentials of the substance must be same on either side of the wall. This is part of the nature of thermodynamic equilibrium, and may be regarded as related to the zeroth law of thermodynamics. In an open system, there is an exchange of energy and matter between the system and the surroundings. The presence of reactants in an open beaker is an example of an open system. Here

5742-473: A temperature gradient, are well enough defined. Thus the description of non-equilibrium thermodynamic systems is a field theory, more complicated than the theory of equilibrium thermodynamics. Non-equilibrium thermodynamics is a growing subject, not an established edifice. Example theories and modeling approaches include the GENERIC formalism for complex fluids, viscoelasticity, and soft materials. In general, it

5916-476: A temperature near absolute zero. The Pauli exclusion principle requires that only two fermions can occupy a quantum state, one spin-up and the other spin-down. Hence, at zero temperature, the fermions fill up sufficient levels to accommodate all the available fermions—and in the case of many fermions, the maximum kinetic energy (called the Fermi energy ) and the pressure of the gas becomes very large, and depends on

6090-478: A thermodynamic system at a given time is described by its state , which can be specified by the values of a set of thermodynamic state variables. A thermodynamic system is in thermodynamic equilibrium when there are no macroscopically apparent flows of matter or energy within it or between it and other systems. Thermodynamic equilibrium is characterized not only by the absence of any flow of mass or energy , but by “the absence of any tendency toward change on

6264-406: A thousand years. In the late 1740s, Carl Linnaeus introduced his system of binomial nomenclature for the classification of species. Linnaeus attempted to improve the composition and reduce the length of the previously used many-worded names by abolishing unnecessary rhetoric, introducing new descriptive terms and precisely defining their meaning. The fungi were originally treated as plants. For

6438-501: Is immortal . Many philosophical definitions of living systems have been proposed, such as self-organizing systems. Viruses in particular make definition difficult as they replicate only in host cells. Life exists all over the Earth in air, water, and soil , with many ecosystems forming the biosphere . Some of these are harsh environments occupied only by extremophiles . Life has been studied since ancient times, with theories such as Empedocles 's materialism asserting that it

6612-461: Is a body of matter and/or radiation separate from its surroundings that can be studied using the laws of thermodynamics . Thermodynamic systems can be passive and active according to internal processes. According to internal processes, passive systems and active systems are distinguished: passive, in which there is a redistribution of available energy, active, in which one type of energy is converted into another. Depending on its interaction with

6786-471: Is a mechanistic explanation for the origin of species by means of natural selection . At the beginning of the 20th century Stéphane Leduc (1853–1939) promoted the idea that biological processes could be understood in terms of physics and chemistry, and that their growth resembled that of inorganic crystals immersed in solutions of sodium silicate. His ideas, set out in his book La biologie synthétique , were widely dismissed during his lifetime, but has incurred

6960-552: Is about 4.54 billion years . Life on Earth has existed for at least 3.5 billion years, with the oldest physical traces of life dating back 3.7 billion years. Estimates from molecular clocks, as summarised in the TimeTree public database, place the origin of life around 4.0 billion years ago. Hypotheses on the origin of life attempt to explain the formation of a universal common ancestor from simple organic molecules via pre-cellular life to protocells and metabolism. In 2016,

7134-462: Is always possible, for example by gravitational forces. It is an axiom of thermodynamics that an isolated system eventually reaches internal thermodynamic equilibrium , when its state no longer changes with time. The walls of a closed system allow transfer of energy as heat and as work, but not of matter, between it and its surroundings. The walls of an open system allow transfer both of matter and of energy. This scheme of definition of terms

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7308-503: Is an acceptable idealization used in constructing mathematical models of certain natural phenomena . In the attempt to justify the postulate of entropy increase in the second law of thermodynamics , Boltzmann's H-theorem used equations , which assumed that a system (for example, a gas ) was isolated. That is all the mechanical degrees of freedom could be specified, treating the walls simply as mirror boundary conditions . This inevitably led to Loschmidt's paradox . However, if

7482-405: Is approximately 12.5  MeV/ c , which is low compared to the mass of a nucleon (approximately 938  MeV/ c ). The bottom line is that most of the mass of everyday objects comes from the interaction energy of its elementary components. The Standard Model groups matter particles into three generations, where each generation consists of two quarks and two leptons. The first generation

7656-471: Is being simulated and explored by scientists and engineers. The definition of life has long been a challenge for scientists and philosophers. This is partially because life is a process, not a substance. This is complicated by a lack of knowledge of the characteristics of living entities, if any, that may have developed outside Earth. Philosophical definitions of life have also been put forward, with similar difficulties on how to distinguish living things from

7830-556: Is capable of forming the various specialised cells that form the adult organism. This specialisation allows multicellular organisms to exploit resources more efficiently than single cells. About 800 million years ago, a minor genetic change in a single molecule, the enzyme GK-PID , may have allowed organisms to go from a single cell organism to one of many cells. Cells have evolved methods to perceive and respond to their microenvironment, thereby enhancing their adaptability. Cell signalling coordinates cellular activities, and hence governs

8004-428: Is considered in a state of thermodynamic equilibrium . The thermodynamic properties of a system in equilibrium are unchanging in time. Equilibrium system states are much easier to describe in a deterministic manner than non-equilibrium states. In some cases, when analyzing a thermodynamic process , one can assume that each intermediate state in the process is at equilibrium. Such a process is called quasistatic. For

8178-420: Is considered the starting point of modern organic chemistry . It is of historical significance because for the first time an organic compound was produced in inorganic reactions. During the 1850s Hermann von Helmholtz , anticipated by Julius Robert von Mayer , demonstrated that no energy is lost in muscle movement, suggesting that there were no "vital forces" necessary to move a muscle. These results led to

8352-402: Is delimited by walls or boundaries, either actual or notional, across which conserved (such as matter and energy) or unconserved (such as entropy) quantities can pass into and out of the system. The space outside the thermodynamic system is known as the surroundings , a reservoir , or the environment . The properties of the walls determine what transfers can occur. A wall that allows transfer of

8526-540: Is direct. A wall can be fixed (e.g. a constant volume reactor) or moveable (e.g. a piston). For example, in a reciprocating engine, a fixed wall means the piston is locked at its position; then, a constant volume process may occur. In that same engine, a piston may be unlocked and allowed to move in and out. Ideally, a wall may be declared adiabatic , diathermal , impermeable, permeable, or semi-permeable . Actual physical materials that provide walls with such idealized properties are not always readily available. The system

