Food energy is chemical energy that animals (including humans ) derive from their food to sustain their metabolism , including their muscular activity.
81-699: Miodowa (lit. Honey Street ) is a street in Warsaw's Old Town . More precisely, it links the Krakowskie (Cracow Suburb) Street in with Krasiński Square . It is also the name of a street in the Kazimierz district in Kraków . In the 16th century Miodowa Street was renowned for its gingerbread shops; hence its name. During the 18th century the Italian painter Bernardo Bellotto , better known as il Canaletto,
162-464: A glassy state and becomes an amorphous solid (noncrystalline). The viscosity of honey is affected greatly by both temperature and water content. The higher the water percentage, the more easily honey flows . Above its melting point, however, water has little effect on viscosity. Aside from water content, the composition of most types of honey also has little effect on viscosity. At 25 °C (77 °F), honey with 14% water content generally has
243-598: A supercooled solution of various sugars in water. These concentrations of sugar can only be achieved near room temperature by evaporation of a less concentrated solution, in this case nectar. For osmotic reasons such high concentrations of sugar are extremely unfavorable to microbiological reproduction and all fermentation is consequently halted. The bees then cap the cells of finished honey with wax. This seals them from contamination and prevents further evaporation. So long as its water concentration does not rise much above 18%, honey has an indefinite shelf life, both within
324-415: A complex dish or meal can be approximated by adding the energy contents of its components. The first determinations of the energy content of food were made by burning a dried sample in a bomb calorimeter and measuring the temperature change in the water surrounding the apparatus, a method known as direct calorimetry . However, the direct calorimetric method generally overestimates the actual energy that
405-408: A few types have non-Newtonian viscous properties. Honeys from heather or manuka display thixotropic properties. These types of honey enter a gel-like state when motionless, but liquefy when stirred. Because honey contains electrolytes , in the form of acids and minerals, it exhibits varying degrees of electrical conductivity . Measurements of the electrical conductivity are used to determine
486-555: A food for their larvae . To this end bees stockpile honey to provide for themselves during ordinary foraging as well as during lean periods, as in overwintering . During foraging bees use part of the nectar they collect to power their flight muscles. The majority of nectar collected is not used to directly nourish the insects but is instead destined for regurgitation , enzymatic digestion , and finally long-term storage as honey. During cold weather or when other food sources are scarce, adult and larval bees consume stored honey, which
567-512: A given mass of food is usually expressed in the metric (SI) unit of energy, the joule (J), and its multiple the kilojoule (kJ); or in the traditional unit of heat energy, the calorie (cal). In nutritional contexts, the latter is often (especially in US) the "large" variant of the unit, also written "Calorie" (with symbol Cal, both with capital "C") or "kilocalorie" (kcal), and equivalent to 4184 J or 4.184 kJ. Thus, for example, fats and ethanol have
648-469: A honey, which may be described as "sweet", "flowery", "citrus", "almond" or "rancid", among other terms. In addition, VOCs play a large role in determining the specific flavor of the honey, both through the aromas and flavor. VOCs from honeys in different geographic regions can be used as floral markers of those regions, and as markers of the bees that foraged the nectars. Honey is classified by its source (floral or not), and divisions are made according to
729-695: A long and varied history, with its beginnings in prehistoric times . Several cave paintings in Cuevas de la Araña in Spain depict humans foraging for honey at least 8,000 years ago. While Apis mellifera is an Old World insect , large-scale meliponiculture of New World stingless bees has been practiced by Mayans since pre-Columbian times. Honey is produced by bees who have collected nectar or honeydew . Bees value honey for its sugars, which they consume to support general metabolic activity , especially that of their flight muscles during foraging , and as
810-439: A manufacturer of rowing equipment shows calories released from "burning" food as four times the actual mechanical work, plus 1,300 kJ (300 kcal) per hour, which amounts to about 20% efficiency at 250 watts of mechanical output. It can take up to 20 hours of little physical output (e.g., walking) to "burn off" 17,000 kJ (4,000 kcal) more than a body would otherwise consume. For reference, each kilogram of body fat
891-436: A possible fire), making them less aggressive, and obscures the pheromones the bees use to communicate. The honeycomb is removed from the hive and the honey may be extracted from it either by crushing or by using a honey extractor . The honey is then usually filtered to remove beeswax and other debris. Before the invention of removable frames, bee colonies were often sacrificed to conduct the harvest. The harvester would take all
