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77-548: The Manchester Terrier is a breed of dog of the smooth-haired terrier type. It was first bred in the 19th century to control vermin, notably rats, at which it excelled. So efficient at the task was it that it often appeared in rat-baiting pits until that sport, which had effectively been illegal in the UK since 1835, finally died out at the beginning of the 20th century. The breed is generally healthy, although dogs can be affected by several inheritable genetic disorders. A crash in

154-456: A breed standard which is a written description of the ideal specimen of the breed. Other uses of the term breed when referring to dogs include pure breeds , cross-breeds , mixed breeds and natural breeds . Prior to the standardization of dog breeds, there were many different types of dogs that were defined by their own, unique functions. Many different terms were used to describe dogs, such as breed, strain, type, kind, and variety. By

231-461: A lncRNA variant in the IGF1 region arisen in wolves from northern latitudes c. 53.000 years ago and fixed by natural and subsequently by human selection. Their behavioral traits include guarding, herding, and hunting, retrieving, and scent detection. Their personality traits include hypersocial behavior, boldness, and aggression, which demonstrates the functional and behavioral diversity of dogs. As

308-456: A "school" book about hawking , hunting, fishing, and heraldry , attributed to Juliana Berners (Barnes), lists dogs of the time mainly by function: "First there is a greyhound, a bastard, a mongrel , a mastiff, a limer, a spaniel, " raches " (small-to-medium-sized scenthounds), "kennets" (small hunting dogs), terriers , "butcher's hounds", dung-heap dogs, "trundel tails" ( lapdogs ?) and prick-eared curs , and small ladies puppies that bear away

385-450: A 0.46 kg (1.0 lb) teacup poodle to a 90 kg (200 lb) giant mastiff . The skull, body, and limb proportions vary significantly between breeds, with dogs displaying more phenotypic diversity than can be found within the entire order of carnivores. These breeds possess distinct traits related to morphology, which include body size, skull shape, tail phenotype, fur type and colour. The difference in size has been attributed to

462-575: A 2010 study ( Afghan Hound , Akita , Alaskan Malamute , American Eskimo , Basenji , Canaan dog , Chow Chow , Dingo , New Guinea singing dog , Saluki , Samoyed , Shar-Pei , Siberian Husky ), plus another three: the Eurasier , Finnish Spitz and Shiba Inu . The study referred to these 16 as basal breeds, as opposed to ancient breeds, as they exhibited genetic divergence but not all of them were historically considered to be "ancient breeds". The 2012 study found that modern breeds only emerged in

539-737: A known foundation stock may still not be recognized by some clubs as breeds. For instance, the feist is a hunting dog raised in the Southern United States for hunting small game. Feists have a consistent set of characteristics that reliably differentiate them from other dog types and breeds. However, the United Kennel Club recognizes one breed of feist, the Treeing Feist , while the American Kennel Club does not recognize any feist breed. A dog

616-453: A membrane-bound H antigen. The I enzyme adds a galactose. The i allele produces no modification. Thus the I and I alleles are each dominant to i ( I I and I i individuals both have type A blood, and I I and I i individuals both have type B blood), but I I individuals have both modifications on their blood cells and thus have type AB blood, so the I and I alleles are said to be co-dominant. Another example occurs at

693-471: A mix of two breeds and is not the result of intentional breeding . In the United States, the term mixed-breed is a favored synonym over mongrel among individuals who wish to avoid negative connotations associated with the latter term. The implication that such dogs must be a mix of defined breeds may stem from an inverted understanding of the origins of dog breeds. Purebred dogs have been, for

770-587: A particular environment and in isolation from other populations of the species. This environment included humans but with little or no selective breeding by humans . Dominance (genetics) Dominance is a key concept in Mendelian inheritance and classical genetics . Letters and Punnett squares are used to demonstrate the principles of dominance in teaching, and the upper-case letters are used to denote dominant alleles and lower-case letters are used for recessive alleles. An often quoted example of dominance

847-406: A pink snapdragon flower. The pink snapdragon is the result of incomplete dominance. A similar type of incomplete dominance is found in the four o'clock plant wherein pink color is produced when true-bred parents of white and red flowers are crossed. In quantitative genetics , where phenotypes are measured and treated numerically, if a heterozygote's phenotype is exactly between (numerically) that of