8700-631: Is expected to be color superconducting . Strange matter is hypothesized to occur in the core of neutron stars , or, more speculatively, as isolated droplets that may vary in size from femtometers ( strangelets ) to kilometers ( quark stars ). In particle physics and astrophysics , the term is used in two ways, one broader and the other more specific. Leptons are particles of spin- 1 ⁄ 2 , meaning that they are fermions . They carry an electric charge of −1  e (charged leptons) or 0  e (neutrinos). Unlike quarks, leptons do not carry colour charge , meaning that they do not experience

8874-417: Is extended by the evolution of novel functions over time. Death is the termination of all vital functions or life processes in an organism or cell. One of the challenges in defining death is in distinguishing it from life. Death would seem to refer to either the moment life ends, or when the state that follows life begins. However, determining when death has occurred is difficult, as cessation of life functions

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9048-459: Is in a state of thermodynamic equilibrium . Truly isolated physical systems do not exist in reality (except perhaps for the universe as a whole), because, for example, there is always gravity between a system with mass and masses elsewhere. However, real systems may behave nearly as an isolated system for finite (possibly very long) times. The concept of an isolated system can serve as a useful model approximating many real-world situations. It

9222-409: Is in terms that approximate, well enough in practice in many cases, equilibrium thermodynamical concepts. This is mostly beyond the scope of the present article, and is set out in other articles, for example the article Flow process . The classification of thermodynamic systems arose with the development of thermodynamics as a science. Theoretical studies of thermodynamic processes in the period from

9396-424: Is its soul (Greek psyche , Latin anima ). There are three kinds of souls: the vegetative soul of plants, which causes them to grow and decay and nourish themselves, but does not cause motion and sensation; the animal soul , which causes animals to move and feel; and the rational soul , which is the source of consciousness and reasoning, which (Aristotle believed) is found only in man. Each higher soul has all of

9570-400: Is made up of atoms . Such atomic matter is also sometimes termed ordinary matter . As an example, deoxyribonucleic acid molecules (DNA) are matter under this definition because they are made of atoms. This definition can be extended to include charged atoms and molecules, so as to include plasmas (gases of ions) and electrolytes (ionic solutions), which are not obviously included in

9744-455: Is made up of neutron stars and white dwarfs. Strange matter is a particular form of quark matter , usually thought of as a liquid of up , down , and strange quarks. It is contrasted with nuclear matter , which is a liquid of neutrons and protons (which themselves are built out of up and down quarks), and with non-strange quark matter, which is a quark liquid that contains only up and down quarks. At high enough density, strange matter

9918-485: Is more subtle than it first appears. All the particles that make up ordinary matter (leptons and quarks) are elementary fermions, while all the force carriers are elementary bosons. The W and Z bosons that mediate the weak force are not made of quarks or leptons, and so are not ordinary matter, even if they have mass. In other words, mass is not something that is exclusive to ordinary matter. The quark–lepton definition of ordinary matter, however, identifies not only

10092-403: Is most likely to exist, within wide bounds of uncertainty. A "Confidence of Life Detection" scale (CoLD) for reporting evidence of life beyond Earth has been proposed. Artificial life is the simulation of any aspect of life, as through computers, robotics , or biochemistry . Synthetic biology is a new area of biotechnology that combines science and biological engineering . The common goal

10266-436: Is natural to phrase the definition as: "ordinary matter is anything that is made of the same things that atoms and molecules are made of". (However, notice that one also can make from these building blocks matter that is not atoms or molecules.) Then, because electrons are leptons, and protons and neutrons are made of quarks, this definition in turn leads to the definition of matter as being "quarks and leptons", which are two of

10440-429: Is no such thing as "anti-mass" or negative mass , so far as is known, although scientists do discuss the concept. Antimatter has the same (i.e. positive) mass property as its normal matter counterpart. Different fields of science use the term matter in different, and sometimes incompatible, ways. Some of these ways are based on loose historical meanings from a time when there was no reason to distinguish mass from simply

10614-480: Is not a substance but rather a quantitative property of matter and other substances or systems; various types of mass are defined within physics – including but not limited to rest mass , inertial mass , relativistic mass , mass–energy . While there are different views on what should be considered matter, the mass of a substance has exact scientific definitions. Another difference is that matter has an "opposite" called antimatter , but mass has no opposite—there

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10788-429: Is not generally accepted. Baryonic matter is the part of the universe that is made of baryons (including all atoms). This part of the universe does not include dark energy , dark matter , black holes or various forms of degenerate matter, such as those that compose white dwarf stars and neutron stars . Microwave light seen by Wilkinson Microwave Anisotropy Probe (WMAP) suggests that only about 4.6% of that part of

10962-473: Is not only plausible, but probable or inevitable, possibly resulting in a biophysical cosmology instead of a mere physical cosmology . Other planets and moons in the Solar System and other planetary systems are being examined for evidence of having once supported simple life, and projects such as SETI are trying to detect radio transmissions from possible alien civilisations. Other locations within

11136-423: Is not possible to find an exactly defined entropy for non-equilibrium problems. For many non-equilibrium thermodynamical problems, an approximately defined quantity called 'time rate of entropy production' is very useful. Non-equilibrium thermodynamics is mostly beyond the scope of the present article. Another kind of thermodynamic system is considered in most engineering. It takes part in a flow process. The account

11310-427: Is not uniformly used, though it is convenient for some purposes. In particular, some writers use 'closed system' where 'isolated system' is here used. Anything that passes across the boundary and effects a change in the contents of the system must be accounted for in an appropriate balance equation. The volume can be the region surrounding a single atom resonating energy, such as Max Planck defined in 1900; it can be

11484-427: Is often not simultaneous across organ systems. Such determination, therefore, requires drawing conceptual lines between life and death. This is problematic because there is little consensus over how to define life. The nature of death has for millennia been a central concern of the world's religious traditions and of philosophical inquiry. Many religions maintain faith in either a kind of afterlife or reincarnation for

11658-483: Is organised into chromosomes . All species of large complex organisms are eukaryotes, including animals, plants and fungi, though with a wide diversity of protist microorganisms . The conventional model is that eukaryotes evolved from prokaryotes, with the main organelles of the eukaryotes forming through endosymbiosis between bacteria and the progenitor eukaryotic cell. The molecular mechanisms of cell biology are based on proteins . Most of these are synthesised by

11832-403: Is the up and down quarks, the electron and the electron neutrino ; the second includes the charm and strange quarks, the muon and the muon neutrino ; the third generation consists of the top and bottom quarks and the tau and tau neutrino . The most natural explanation for this would be that quarks and leptons of higher generations are excited states of