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#1732771846367972-631: A primary role in determining honey flavors and aromas . VOCs are carbon-based compounds that readily vaporize into the air, providing aroma, including the scents of flowers, essential oils, or ripening fruit. The typical chemical families of VOCs found in honey include hydrocarbons , aldehydes , alcohols , ketones , esters , acids , benzenes , furans , pyrans , norisoprenoids , and terpenes , among many others and their derivatives. The specific VOCs and their amounts vary considerably between different types of honey obtained by bees foraging on different plant sources. By example, when comparing
1053-449: A primary role in determining the exact temperature. Of these acids, the amino acids, which occur in very small amounts, play an important role in the darkening of honey. The amino acids form darkened compounds called melanoidins , during a Maillard reaction . The Maillard reaction occurs slowly at room temperature, taking from a few to several months to show visible darkening, but speeds up dramatically with increasing temperatures. However,
1134-443: A similar table. Other components of the human diet are either noncaloric, or are usually consumed in such small amounts that they can be neglected. The food energy actually obtained by respiration is used by the human body for a wide range of purposes, including basal metabolism of various organs and tissues, maintaining the internal body temperature , and exerting muscular force to maintain posture and produce motion. About 20%
1215-518: A solid), so melting crystallized honey can easily result in localized caramelization if the heat source is too hot or not evenly distributed. However, honey takes substantially longer to liquefy when just above the melting point than at elevated temperatures. Melting 20 kg (44 lb) of crystallized honey at 40 °C (104 °F) can take up to 24 hours, while 50 kg (110 lb) may take twice as long. These times can be cut nearly in half by heating at 50 °C (122 °F); however, many of
1296-418: A solution of fructose and other ingredients. The density of honey typically ranges between 1.38 and 1.45 kg/L at 20 °C. The melting point of crystallized honey is between 40 and 50 °C (104 and 122 °F), depending on its composition. Below this temperature, honey can be either in a metastable state, meaning that it will not crystallize until a seed crystal is added, or, more often, it
1377-463: A viscosity around 400 poise , while a honey containing 20% water has a viscosity around 20 poise. Viscosity increases very slowly with moderate cooling; a honey containing 16% water, at 70 °C (158 °F), has a viscosity around 2 poise, while at 30 °C (86 °F), the viscosity is around 70 poise. With further cooling, the increase in viscosity is more rapid, reaching 600 poise at around 14 °C (57 °F). However, while honey
1458-414: Is also affected by water content, because a high percentage of water inhibits crystallization, as does a high dextrin content. Temperature also affects the rate of crystallization, with the fastest growth occurring between 13 and 17 °C (55 and 63 °F). Crystal nuclei (seeds) tend to form more readily if the honey is disturbed, by stirring, shaking, or agitating, rather than if left at rest. However,
1539-607: Is an important food for virtually all hunter-gatherer cultures in warm climates, with the Hadza people ranking honey as their favorite food. Honey hunters in Africa have a mutualistic relationship with certain species of honeyguide birds. Possibly the world's oldest fermented beverage , dating from 9,000 years ago, mead ("honey wine") is the alcoholic product made by adding yeast to honey-water must and fermenting it for weeks or months. The yeast Saccharomyces cerevisiae
1620-408: Is collected from wild bee colonies or from domesticated beehives. On average, a hive will produce about 29 kilograms (65 lb) of honey per year. Wild bee nests are sometimes located by following a honeyguide bird. To safely collect honey from a hive, beekeepers typically pacify the bees using a bee smoker . The smoke triggers a feeding instinct (an attempt to save the resources of the hive from
1701-570: Is commonly used in modern mead production. Mead varieties include drinks called metheglin (with spices or herbs), melomel (with fruit juices, such as grape, specifically called pyment ), hippocras (with cinnamon ), and sack mead (high concentration of honey), many of which have been developed as commercial products numbering in the hundreds in the United States. Honey is also used to make mead beer , called "braggot". The physical properties of honey vary, depending on water content,
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#17327718463671782-473: Is equivalent to walking around 2.5 to 5 km ( 1 + 1 ⁄ 2 to 3 mi) per day in addition to the activities of sedentary living. These estimates are for a "reference woman" who is 1.63 m (5 ft 4 in) tall and weighs 57 kg (126 lb) and a "reference man" who is 1.78 m (5 ft 10 in) tall and weighs 70 kg (154 lb). Because caloric requirements vary by height, activity, age, pregnancy status, and other factors,
1863-594: Is in a "labile" state, being saturated with enough sugars to crystallize spontaneously. The rate of crystallization is affected by many factors, but the primary factor is the ratio of the main sugars: fructose to glucose. Honeys that are supersaturated with a very high percentage of glucose, such as brassica honey, crystallize almost immediately after harvesting, while honeys with a low percentage of glucose, such as chestnut or tupelo honey, do not crystallize. Some types of honey may produce few but very large crystals, while others produce many small crystals. Crystallization