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924-566: A result, today dogs are the most abundant carnivore species and are dispersed around the world. The most striking example of this dispersal is that of the numerous modern breeds of European lineage during the Victorian era . A genetic study identified 51 regions of the dog genome which were associated with phenotype variation among breeds in the 57 traits studied, which included body, cranial, dental, and long bone shape and size. There were three quantitative trait loci that explained most of

1001-488: A select set of ancestors who had the same characteristics. Dogs of a specific breed breed true , producing young that are very similar to their parents. An individual dog is identified as a member of a breed through proof of ancestry, using genetic analysis or written records of ancestry. Without such proof, identification of a specific breed is not reliable. Such records, called stud books , may be maintained by individuals, clubs, or other organizations. Kennel clubs provide

1078-728: A sled dog is 20–25 kg (44–55 lb) based on thermo-regulation, and the ancient sled dogs were between 16–25 kg (35–55 lb). The same standard has been found in the remains of sled dogs from this region 2,000 years ago and in the modern Siberian Husky breed standard. Other dogs were larger at 30 kg (66 lb) and appear to be dogs that had been crossed with wolves and used for polar-bear hunting. Between 3,000 and 4,000 years ago greyhound -types of dogs were depicted on pottery and paintings in Egypt and Western Asia. Mastiff -type dogs were kept for guarding and hunting, and short-legged dogs were also bred. Most modern dog breeds are

1155-407: A study indicated that some breed-temperaments, such as anxiety or fear, may be linked to gene mutations. Other temperaments may be due to the legacies of 'ancient' ancestry. Groups of owners that have dogs of the same breed and have an interest in dog breeding can form national Kennel clubs . Kennel Clubs maintain breed standards , record pedigrees in a breed registry (or studbook), and issue

1232-437: A third, and co-dominant with a fourth. Additionally, one allele may be dominant for one trait but not others. Dominance differs from epistasis , the phenomenon of an allele of one gene masking the effect of alleles of a different gene. Gregor Johann Mendel , "The Father of Genetics", promulgated the idea of dominance in the 1860s. However, it was not widely known until the early twentieth century. Mendel observed that, for

1309-445: A variety of traits of garden peas having to do with the appearance of seeds, seed pods, and plants, there were two discrete phenotypes, such as round versus wrinkled seeds, yellow versus green seeds, red versus white flowers or tall versus short plants. When bred separately, the plants always produced the same phenotypes, generation after generation. However, when lines with different phenotypes were crossed (interbred), one and only one of

1386-475: Is a global organisation with 98 members and contract partners (one member per country) that recognize 356 purebreds. A dog crossbreed is the result of mating two different breeds. "Designer Dog" became a fad in the late 20th century. Dog crossbreeds are combinations of lineages of the domestic dog ; they are distinguished from canid hybrids , which are interspecific crosses between Canis species (wolves, coyotes, jackals, etc.). Notable canid hybrids include

1463-627: Is a particular type of dog that was purposefully bred by humans to perform specific tasks, such as herding, hunting, and guarding. Dogs are the most variable mammal on Earth, with artificial selection producing upward of 360 globally recognized breeds. These breeds possess distinct traits related to morphology, which include body size and shape, tail phenotype , fur type, etc, but are only one species of dog. Their behavioral traits include guarding, herding, and hunting, and personality traits such as hyper-social behavior, boldness, and aggression. Most breeds were derived from small numbers of founders within

1540-740: Is a term formerly, but no longer, used for a particular group of dog breeds by the American Kennel Club. These breeds were referred to as "ancient", as opposed to modern, breeds because historically it was believed their origins dated back more than 500 years. In 2004, a study looked at the microsatellites of 414 purebred dogs representing 85 breeds. The study found that dog breeds were so genetically distinct that 99% of individual dogs could be correctly assigned to their breed based on their genotype , indicating that breeding barriers (purebred breeding) have led to distinct genetic units. The study identified 9 breeds that could be represented on

1617-482: Is as "feisty and affectionate" as he is "courageous and tough", and makes a "loyal, eager-to-please" companion. The Canadian and American Kennel Clubs recognise two varieties of Manchester Terrier, the original and a miniaturised variant, the Toy Manchester Terrier . The latter was recognised as a separate breed by the American Kennel Club in 1938, and is still considered to be a separate breed by