12006-417: Is the design and construction of new biological functions and systems not found in nature. Synthetic biology includes the broad redefinition and expansion of biotechnology, with the ultimate goals of being able to design and build engineered biological systems that process information, manipulate chemicals, fabricate materials and structures, produce energy, provide food, and maintain and enhance human health and

12180-611: Is the essential, characteristic, and most fundamental postulate of thermodynamics, though it is only rarely cited as a numbered law. According to Bailyn, the commonly rehearsed statement of the zeroth law of thermodynamics is a consequence of this fundamental postulate. In reality, practically nothing in nature is in strict thermodynamic equilibrium, but the postulate of thermodynamic equilibrium often provides very useful idealizations or approximations, both theoretically and experimentally; experiments can provide scenarios of practical thermodynamic equilibrium. In equilibrium thermodynamics

12354-431: Is to define life as "a self-sustained chemical system capable of undergoing Darwinian evolution ", a definition adopted by a NASA committee attempting to define life for the purposes of exobiology , based on a suggestion by Carl Sagan . This definition, however, has been widely criticised because according to it, a single sexually reproducing individual is not alive as it is incapable of evolving on its own. Others take

12528-506: Is unlikely or impossible. Organisms that have a wide range of tolerance are more widely distributed than organisms with a narrow range of tolerance. To survive, some microorganisms have evolved to withstand freezing , complete desiccation , starvation , high levels of radiation exposure , and other physical or chemical challenges. These extremophile microorganisms may survive exposure to such conditions for long periods. They excel at exploiting uncommon sources of energy. Characterization of

12702-711: The International Ocean Discovery Program found unicellular life in 120 °C sediment 1.2 km below seafloor in the Nankai Trough subduction zone. According to one researcher, "You can find microbes everywhere—they're extremely adaptable to conditions, and survive wherever they are." The inert components of an ecosystem are the physical and chemical factors necessary for life—energy (sunlight or chemical energy ), water, heat, atmosphere , gravity , nutrients , and ultraviolet solar radiation protection . In most ecosystems,

12876-503: The Nuvvuagittuq Belt of Quebec, Canada that were as old as 4.28 billion years, the oldest record of life on Earth, suggesting "an almost instantaneous emergence of life" after ocean formation 4.4 billion years ago , and not long after the formation of the Earth 4.54 billion years ago. Evolution is the change in heritable characteristics of biological populations over successive generations. It results in

13050-477: The Pauli exclusion principle , which applies to fermions . Two particular examples where the exclusion principle clearly relates matter to the occupation of space are white dwarf stars and neutron stars, discussed further below. Thus, matter can be defined as everything composed of elementary fermions. Although we do not encounter them in everyday life, antiquarks (such as the antiproton ) and antileptons (such as

13224-453: The Solar System that may host microbial life include the subsurface of Mars , the upper atmosphere of Venus , and subsurface oceans on some of the moons of the giant planets . Investigation of the tenacity and versatility of life on Earth, as well as an understanding of the molecular systems that some organisms utilise to survive such extremes, is important for the search for extraterrestrial life. For example, lichen could survive for

13398-499: The Standard Model of particle physics , matter is not a fundamental concept because the elementary constituents of atoms are quantum entities which do not have an inherent "size" or " volume " in any everyday sense of the word. Due to the exclusion principle and other fundamental interactions , some " point particles " known as fermions ( quarks , leptons ), and many composites and atoms, are effectively forced to keep

13572-570: The energy–momentum tensor that quantifies the amount of matter. This tensor gives the rest mass for the entire system. Matter, therefore, is sometimes considered as anything that contributes to the energy–momentum of a system, that is, anything that is not purely gravity. This view is commonly held in fields that deal with general relativity such as cosmology . In this view, light and other massless particles and fields are all part of matter. In particle physics, fermions are particles that obey Fermi–Dirac statistics . Fermions can be elementary, like

13746-645: The laws of nature . They coupled their ideas of soul, or lack thereof, into their theory of matter. The strongest developers and defenders of this theory were the Nyaya - Vaisheshika school, with the ideas of the Indian philosopher Kanada being the most followed. Buddhist philosophers also developed these ideas in late 1st-millennium CE, ideas that were similar to the Vaisheshika school, but ones that did not include any soul or conscience. Jain philosophers included

13920-556: The positron ) are the antiparticles of the quark and the lepton, are elementary fermions as well, and have essentially the same properties as quarks and leptons, including the applicability of the Pauli exclusion principle which can be said to prevent two particles from being in the same place at the same time (in the same state), i.e. makes each particle "take up space". This particular definition leads to matter being defined to include anything made of these antimatter particles as well as

14094-570: The soul ( jiva ), adding qualities such as taste, smell, touch, and color to each atom. They extended the ideas found in early literature of the Hindus and Buddhists by adding that atoms are either humid or dry, and this quality cements matter. They also proposed the possibility that atoms combine because of the attraction of opposites, and the soul attaches to these atoms, transforms with karma residue, and transmigrates with each rebirth . In ancient Greece , pre-Socratic philosophers speculated

14268-473: The soul , or resurrection of the body at a later date. Whether or not viruses should be considered as alive is controversial. They are most often considered as just gene coding replicators rather than forms of life. They have been described as "organisms at the edge of life" because they possess genes , evolve by natural selection, and replicate by making multiple copies of themselves through self-assembly. However, viruses do not metabolise and they require

14442-445: The stochastic behavior of the molecules in actual walls is considered, along with the randomizing effect of the ambient, background thermal radiation , Boltzmann's assumption of molecular chaos can be justified. The second law of thermodynamics for isolated systems states that the entropy of an isolated system not in equilibrium tends to increase over time, approaching maximum value at equilibrium. Overall, in an isolated system,

14616-551: The strong interaction . Leptons also undergo radioactive decay, meaning that they are subject to the weak interaction . Leptons are massive particles, therefore are subject to gravity. In bulk , matter can exist in several different forms, or states of aggregation, known as phases , depending on ambient pressure , temperature and volume . A phase is a form of matter that has a relatively uniform chemical composition and physical properties (such as density , specific heat , refractive index , and so forth). These phases include

14790-466: The structure and metabolic diversity of microbial communities in such extreme environments is ongoing. The first classification of organisms was made by the Greek philosopher Aristotle (384–322 BC), who grouped living things as either plants or animals, based mainly on their ability to move. He distinguished animals with blood from animals without blood, which can be compared with the concepts of vertebrates and invertebrates respectively, and divided