1944-415: Is in their own honey stomachs, the hive bees regurgitate the nectar, repeatedly forming bubbles between their mandibles , speeding its digestion and concentration. These bubbles create a large surface area per volume and by this means the bees evaporate a portion of the nectar's water into the warm air of the hive. Hive bees form honey-processing groups. These groups work in relay, with one bee subjecting
2025-605: Is limiting access to humidity. In its cured state, honey has a sufficiently high sugar content to inhibit fermentation. If exposed to moist air, its hydrophilic properties pull moisture into the honey, eventually diluting it to the point that fermentation can begin. The long shelf life of honey is attributed to an enzyme found in the stomach of bees. The bees mix glucose oxidase with expelled nectar they previously consumed, creating two byproducts – gluconic acid and hydrogen peroxide , which are partially responsible for honey acidity and suppression of bacterial growth. Honey
2106-405: Is made and stored to nourish bee colonies. Bees produce honey by gathering and then refining the sugary secretions of plants (primarily floral nectar ) or the secretions of other insects, like the honeydew of aphids. This refinement takes place both within individual bees, through regurgitation and enzymatic activity, and during storage in the hive, through water evaporation that concentrates
2187-399: Is many times as energy-dense as the nectar from which it is made. After leaving the hive, a foraging bee collects sugar-rich nectar or honeydew. Nectar from the flower generally has a water content of 70 to 80% and is much less viscous than finished honey, which usually has a water content around 18%. The water content of honeydew from aphids and other true bugs is generally very close to
2268-421: Is normal. At very low temperatures, honey does not freeze solid; rather its viscosity increases. Like most viscous liquids , the honey becomes thick and sluggish with decreasing temperature. At −20 °C (−4 °F), honey may appear or even feel solid, but it continues to flow at very low rates. Honey has a glass transition between −42 and −51 °C (−44 and −60 °F). Below this temperature, honey enters
2349-573: Is often pasteurized , to kill any yeast, by heating it above 70 °C (158 °F). Like all sugar compounds, honey caramelizes if heated sufficiently, becoming darker in color, and eventually burns. However, honey contains fructose, which caramelizes at lower temperatures than glucose. The temperature at which caramelization begins varies, depending on the composition, but is typically between 70 and 110 °C (158 and 230 °F). Honey also contains acids, which act as catalysts for caramelization. The specific types of acids and their amounts play
2430-598: Is preferred because all bee colonies, even those in very large agricultural apiculture operations, readily leave the protection of humans in swarms that can establish successful wild colonies. Much of the effort in commercial beekeeping is dedicated to persuading a hive that is ready to swarm to produce more honeycomb in its present location. This is usually done by adding more space to the colony with honey supers , empty boxes placed on top of an existing colony. The bees can then usually be enticed to develop this empty space instead of dividing their colony through swarming. Honey
2511-454: Is roughly equivalent to 32,300 kilojoules of food energy (i.e., 3,500 kilocalories per pound or 7,700 kilocalories per kilogram). Many countries and health organizations have published recommendations for healthy levels of daily intake of food energy. For example, the United States government estimates 8,400 and 10,900 kJ (2,000 and 2,600 kcal) needed for women and men, respectively, between ages 26 and 45, whose total physical activity
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2592-441: Is sometimes adulterated by the addition of other sugars, syrups, or compounds to change its flavor or viscosity, reduce cost, or increase the fructose content to inhibit crystallization . Honey has been adulterated since ancient times, when honey was sometimes blended with plant syrups such as maple , birch , or sorghum and sold to customers as pure honey. Sometimes crystallized honey was mixed with flour or other fillers, hiding
2673-605: Is sweet because of its high concentrations of the monosaccharides fructose and glucose . It has about the same relative sweetness as sucrose (table sugar). One standard tablespoon (15 mL) of honey provides around 190 kilojoules (46 kilocalories) of food energy . It has attractive chemical properties for baking and a distinctive flavor when used as a sweetener. Most microorganisms cannot grow in honey and sealed honey therefore does not spoil . Samples of honey discovered in archaeological contexts have proven edible even after millennia. Honey use and production has
2754-460: Is the result of about 40% efficiency of generating ATP from the respiration of food, losses in converting energy from ATP into mechanical work inside the muscle, and mechanical losses inside the body. The latter two losses are dependent on the type of exercise and the type of muscle fibers being used (fast-twitch or slow-twitch). For an overall efficiency of 20%, one watt of mechanical power is equivalent to 18 kJ/h (4.3 kcal/h). For example,
2835-447: Is used for brain metabolism. The conversion efficiency of energy from respiration into muscular (physical) power depends on the type of food and on the type of physical energy usage (e.g., which muscles are used, whether the muscle is used aerobically or anaerobically ). In general, the efficiency of muscles is rather low: only 18 to 26% of the energy available from respiration is converted into mechanical energy. This low efficiency