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1694-565: Is neither a breed registry nor does it issue pedigrees or keep addresses for breeders. It is a global canine organization with member and contract partners (one member per country) that conduct international conformation shows, working/hunting/herding trials, and various other events. The results are submitted to FCI for processing, approval and international recognition. Each of the member and contract partners issue and maintain their own pedigrees and respective breed standards, and train their own judges. FCI ensures that each member mutually recognizes

1771-598: Is said to be purebred if their parents were purebred and if the dog meets the standards of the breed. The American Kennel Club allows mixed-breed dogs to be shown but under the condition the animals have been spayed or neutered, are not a wolf hybrid, and not eligible for the AKC Foundation Stock Service Program or an AKC Purebred Alternative Listing (PAL). In Canada, the Animal Pedigree Act lays out strict standards for

1848-411: Is the inheritance of seed shape in peas . Peas may be round, associated with allele R , or wrinkled, associated with allele r . In this case, three combinations of alleles (genotypes) are possible: RR , Rr , and rr . The RR ( homozygous ) individuals have round peas, and the rr (homozygous) individuals have wrinkled peas. In Rr ( heterozygous ) individuals, the R allele masks

1925-517: The Fédération Cynologique Internationale ." In 2017, a study showed that 9,000 years ago the domestic dog was present at what is now Zhokhov Island , arctic north-eastern Siberia, which was connected to the mainland at that time. The dogs were selectively bred as either sled dogs or as hunting dogs, which implies that a sled dog standard and a hunting dog standard existed at that time. The optimal maximum size for

2002-632: The Italian Greyhound and the Dachshund , may also have been introduced into the bloodline. The Manchester Terrier gained the nickname of the Gentleman's Terrier in Victorian times, because of its dignified demeanour. By the early 19th century a closer facsimile to the current Manchester terrier had evolved. In The Dog in Health and Disease by J. A. Walsh, a full chapter was devoted to

2079-484: The coydog , coywolf , dingo–dog hybrid , jackal–dog hybrid , and wolfdog .   Historically, crosses between dogs of different types were more well accepted at a time when modern purebred breeds (based on the eugenics principles) did not yet exist. Some crossbred dogs, created by breeding two purebred dogs of different breeds, may have the advantage of heterosis , or crossbreed vigor. This advantage can be progressively diluted when two crossbreeds are bred in

2156-418: The fleas and diverse small sorts." A study in 2012 examined 49,000 single nucleotide polymorphisms that gave a genome -wide coverage of 1,375 dogs representing 35 breeds, 19 wolves, and previous published genetic signatures of other breeds, giving a total of 121 breeds covered. The study found a deep genetic split between old-world and new-world wolves, and confirmed the genetic divergence of 13 breeds from

2233-509: The 19th century and that claims of their antiquity are based on little or no historical or empirical evidence. The study indicated that throughout history, global dog populations experienced numerous episodes of diversification and homogenization, with each round further reducing the power of genetic data derived from modern breeds to help infer their early history. Of the basal breeds, the American Eskimo Dog and Eurasier were

2310-411: The 3:1 phenotype ratio. Mendel did not use the terms gene, allele, phenotype, genotype, homozygote, and heterozygote, all of which were introduced later. He did introduce the notation of capital and lowercase letters for dominant and recessive alleles, respectively, still in use today. In 1928, British population geneticist Ronald Fisher proposed that dominance acted based on natural selection through

2387-691: The British Kennel Club, the English Toy Terrier (Black & Tan) . Where the Manchester Terrier may have button ears, erect ears, or cropped ears, the Toy Manchester Terrier only has erect ears. Other than the size difference and ear type, the Manchester Terrier and the Toy Manchester Terrier have the same overall appearance. Since 1958 the American Kennel Club has considered both to be varieties of

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2464-454: The English language to mention numbers of dog types are from the "Cynegetica" (hunting literature), namely, The Art of Venery (1327) by Twiti (Twici), a treatise that describes hunting with the limer (a leashed bloodhound type); the pack of running hounds , which included barcelets and brachetz (both scent hounds ); and the sighthound and greyhound . More significant in recording