14964-419: The 1740s. Living things are composed of biochemical molecules , formed mainly from a few core chemical elements . All living things contain two types of large molecule, proteins and nucleic acids , the latter usually both DNA and RNA : these carry the information needed by each species, including the instructions to make each type of protein. The proteins, in turn, serve as the machinery which carries out

15138-458: The Sun-like "main sequence" of stellar evolution for a shorter time interval. Small red dwarfs have the opposite problem, with a smaller habitable zone that is subject to higher levels of magnetic activity and the effects of tidal locking from close orbits. Hence, stars in the intermediate mass range such as the Sun may have a greater likelihood for Earth-like life to develop. The location of

15312-401: The abandonment of scientific interest in vitalistic theories, especially after Eduard Buchner 's demonstration that alcoholic fermentation could occur in cell-free extracts of yeast. Nonetheless, belief still exists in pseudoscientific theories such as homoeopathy , which interprets diseases and sickness as caused by disturbances in a hypothetical vital force or life force. The age of Earth

15486-410: The annihilation. In short, matter, as defined in physics, refers to baryons and leptons. The amount of matter is defined in terms of baryon and lepton number. Baryons and leptons can be created, but their creation is accompanied by antibaryons or antileptons; and they can be destroyed by annihilating them with antibaryons or antileptons. Since antibaryons/antileptons have negative baryon/lepton numbers,

15660-478: The antiparticle partners of one another. In October 2017, scientists reported further evidence that matter and antimatter , equally produced at the Big Bang , are identical, should completely annihilate each other and, as a result, the universe should not exist. This implies that there must be something, as yet unknown to scientists, that either stopped the complete mutual destruction of matter and antimatter in

15834-461: The appearance of new species and often the disappearance of old ones. Evolution occurs when evolutionary processes such as natural selection (including sexual selection ) and genetic drift act on genetic variation, resulting in certain characteristics increasing or decreasing in frequency within a population over successive generations. The process of evolution has given rise to biodiversity at every level of biological organisation . Fossils are

16008-455: The atomic nuclei are composed) are destroyed—there are as many baryons after as before the reaction, so none of these matter particles are actually destroyed and none are even converted to non-matter particles (like photons of light or radiation). Instead, nuclear (and perhaps chromodynamic) binding energy is released, as these baryons become bound into mid-size nuclei having less energy (and, equivalently , less mass) per nucleon compared to

16182-650: The atoms definition. Alternatively, one can adopt the protons, neutrons, and electrons definition. A definition of "matter" more fine-scale than the atoms and molecules definition is: matter is made up of what atoms and molecules are made of , meaning anything made of positively charged protons , neutral neutrons , and negatively charged electrons . This definition goes beyond atoms and molecules, however, to include substances made from these building blocks that are not simply atoms or molecules, for example electron beams in an old cathode ray tube television, or white dwarf matter—typically, carbon and oxygen nuclei in

16356-449: The attributes of the lower ones. Aristotle believed that while matter can exist without form, form cannot exist without matter, and that therefore the soul cannot exist without the body. This account is consistent with teleological explanations of life , which account for phenomena in terms of purpose or goal-directedness. Thus, the whiteness of the polar bear's coat is explained by its purpose of camouflage. The direction of causality (from

16530-493: The basic element is fire, though perhaps he means that all is change. Empedocles (c. 490–430 BCE) spoke of four elements of which everything was made: earth, water, air, and fire. Meanwhile, Parmenides argued that change does not exist, and Democritus argued that everything is composed of minuscule, inert bodies of all shapes called atoms, a philosophy called atomism . All of these notions had deep philosophical problems. Thermodynamic system A thermodynamic system

16704-398: The basic functions of multicellular organisms. Signaling between cells can occur through direct cell contact using juxtacrine signalling , or indirectly through the exchange of agents as in the endocrine system . In more complex organisms, coordination of activities can occur through a dedicated nervous system . Though life is confirmed only on Earth, many think that extraterrestrial life

16878-430: The biosphere, including soil , hot springs , inside rocks at least 19 km (12 mi) deep underground, the deepest parts of the ocean, and at least 64 km (40 mi) high in the atmosphere. For example, spores of Aspergillus niger have been detected in the mesosphere at an altitude of 48 to 77 km. Under test conditions, life forms have been observed to survive in the vacuum of space. Life forms thrive in

17052-457: The blooded animals into five groups: viviparous quadrupeds ( mammals ), oviparous quadrupeds (reptiles and amphibians ), birds, fishes and whales . The bloodless animals were divided into five groups: cephalopods , crustaceans , insects (which included the spiders, scorpions , and centipedes ), shelled animals (such as most molluscs and echinoderms ), and " zoophytes " (animals that resemble plants). This theory remained dominant for more than

17226-416: The boundary is an imaginary surface enclosing the beaker and reactants. It is named closed , if borders are impenetrable for substance, but allow transit of energy in the form of heat, and isolated , if there is no exchange of heat and substances. The open system cannot exist in the equilibrium state. To describe deviation of the thermodynamic system from equilibrium, in addition to constitutive variables that

17400-414: The boundary of the system. Whether a system can exchange heat, work, or both is dependent on the property of its boundary. One example is fluid being compressed by a piston in a cylinder. Another example of a closed system is a bomb calorimeter , a type of constant-volume calorimeter used in measuring the heat of combustion of a particular reaction. Electrical energy travels across the boundary to produce

17574-411: The conditions vary during the day and from one season to the next. To live in most ecosystems, then, organisms must be able to survive a range of conditions, called the "range of tolerance". Outside that are the "zones of physiological stress", where the survival and reproduction are possible but not optimal. Beyond these zones are the "zones of intolerance", where survival and reproduction of that organism

17748-416: The deep Mariana Trench , and inside rocks up to 580 m (1,900 ft; 0.36 mi) below the sea floor under 2,590 m (8,500 ft; 1.61 mi) of ocean off the coast of the northwestern United States, and 2,400 m (7,900 ft; 1.5 mi) beneath the seabed off Japan. In 2014, life forms were found living 800 m (2,600 ft; 0.50 mi) below the ice of Antarctica. Expeditions of

17922-418: The definition of an intensive state variable, with respect to a reference state of the surroundings, for that substance. The intensive variable is called the chemical potential; for component substance i it is usually denoted μ i . The corresponding extensive variable can be the number of moles N i of the component substance in the system. For a contact equilibrium across a wall permeable to

18096-414: The difference between the rest mass of the products of the annihilation and the rest mass of the original particle–antiparticle pair, which is often quite large. Depending on which definition of "matter" is adopted, antimatter can be said to be a particular subclass of matter, or the opposite of matter. Antimatter is not found naturally on Earth, except very briefly and in vanishingly small quantities (as

18270-691: The disappearance of antimatter requires an asymmetry in physical laws called CP (charge–parity) symmetry violation , which can be obtained from the Standard Model, but at this time the apparent asymmetry of matter and antimatter in the visible universe is one of the great unsolved problems in physics . Possible processes by which it came about are explored in more detail under baryogenesis . Formally, antimatter particles can be defined by their negative baryon number or lepton number , while "normal" (non-antimatter) matter particles have positive baryon or lepton number. These two classes of particles are

18444-399: The early forming universe, or that gave rise to an imbalance between the two forms. Two quantities that can define an amount of matter in the quark–lepton sense (and antimatter in an antiquark–antilepton sense), baryon number and lepton number , are conserved in the Standard Model. A baryon such as the proton or neutron has a baryon number of one, and a quark, because there are three in

18618-414: The early phase of the universe and still floating about. In cosmology , dark energy is the name given to the source of the repelling influence that is accelerating the rate of expansion of the universe . Its precise nature is currently a mystery, although its effects can reasonably be modeled by assigning matter-like properties such as energy density and pressure to the vacuum itself. Fully 70% of

18792-448: The early universe and the Big Bang theory require that this matter have energy and mass, but not be composed of ordinary baryons (protons and neutrons). The commonly accepted view is that most of the dark matter is non-baryonic in nature . As such, it is composed of particles as yet unobserved in the laboratory. Perhaps they are supersymmetric particles , which are not Standard Model particles but relics formed at very high energies in

18966-428: The electron—or composite, like the proton and neutron. In the Standard Model , there are two types of elementary fermions: quarks and leptons, which are discussed next. Quarks are massive particles of spin- 1 ⁄ 2 , implying that they are fermions . They carry an electric charge of − 1 ⁄ 3   e (down-type quarks) or + 2 ⁄ 3   e (up-type quarks). For comparison, an electron has

19140-438: The elementary building blocks of matter, but also includes composites made from the constituents (atoms and molecules, for example). Such composites contain an interaction energy that holds the constituents together, and may constitute the bulk of the mass of the composite. As an example, to a great extent, the mass of an atom is simply the sum of the masses of its constituent protons, neutrons and electrons. However, digging deeper,

19314-458: The entire universe). 'Closed system' is often used in thermodynamics discussions when 'isolated system' would be correct – i.e. there is an assumption that energy does not enter or leave the system. For a thermodynamic process, the precise physical properties of the walls and surroundings of the system are important, because they determine the possible processes. An open system has one or several walls that allow transfer of matter. To account for

19488-403: The environment, a thermodynamic system may be an isolated system , a closed system , or an open system . An isolated system does not exchange matter or energy with its surroundings. A closed system may exchange heat, experience forces, and exert forces, but does not exchange matter. An open system can interact with its surroundings by exchanging both matter and energy. The physical condition of

19662-1027: The environment. Matter In classical physics and general chemistry , matter is any substance that has mass and takes up space by having volume . All everyday objects that can be touched are ultimately composed of atoms , which are made up of interacting subatomic particles , and in everyday as well as scientific usage, matter generally includes atoms and anything made up of them, and any particles (or combination of particles ) that act as if they have both rest mass and volume . However it does not include massless particles such as photons , or other energy phenomena or waves such as light or heat . Matter exists in various states (also known as phases ). These include classical everyday phases such as solid , liquid , and gas – for example water exists as ice , liquid water, and gaseous steam – but other states are possible, including plasma , Bose–Einstein condensates , fermionic condensates , and quark–gluon plasma . Usually atoms can be imagined as

19836-522: The exception being sulfur. The latter is a component of the amino acids cysteine and methionine . The most abundant of these elements in organisms is carbon, which has the desirable attribute of forming multiple, stable covalent bonds . This allows carbon-based (organic) molecules to form the immense variety of chemical arrangements described in organic chemistry . Alternative hypothetical types of biochemistry have been proposed that eliminate one or more of these elements, swap out an element for one not on

20010-518: The field of thermodynamics . In nanomaterials, the vastly increased ratio of surface area to volume results in matter that can exhibit properties entirely different from those of bulk material, and not well described by any bulk phase (see nanomaterials for more details). Phases are sometimes called states of matter , but this term can lead to confusion with thermodynamic states . For example, two gases maintained at different pressures are in different thermodynamic states (different pressures), but in

20184-447: The first generations. If this turns out to be the case, it would imply that quarks and leptons are composite particles , rather than elementary particles . This quark–lepton definition of matter also leads to what can be described as "conservation of (net) matter" laws—discussed later below. Alternatively, one could return to the mass–volume–space concept of matter, leading to the next definition, in which antimatter becomes included as

20358-527: The first theory of heat engines (Saadi Carnot, France, 1824) to the theory of dissipative structures (Ilya Prigozhin, Belgium, 1971) mainly concerned the patterns of interaction of thermodynamic systems with the environment. At the same time, thermodynamic systems were mainly classified as isolated, closed and open, with corresponding properties in various thermodynamic states, for example, in states close to equilibrium, nonequilibrium and strongly nonequilibrium. In 2010, Boris Dobroborsky (Israel, Russia) proposed

20532-409: The formation of colonies of identical cells. These cells can form group organisms through cell adhesion . The individual members of a colony are capable of surviving on their own, whereas the members of a true multi-cellular organism have developed specialisations, making them dependent on the remainder of the organism for survival. Such organisms are formed clonally or from a single germ cell that

20706-472: The formation of Earth's major animal and plant species. This interplay between organisms and their environment is an inherent feature of living systems. The biosphere is the global sum of all ecosystems. It can also be termed as the zone of life on Earth, a closed system (apart from solar and cosmic radiation and heat from the interior of the Earth), and largely self-regulating. Organisms exist in every part of

20880-470: The four types of elementary fermions (the other two being antiquarks and antileptons, which can be considered antimatter as described later). Carithers and Grannis state: "Ordinary matter is composed entirely of first-generation particles, namely the [up] and [down] quarks, plus the electron and its neutrino." (Higher generations particles quickly decay into first-generation particles, and thus are not commonly encountered. ) This definition of ordinary matter