2541-642: The F1 generation are self-pollinated, the phenotypic and genotypic ratio of the F2 generation will be 1:2:1 (Red:Spotted:White). These ratios are the same as those for incomplete dominance. Again, this classical terminology is inappropriate – in reality, such cases should not be said to exhibit dominance at all. Dominance can be influenced by various genetic interactions and it is essential to evaluate them when determining phenotypic outcomes. Multiple alleles , epistasis and pleiotropic genes are some factors that might influence

2618-426: The Manchester Terrier. Secondary glaucoma, also known as lens luxation , is caused by excessive pressure within the eyeball forcing the lens of the eye out of alignment. The only effective treatment for this condition is surgical removal of the lens. Von Willebrands disease is a blood-clotting disorder that affects humans and several breeds of dog, including the Manchester Terrier. The genetic mutation occurs in both

2695-445: The attempt to create a breed, narrowing the gene pool. The best way to continue taking advantage of crossbreed vigor is from the breeding of dogs of purebred ancestry, as this vigor is typically seen only in the first generation cross of two purebred animals of separate breeds, thus taking advantage of genetic diversity . A mongrel, mixed-breed dog or mutt is a dog that does not belong to one officially recognized breed but can be

2772-439: The beginning of the 19th century Sydenham Edwards described what he called a Manchester Terrier in his Cynographia Britannica . The Black and Tan Terrier had an almost legendary status as a rat killer, an ability that was much prized in overcrowded and rat-infested cities such as 19th-century Manchester. The sport of rat baiting , in which a terrier was thrown into an enclosure filled with rats, and timed to see how long it took

2849-420: The black and tan, for the first time recognising it as an established breed. The description given by Walsh is still relevant: Smooth haired, long tapering nose, narrow flat skull, eyes small and bright, chest rather deep than wide, only true colour black and tan. Some terrier breeds have a reputation for being "stoic and dour", but such terms do not apply to the intelligent and inquisitive Manchester Terrier, who

2926-563: The branches of a phylogenetic tree which grouped together with strong statistical support and could be separated from the other breeds with a modern European origin. These 9 breeds had been referred to as "ancient breeds". The study found that the Pharaoh Hound and Ibizan Hound were not as old as once believed; rather, they had been recreated from combinations of other breeds, and that the Norwegian Elkhound grouped with

3003-882: The breed club to be unacceptable (called faults ). In addition, most breed standards include a historical section, describing the place of origin and the original work done by the breed or its ancestor types . Dogs with a breed standard may be accepted into one or more of the major registries ( kennel clubs ) of dog breeds, including The Kennel Club (1873, UK), American Kennel Club (1884), New Zealand Kennel Club (1886), Canadian Kennel Club (1888), United Kennel Club (1898), United Kennel Clubs International (UCI, Germany 1976), Australian National Kennel Council (1958), and other national breed registries. Recognized dog breeds are classified by groups , such as Hound, Terrier, Working, Herding, Sporting, Non-Sporting, Toy and Miscellaneous; some groups may be further subdivided by some registries. Fédération Cynologique Internationale (FCI)

3080-505: The breeding, exhibiting and judging of pure-bred dogs. It now has 99 member countries. According to BigThink , over 40% of the world’s dog breeds come from the United Kingdom , France and Germany . It states: " Great Britain and France are the ground zero of dog fancying, with 57 registered breeds each. Germany is not far behind, with 47 breeds. These three countries alone represent more than 40% of all dog breeds recognized by

3157-401: The contribution of modifier genes . In 1929, American geneticist Sewall Wright responded by stating that dominance is simply a physiological consequence of metabolic pathways and the relative necessity of the gene involved. In complete dominance, the effect of one allele in a heterozygous genotype completely masks the effect of the other. The allele that masks are considered dominant to

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3234-461: The documenting of what it calls "evolving breeds". The breed standard for each breed of dog is a detailed description of the appearance and behaviour of an idealized dog of that breed. Included in the breed standard description are externally observable aspects of appearance and behaviour that are considered by the breed club to be the most important for the breed, and externally observable details of appearance or temperament that are considered by

3311-451: The dog to kill them all, had become popular by the early 19th century along with rabbit coursing . In an effort to produce a dog that could excel at both sports, the breeder John Hulme decided to cross a Black and Tan Terrier with a Whippet . Similar dogs were bred in various other parts of England, but by 1860 Manchester had become the breed centre for these new terriers, and so they became known as Manchester Terriers. Other breeds, including

3388-556: The dog–wolf genetic divergence are not known. Dogs are the most variable mammal on earth with around 450 globally recognized dog breeds. In the Victorian era , directed human selection developed the modern dog breeds, which resulted in a vast range of phenotypes. Most breeds were derived from small numbers of founders within the last 200 years, and since then dogs have undergone rapid phenotypic change and were formed into today's modern breeds due to artificial selection imposed by humans. These breeds can vary in size and weight from

3465-541: The dominant gene. However, if the F1-generation is further crossed with the F1-generation (heterozygote crossed with heterozygote) the offspring (F2-generation) will present the phenotype associated with the dominant gene ¾ times. Although heterozygote monohybrid crossing can result in two phenotype variants, it can result in three genotype variants -  homozygote dominant, heterozygote and homozygote recessive, respectively. In dihybrid inheritance we look at

3542-662: The end of the Second World War in 1945 there were only 11 Manchester Terriers registered with the Kennel Club, but thanks to the efforts of the British Manchester Terrier Club, formed in 1937, and others, the breed was saved from extinction. The Kennel Club has nevertheless classified the Manchester Terrier as a vulnerable native breed. Between 2017 and 2021, an average of 177 births per year were registered. Dog breed A dog breed

3619-509: The end of the Victorian era, society had changed and so did the role of dogs. The form was given a more prominent role than function. Different types or breeds of dog were being developed by breeders who wanted to define specific characteristics and desirable features in their dogs. Driven by dog shows and the groups that hosted them, the term dog breed took on an entirely new meaning. Dog show competitions included best-in-breed winners, and

3696-417: The inheritance of two pairs of genes simultaneous. Assuming here that the two pairs of genes are located at non-homologous chromosomes, such that they are not coupled genes (see genetic linkage ) but instead inherited independently. Consider now the cross between parents (P-generation) of genotypes homozygote dominant and recessive, respectively. The offspring (F1-generation) will always heterozygous and present

3773-437: The last 200 years. As a result of their adaptability to many environments and breedability for human needs, today dogs are the most abundant carnivore species and are dispersed around the world. A dog breed will consistently produce the physical traits, movement and temperament that were developed over decades of selective breeding. For each breed they recognize, kennel clubs and breed registries usually maintain and publish

3850-485: The level of dominance the alleles expresses towards each other. Pleiotropic genes are genes where one single gene affects two or more characters (phenotype). This means that a gene can have a dominant effect on one trait, but a more recessive effect on another trait. Epistasis is interactions between multiple alleles at different loci. Easily said, several genes for one phenotype. The dominance relationship between alleles involved in epistatic interactions can influence

3927-450: The locus for the beta-globin component of hemoglobin , where the three molecular phenotypes of Hb /Hb , Hb /Hb , and Hb /Hb are all distinguishable by protein electrophoresis . (The medical condition produced by the heterozygous genotype is called sickle-cell trait and is a milder condition distinguishable from sickle-cell anemia , thus the alleles show incomplete dominance concerning anemia, see above). For most gene loci at

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4004-454: The modern breeds because the genetic heritage of the modern breeds has become blurred due to admixture, and the basal breeds have mostly avoided admixture with them due to geographic or cultural barriers. As dogs are a subspecies but their breeds are distinct genetic units, and because only certain breeds share the same type of cancers as humans, the differences in the genes of different breeds may be useful in human medical research. In 2014,

4081-455: The molecular level, both alleles are expressed co-dominantly, because both are transcribed into RNA . Co-dominance, where allelic products co-exist in the phenotype, is different from incomplete dominance, where the quantitative interaction of allele products produces an intermediate phenotype. For example, in co-dominance, a red homozygous flower and a white homozygous flower will produce offspring that have red and white spots. When plants of

4158-416: The most part, artificially created from random-bred populations by human selective breeding with the purpose of enhancing desired physical, behavioral, or temperamental characteristics. Dogs that are not purebred are not necessarily a mix of such defined breeds. Therefore, among some experts and fans of such dogs, mongrel is still the preferred term. Natural breeds rose through time in response to