21054-408: The fractions of energy in the universe contributed by different sources. Ordinary matter is divided into luminous matter (the stars and luminous gases and 0.005% radiation) and nonluminous matter (intergalactic gas and about 0.1% neutrinos and 0.04% supermassive black holes). Ordinary matter is uncommon. Modeled after Ostriker and Steinhardt. For more information, see NASA . Ordinary matter, in

21228-405: The fungi are more closely related to animals than to plants. As advances in microscopy enabled detailed study of cells and microorganisms, new groups of life were revealed, and the fields of cell biology and microbiology were created. These new organisms were originally described separately in protozoa as animals and protophyta/thallophyta as plants, but were united by Ernst Haeckel in

21402-479: The future to the past) is in contradiction with the scientific evidence for natural selection, which explains the consequence in terms of a prior cause. Biological features are explained not by looking at future optimal results, but by looking at the past evolutionary history of a species, which led to the natural selection of the features in question. Spontaneous generation was the belief that living organisms can form without descent from similar organisms. Typically,

21576-417: The given environment. This implies all or most of the following traits: From a physics perspective, an organism is a thermodynamic system with an organised molecular structure that can reproduce itself and evolve as survival dictates. Thermodynamically, life has been described as an open system which makes use of gradients in its surroundings to create imperfect copies of itself. Another way of putting this

21750-432: The idea was that certain forms such as fleas could arise from inanimate matter such as dust or the supposed seasonal generation of mice and insects from mud or garbage. The theory of spontaneous generation was proposed by Aristotle , who compiled and expanded the work of prior natural philosophers and the various ancient explanations of the appearance of organisms; it was considered the best explanation for two millennia. It

21924-492: The information to be preserved during reproduction and cell division. Within cells, DNA is organised into long structures called chromosomes . During cell division these chromosomes are duplicated in the process of DNA replication , providing each cell its own complete set of chromosomes. Eukaryotes store most of their DNA inside the cell nucleus . Cells are the basic unit of structure in every living thing, and all cells arise from pre-existing cells by division . Cell theory

22098-457: The internal energy is constant and the entropy can never decrease. A closed system's entropy can decrease e.g. when heat is extracted from the system. Isolated systems are not equivalent to closed systems. Closed systems cannot exchange matter with the surroundings, but can exchange energy. Isolated systems can exchange neither matter nor energy with their surroundings, and as such are only theoretical and do not exist in reality (except, possibly,

22272-454: The internal energy of the open system, this requires energy transfer terms in addition to those for heat and work. It also leads to the idea of the chemical potential . A wall selectively permeable only to a pure substance can put the system in diffusive contact with a reservoir of that pure substance in the surroundings. Then a process is possible in which that pure substance is transferred between system and surroundings. Also, across that wall

22446-640: The kingdom Protista ; later, the prokaryotes were split off in the kingdom Monera , which would eventually be divided into two separate groups, the Bacteria and the Archaea . This led to the six-kingdom system and eventually to the current three-domain system , which is based on evolutionary relationships. However, the classification of eukaryotes, especially of protists, is still controversial. As microbiology developed, viruses, which are non-cellular, were discovered. Whether these are considered alive has been

22620-561: The list, or change required chiralities or other chemical properties. Deoxyribonucleic acid or DNA is a molecule that carries most of the genetic instructions used in the growth, development, functioning and reproduction of all known living organisms and many viruses. DNA and RNA are nucleic acids ; alongside proteins and complex carbohydrates , they are one of the three major types of macromolecule that are essential for all known forms of life. Most DNA molecules consist of two biopolymer strands coiled around each other to form

22794-423: The many chemical processes of life. The cell is the structural and functional unit of life. Smaller organisms, including prokaryotes (bacteria and archaea ), consist of small single cells. Larger organisms , mainly eukaryotes , can consist of single cells or may be multicellular with more complex structure. Life is only known to exist on Earth but extraterrestrial life is thought probable . Artificial life

22968-414: The mass–energy density of the universe. Hadronic matter can refer to 'ordinary' baryonic matter, made from hadrons (baryons and mesons ), or quark matter (a generalisation of atomic nuclei), i.e. the 'low' temperature QCD matter . It includes degenerate matter and the result of high energy heavy nuclei collisions. In physics, degenerate matter refers to the ground state of a gas of fermions at

23142-545: The matter density in the universe appears to be in the form of dark energy. Twenty-six percent is dark matter. Only 4% is ordinary matter. So less than 1 part in 20 is made out of matter we have observed experimentally or described in the standard model of particle physics. Of the other 96%, apart from the properties just mentioned, we know absolutely nothing. Exotic matter is a concept of particle physics , which may include dark matter and dark energy but goes further to include any hypothetical material that violates one or more of

23316-505: The middle of the 19th century. It appealed to philosophers such as Henri Bergson , Friedrich Nietzsche , and Wilhelm Dilthey , anatomists like Xavier Bichat , and chemists like Justus von Liebig . Vitalism included the idea that there was a fundamental difference between organic and inorganic material, and the belief that organic material can only be derived from living things. This was disproved in 1828, when Friedrich Wöhler prepared urea from inorganic materials. This Wöhler synthesis

23490-445: The non-living. Legal definitions of life have been debated, though these generally focus on the decision to declare a human dead, and the legal ramifications of this decision. At least 123 definitions of life have been compiled. Since there is no consensus for a definition of life, most current definitions in biology are descriptive. Life is considered a characteristic of something that preserves, furthers or reinforces its existence in

23664-414: The number of atoms of element i {\displaystyle i} in molecule j {\displaystyle j} , and b i 0 {\displaystyle b_{i}^{0}} the total number of atoms of element i {\displaystyle i} in the system, which remains constant, since the system is closed. There is one such equation for each element in

23838-412: The number of fermions rather than the temperature, unlike normal states of matter. Degenerate matter is thought to occur during the evolution of heavy stars. The demonstration by Subrahmanyan Chandrasekhar that white dwarf stars have a maximum allowed mass because of the exclusion principle caused a revolution in the theory of star evolution. Degenerate matter includes the part of the universe that

24012-563: The oldest from the Archaean Eon, up to 3.4 billion years old. Extinction is the process by which a species dies out. The moment of extinction is the death of the last individual of that species. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively after a period of apparent absence. Species become extinct when they are no longer able to survive in changing habitat or against superior competition. Over 99% of all