4235-637: The number of registered Manchester Terriers following the Second World War has resulted in the Kennel Club categorising it as a vulnerable native breed, in danger of extinction. The Manchester Terrier was developed from the Black and Tan Terrier and the Whippet .. Writing in the early 16th century, John Caius describes a Manchester-terrier type of dog in De Canibus Britannicis , and at

4312-409: The offspring, with the result that all of these hybrids were heterozygotes (Aa), and that one of the two alleles in the hybrid cross dominated expression of the other: A masked a. The final cross between two heterozygotes (Aa X Aa) would produce AA, Aa, and aa offspring in a 1:2:1 genotype ratio with the first two classes showing the (A) phenotype, and the last showing the (a) phenotype, thereby producing

4389-458: The other European dogs despite reports of direct Scandinavian origins dating back 5,000 years. Dog types are broad categories of dogs based on form, function or style of work, lineage, or appearance. In contrast, modern dog breeds are particular breed standards, sharing a common set of heritable characteristics, determined by the kennel club that recognizes the breed. The spread of modern dog breeds has been difficult to resolve because many are

4466-418: The other allele, and the masked allele is considered recessive . When we only look at one trait determined by one pair of genes, we call it monohybrid inheritance . If the crossing is done between parents (P-generation, F0-generation) who are homozygote dominant and homozygote recessive, the offspring (F1-generation) will always have the heterozygote genotype and always present the phenotype associated with

4543-454: The parental phenotypes showed up in the offspring (green, round, red, or tall). However, when these hybrid plants were crossed, the offspring plants showed the two original phenotypes, in a characteristic 3:1 ratio, the more common phenotype being that of the parental hybrid plants. Mendel reasoned that each parent in the first cross was a homozygote for different alleles (one parent AA and the other parent aa), that each contributed one allele to

4620-441: The pedigrees and judges of all FCI members. Purebred dogs have more health problems than mongrel dogs, and require more veterinary visits, and tend to have lower longevity. Indeed, studies have reported lifespans that are shorter by between one and almost two years. Notably, dog breeds with flat faces and short noses have breathing difficulties, eye trouble and other health issues. The Fédération Cynologique Internationale

4697-499: The phenotype and neither allele masks another. For example, in the ABO blood group system , chemical modifications to a glycoprotein (the H antigen) on the surfaces of blood cells are controlled by three alleles, two of which are co-dominant to each other ( I , I ) and dominant over the recessive i at the ABO locus . The I and I alleles produce different modifications. The enzyme coded for by I adds an N-acetylgalactosamine to

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4774-469: The phenotype associated with the dominant allele variant. However, when crossing the F1-generation there are four possible phenotypic possibilities and the phenotypical ratio for the F2-generation will always be 9:3:3:1. Incomplete dominance (also called partial dominance , semi-dominance , intermediate inheritance , or occasionally incorrectly co-dominance in reptile genetics ) occurs when

4851-404: The phenotype of the heterozygous genotype is distinct from and often intermediate to the phenotypes of the homozygous genotypes. The phenotypic result often appears as a blended form of characteristics in the heterozygous state. For example, the snapdragon flower color is homozygous for either red or white. When the red homozygous flower is paired with the white homozygous flower, the result yields

4928-412: The phenotype variation. Indicators of recent selection were shown by many of the 51 genomic regions that were associated with traits that define a breed, which include body size, coat characteristics, and ear floppiness. Wolf Shar Pei Shiba Inu Chow Chow Akita Inu Basenji Siberian Husky Alaskan Malamute Afghan Hound Saluki other breeds in the study "Ancient breed"

5005-535: The phenotypic outcome. Although any individual of a diploid organism has at most two different alleles at a given locus, most genes exist in a large number of allelic versions in the population as a whole. This is called polymorphism , and is caused by mutations. Polymorphism can have an effect on the dominance relationship and phenotype, which is observed in the ABO blood group system . The gene responsible for human blood type have three alleles; A, B, and O, and their interactions result in different blood types based on

5082-407: The presence of the r allele, so these individuals also have round peas. Thus, allele R is dominant over allele r , and allele r is recessive to allele R . Dominance is not inherent to an allele or its traits ( phenotype ). It is a strictly relative effect between two alleles of a given gene of any function; one allele can be dominant over a second allele of the same gene, recessive to