24186-414: The ordinary quark and lepton, and thus also anything made of mesons , which are unstable particles made up of a quark and an antiquark. In the context of relativity , mass is not an additive quantity, in the sense that one cannot add the rest masses of particles in a system to get the total rest mass of the system. In relativity, usually a more general view is that it is not the sum of rest masses , but

24360-401: The original small (hydrogen) and large (plutonium etc.) nuclei. Even in electron–positron annihilation , there is no net matter being destroyed, because there was zero net matter (zero total lepton number and baryon number) to begin with before the annihilation—one lepton minus one antilepton equals zero net lepton number—and this net amount matter does not change as it simply remains zero after

24534-447: The overall baryon/lepton numbers are not changed, so matter is conserved. However, baryons/leptons and antibaryons/antileptons all have positive mass, so the total amount of mass is not conserved. Further, outside of natural or artificial nuclear reactions, there is almost no antimatter generally available in the universe (see baryon asymmetry and leptogenesis ), so particle annihilation is rare in normal circumstances. Pie chart showing

24708-608: The parent cell divides into two or more daughter cells. For prokaryotes, cell division occurs through a process of fission in which the DNA is replicated, then the two copies are attached to parts of the cell membrane. In eukaryotes , a more complex process of mitosis is followed. However, the result is the same; the resulting cell copies are identical to each other and to the original cell (except for mutations ), and both are capable of further division following an interphase period. Multicellular organisms may have first evolved through

24882-408: The particulate theory of matter include the ancient Indian philosopher Kanada (c. 6th–century BCE or after), pre-Socratic Greek philosopher Leucippus (~490 BCE), and pre-Socratic Greek philosopher Democritus (~470–380 BCE). Matter should not be confused with mass, as the two are not the same in modern physics. Matter is a general term describing any 'physical substance'. By contrast, mass

25056-472: The preserved remains or traces of organisms from the remote past. The totality of fossils, both discovered and undiscovered, and their placement in layers ( strata ) of sedimentary rock is known as the fossil record . A preserved specimen is called a fossil if it is older than the arbitrary date of 10,000 years ago. Hence, fossils range in age from the youngest at the start of the Holocene Epoch to

25230-472: The properties of known forms of matter. Some such materials might possess hypothetical properties like negative mass . In ancient India , the Buddhist , Hindu , and Jain philosophical traditions each posited that matter was made of atoms ( paramanu , pudgala ) that were "eternal, indestructible, without parts, and innumerable" and which associated or dissociated to form more complex matter according to

25404-399: The protons and neutrons are made up of quarks bound together by gluon fields (see dynamics of quantum chromodynamics ) and these gluon fields contribute significantly to the mass of hadrons. In other words, most of what composes the "mass" of ordinary matter is due to the binding energy of quarks within protons and neutrons. For example, the sum of the mass of the three quarks in a nucleon

25578-503: The quarks and leptons definition, constitutes about 4% of the energy of the observable universe . The remaining energy is theorized to be due to exotic forms, of which 23% is dark matter and 73% is dark energy . In astrophysics and cosmology , dark matter is matter of unknown composition that does not emit or reflect enough electromagnetic radiation to be observed directly, but whose presence can be inferred from gravitational effects on visible matter. Observational evidence of

25752-439: The reaction process. In this case, the fact that the system is closed is expressed by stating that the total number of each elemental atom is conserved, no matter what kind of molecule it may be a part of. Mathematically: where N j {\displaystyle N_{j}} denotes the number of j {\displaystyle j} -type molecules, a i j {\displaystyle a_{ij}}

25926-408: The result of radioactive decay , lightning or cosmic rays ). This is because antimatter that came to exist on Earth outside the confines of a suitable physics laboratory would almost instantly meet the ordinary matter that Earth is made of, and be annihilated. Antiparticles and some stable antimatter (such as antihydrogen ) can be made in tiny amounts, but not in enough quantity to do more than test

26100-458: The ribosomes through an enzyme-catalyzed process called protein biosynthesis . A sequence of amino acids is assembled and joined based upon gene expression of the cell's nucleic acid. In eukaryotic cells, these proteins may then be transported and processed through the Golgi apparatus in preparation for dispatch to their destination. Cells reproduce through a process of cell division in which

26274-555: The same phase (both are gases). Antimatter is matter that is composed of the antiparticles of those that constitute ordinary matter. If a particle and its antiparticle come into contact with each other, the two annihilate ; that is, they may both be converted into other particles with equal energy in accordance with Albert Einstein 's equation E = mc . These new particles may be high-energy photons ( gamma rays ) or other particle–antiparticle pairs. The resulting particles are endowed with an amount of kinetic energy equal to

26448-574: The scale of elementary particles, a definition that follows this tradition can be stated as: "ordinary matter is everything that is composed of quarks and leptons ", or "ordinary matter is everything that is composed of any elementary fermions except antiquarks and antileptons". The connection between these formulations follows. Leptons (the most famous being the electron ), and quarks (of which baryons , such as protons and neutrons , are made) combine to form atoms , which in turn form molecules . Because atoms and molecules are said to be matter, it

26622-421: The so-called wave–particle duality . A chemical substance is a unique form of matter with constant chemical composition and characteristic properties . Chemical substances may take the form of a single element or chemical compounds . If two or more chemical substances can be combined without reacting , they may form a chemical mixture . If a mixture is separated to isolate one chemical substance to

26796-433: The soul, like everything else, was composed of fiery atoms. He elaborated on fire because of the apparent connection between life and heat, and because fire moves. Plato , in contrast, held that the world was organised by permanent forms , reflected imperfectly in matter; forms provided direction or intelligence, explaining the regularities observed in the world. The mechanistic materialism that originated in ancient Greece

26970-437: The special context of thermodynamics. The possible equilibria between bodies are determined by the physical properties of the walls that separate the bodies. Equilibrium thermodynamics in general does not measure time. Equilibrium thermodynamics is a relatively simple and well settled subject. One reason for this is the existence of a well defined physical quantity called 'the entropy of a body'. Non-equilibrium thermodynamics, as

27144-484: The species that have ever lived are now extinct. Mass extinctions may have accelerated evolution by providing opportunities for new groups of organisms to diversify. The diversity of life on Earth is a result of the dynamic interplay between genetic opportunity , metabolic capability, environmental challenges, and symbiosis . For most of its existence, Earth's habitable environment has been dominated by microorganisms and subjected to their metabolism and evolution. As