5159-598: The product of the controlled breeding practices of the Victorian era (1830–1900). In 2010, a study looked at 48,000  single nucleotide polymorphisms that gave a genome -wide coverage of 912 dogs representing 85 breeds. The study found distinct genetic clusters within modern dogs that largely corresponded to phenotype or function. These included spitz-breeds, toy dogs, spaniels, Mastiff-like breeds, small terriers, retrievers, herding dogs, scent-hounds, and sight-hounds. There were 17 breeds that conflicted with phenotype or function and these were thought to be

5236-510: The products of the controlled breeding practices of the Victorian era (1830-1900), and the accurate documenting of pedigrees with the establishment of the English Kennel Club in 1873 in imitation of other stud book registries for cattle and horses. The domestic dog is the first species, and the only large carnivore , to have been domesticated. The first dogs were certainly wolflike, but the phenotypic changes that coincided with

5313-425: The purebreds were winning. Breed standards are the reason the breed came to be, and with those standards are key features, including form, function and fitness for purpose. The Kennel Club in the UK was founded in 1873, and was the world's first national kennel club and breed registry. The International Canine Federation was founded in 1911 as a worldwide organization. Its objective is to bring global uniformity to

5390-553: The recognition of distinct dog breeds, but there are many independent clubs with differing, and sometimes inconsistent standards and they need not apply scientific standards. Four varieties of the Belgian Shepherd Dog are recognised as four distinct breeds by the New Zealand Kennel Club . Further, some groups of dogs which clearly share a persistent set of characteristics and documented descent from

5467-638: The result of crossing some of the other phenotypes. As in a 2004 study that found 9 'ancient breeds' to be genetically divergent, the study found 13 breeds that were genetically divergent from the modern breeds: the Basenji , Saluki , Afghan hound , Samoyed , Canaan dog , New Guinea singing dog , dingo , Chow Chow , Chinese Shar Pei , Akita , Alaskan malamute , Siberian husky and American Eskimo dog . The study found that there were three well-supported groups that were highly divergent and distinct from modern domestic dogs. The earliest books in

5544-455: The rules for conformation dog shows and trials and accreditation of judges. They often serve as registries , which are lists of adult purebred dogs and lists of litters of puppies born to purebred parents. A dog breed is represented by a sufficient number of individuals to stably transfer its specific characteristics over generations. Dogs of same breed have similar characteristics of appearance and behavior, primarily because they come from

5621-464: The same breed. According to the Kennel Club's Purebred Health Survey, carried out in 2004, the Manchester Terrier has a median lifespan of around 12 years 10 months. Slightly more than 31 per cent of the animals whose deaths were reported in the survey died of old age. There are some recognised health problems in the breed associated with inheritable genetic mutations. Glaucoma is a fairly common genetic condition in several breeds of terrier, including

5698-477: The standard and toy varieties, but is seen more commonly in the toy. In the Manchester Terrier the gene is autosomal recessive , meaning that for any offspring to be affected the faulty gene must be carried by both parents. Affected dogs are prone to nose bleeds and bleeding from the gums. DNA tests are available to check for the condition, and the Kennel Club maintains a register of the results of those tests, allowing breeders to avoid breeding from affected dogs. At

5775-420: The two homozygotes, the phenotype is said to exhibit no dominance at all, i.e. dominance exists only when the heterozygote's phenotype measure lies closer to one homozygote than the other. When plants of the F 1 generation are self-pollinated, the phenotypic and genotypic ratio of the F 2 generation will be 1:2:1 (Red:Pink:White). Co-dominance occurs when the contributions of both alleles are visible in

5852-546: The use and description of various dog types is The Master of Game (circa 1406) by Edward of York , a treatise that describes dogs and their work, such as the alaunt , greyhound, pack scent hounds, spaniel , and mastiff , used by the privileged and wealthy for hunting purposes. The Master of Game is a combination of the earlier Art of Venery and the French hunting treatise Livre de Chasse by Gaston Phoebus (circa 1387). The Book of Saint Albans , published in 1486,

5929-478: The very recent product of cross-breeding other basal breeds. Most basal breeds have hybridized with other lineages in the past. If those other lineages were other basal breeds then a basal genetic signature remains. The combination of introgression and past population bottlenecks suggested that basal breeds have little or no genetic connections to their ancestral populations and that their genetic distinctiveness does not signify ancient heritage. They are distinctive from

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