27318-487: The star within a galaxy may also affect the likelihood of life forming. Stars in regions with a greater abundance of heavier elements that can form planets, in combination with a low rate of potentially habitat -damaging supernova events, are predicted to have a higher probability of hosting planets with complex life. The variables of the Drake equation are used to discuss the conditions in planetary systems where civilisation

27492-484: The state variables do not include fluxes because in a state of thermodynamic equilibrium all fluxes have zero values by definition. Equilibrium thermodynamic processes may involve fluxes but these must have ceased by the time a thermodynamic process or operation is complete bringing a system to its eventual thermodynamic state. Non-equilibrium thermodynamics allows its state variables to include non-zero fluxes, which describe transfers of mass or energy or entropy between

27666-408: The sugar of one nucleotide and the phosphate of the next, resulting in an alternating sugar-phosphate backbone . According to base pairing rules (A with T, and C with G), hydrogen bonds bind the nitrogenous bases of the two separate polynucleotide strands to make double-stranded DNA. This has the key property that each strand contains all the information needed to recreate the other strand, enabling

27840-478: The system, W {\displaystyle W} the work done by the system. For infinitesimal changes the first law for closed systems may stated: If the work is due to a volume expansion by d V {\displaystyle \mathrm {d} V} at a pressure P {\displaystyle P} then: For a quasi-reversible heat transfer, the second law of thermodynamics reads: where T {\displaystyle T} denotes

28014-492: The system. An isolated system is more restrictive than a closed system as it does not interact with its surroundings in any way. Mass and energy remains constant within the system, and no energy or mass transfer takes place across the boundary. As time passes in an isolated system, internal differences in the system tend to even out and pressures and temperatures tend to equalize, as do density differences. A system in which all equalizing processes have gone practically to completion

28188-444: The thermodynamic system will always tend to a non-equilibrium state with respect to the environment. In isolated systems it is consistently observed that as time goes on internal rearrangements diminish and stable conditions are approached. Pressures and temperatures tend to equalize, and matter arranges itself into one or a few relatively homogeneous phases . A system in which all processes of change have gone practically to completion

28362-478: The thermodynamic temperature and S {\displaystyle S} the entropy of the system. With these relations the fundamental thermodynamic relation , used to compute changes in internal energy, is expressed as: For a simple system, with only one type of particle (atom or molecule), a closed system amounts to a constant number of particles. For systems undergoing a chemical reaction , there may be all sorts of molecules being generated and destroyed by

28536-445: The three familiar ones ( solids , liquids , and gases ), as well as more exotic states of matter (such as plasmas , superfluids , supersolids , Bose–Einstein condensates , ...). A fluid may be a liquid, gas or plasma. There are also paramagnetic and ferromagnetic phases of magnetic materials . As conditions change, matter may change from one phase into another. These phenomena are called phase transitions and are studied in

28710-472: The type of system, it may interact with the system by exchanging mass, energy (including heat and work), momentum , electric charge , or other conserved properties . The environment is ignored in the analysis of the system, except in regards to these interactions. In a closed system, no mass may be transferred in or out of the system boundaries. The system always contains the same amount of matter, but (sensible) heat and (boundary) work can be exchanged across

28884-472: The underlying nature of the visible world. Thales (c. 624 BCE–c. 546 BCE) regarded water as the fundamental material of the world. Anaximander (c. 610 BCE–c. 546 BCE) posited that the basic material was wholly characterless or limitless: the Infinite ( apeiron ). Anaximenes (flourished 585 BCE, d. 528 BCE) posited that the basic stuff was pneuma or air. Heraclitus (c. 535 BCE–c. 475 BCE) seems to say

29058-412: The universe is made up of a combination of four eternal "elements" or "roots of all": earth, water, air, and fire. All change is explained by the arrangement and rearrangement of these four elements. The various forms of life are caused by an appropriate mixture of elements. Democritus (460 BC) was an atomist ; he thought that the essential characteristic of life was having a soul ( psyche ), and that

29232-411: The universe within range of the best telescopes (that is, matter that may be visible because light could reach us from it) is made of baryonic matter. About 26.8% is dark matter, and about 68.3% is dark energy. The great majority of ordinary matter in the universe is unseen, since visible stars and gas inside galaxies and clusters account for less than 10 per cent of the ordinary matter contribution to

29406-442: The warmer part decreases. As a result, after some time, the temperature in the rod will equalize – the rod will come to a state of thermodynamic equilibrium. If the process of converting one type of energy into another takes place inside a thermodynamic system, for example, in chemical reactions, in electric or pneumatic motors, when one solid body rubs against another, then the processes of energy release or absorption will occur, and

29580-535: Was composed of four eternal elements , and Aristotle 's hylomorphism asserting that living things have souls and embody both form and matter. Life originated at least 3.5 billion years ago, resulting in a universal common ancestor . This evolved into all the species that exist now, by way of many extinct species, some of which have left traces as fossils . Attempts to classify living things, too, began with Aristotle . Modern classification began with Carl Linnaeus 's system of binomial nomenclature in

29754-409: Was decisively dispelled by the experiments of Louis Pasteur in 1859, who expanded upon the investigations of predecessors such as Francesco Redi . Disproof of the traditional ideas of spontaneous generation is no longer controversial among biologists. Vitalism is the belief that there is a non-material life-principle. This originated with Georg Ernst Stahl (17th century), and remained popular until

29928-731: Was described above, a set of internal variables ξ 1 , ξ 2 , … {\displaystyle \xi _{1},\xi _{2},\ldots } have been introduced. The equilibrium state is considered to be stable and the main property of the internal variables, as measures of non-equilibrium of the system, is their trending to disappear; the local law of disappearing can be written as relaxation equation for each internal variable where τ i = τ i ( T , x 1 , x 2 , … , x n ) {\displaystyle \tau _{i}=\tau _{i}(T,x_{1},x_{2},\ldots ,x_{n})}

30102-493: Was formulated by Henri Dutrochet , Theodor Schwann , Rudolf Virchow and others during the early nineteenth century, and subsequently became widely accepted. The activity of an organism depends on the total activity of its cells, with energy flow occurring within and between them. Cells contain hereditary information that is carried forward as a genetic code during cell division. There are two primary types of cells, reflecting their evolutionary origins. Prokaryote cells lack

30276-485: Was revived and revised by the French philosopher René Descartes (1596–1650), who held that animals and humans were assemblages of parts that together functioned as a machine. This idea was developed further by Julien Offray de La Mettrie (1709–1750) in his book L'Homme Machine . In the 19th century the advances in cell theory in biological science encouraged this view. The evolutionary theory of Charles Darwin (1859)

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