2916-399: Is viscous, it has low surface tension of 50–60 mJ/m , making its wettability similar to water, glycerin , or most other liquids. The high viscosity and wettability of honey cause stickiness , which is a time-dependent process in supercooled liquids between the glass-transition temperature (T g ) and the crystalline-melting temperature. Most types of honey are Newtonian liquids , but
2997-443: The nucleation of microscopic seed-crystals is greatest between 5 and 8 °C (41 and 46 °F). Therefore, larger but fewer crystals tend to form at higher temperatures, while smaller but more-numerous crystals usually form at lower temperatures. Below 5 °C, the honey will not crystallize, thus the original texture and flavor can be preserved indefinitely. Honey is a supercooled liquid when stored below its melting point, as
3078-415: The stingless bee . Honey for human consumption is collected from wild bee colonies, or from the hives of domesticated bees. The honey produced by honey bees is the most familiar to humans, thanks to its worldwide commercial production and availability. The husbandry of bees is known as beekeeping or apiculture, with the cultivation of stingless bees usually referred to as meliponiculture . Honey
3159-608: The EU regulations and UK regulations, are: (1) Some polyols, like erythritol , are not digested and should be excluded from the count. (2) This entry exists in the EU regulations of 2008, but not in the UK regulations, according to which fibre shall not be counted. More detailed tables for specific foods have been published by many organizations, such as the United Nations Food and Agriculture Organization also has published
3240-670: The USDA created the DRI Calculator for Healthcare Professionals in order to determine individual caloric needs. According to the Food and Agriculture Organization of the United Nations , the average minimum energy requirement per person per day is about 7,500 kJ (1,800 kcal). Although the U.S. has changed over time with a growth in population and processed foods or food in general, Americans today have available roughly
3321-580: The United States, according to the National Honey Board, "Ensuring honey authenticity is one of the great challenges facing the honey industry today. Over the past half century, a number of honey testing methods have been developed to detect food fraud. To date, there is no single universal analytical method available which is capable of detecting all types of adulteration with adequate sensitivity." Isotope ratio mass spectrometry can be used to detect addition of corn syrup and cane sugar by
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3402-430: The ability to absorb moisture directly from the air, a phenomenon called hygroscopy . The amount of water the honey absorbs is dependent on the relative humidity of the air. Because honey contains yeast, this hygroscopic nature requires that honey be stored in sealed containers to prevent fermentation, which usually begins if the honey's water content rises much above 25%. Honey tends to absorb more water in this manner than
3483-555: The adulteration from buyers until the honey was liquefied. In modern times, the most common adulterant became clear, almost-flavorless corn syrup; the adulterated mixture can be very difficult to distinguish from pure honey. According to the Codex Alimentarius of the United Nations, any product labeled as "honey" or "pure honey" must be a wholly natural product, although labeling laws differ between countries. In
3564-438: The available honey and replace the entire colony the next spring. Since the invention of removable frames, the principles of husbandry led most beekeepers to ensure that their bees have enough stores to survive the winter, either by leaving some honey in the beehive or by providing the colony with a honey substitute such as sugar water or crystalline sugar (often in the form of a "candyboard"). The amount of food necessary to survive
3645-541: The average daily energy intake of 8,800 kJ (2,100 kcal). The minimum food energy intake is also higher in cold environments. Increased mental activity has been linked with moderately increased brain energy consumption . Many governments require food manufacturers to label the energy content of their products, to help consumers control their energy intake. To facilitate evaluation by consumers, food energy values (and other nutritional properties) in package labels or tables are often quoted for convenient amounts of
3726-450: The body can obtain from the food, because it also counts the energy contents of dietary fiber and other indigestible components, and does not allow for partial absorption and/or incomplete metabolism of certain substances. For this reason, today the energy content of food is instead obtained indirectly, by using chemical analysis to determine the amount of each digestible dietary component (such as protein, carbohydrates, and fats), and adding
3807-429: The body. For example, leucine plays an important role in the regulation of protein metabolism and suppresses an individual's appetite. Small amounts of essential fatty acids , constituents of some fats that cannot be synthesized by the human body, are used (and necessary) for other biochemical processes. The approximate food energy contents of various human diet components, to be used in package labeling according to
3888-481: The carbon isotopic signature . Addition of sugars originating from corn or sugar cane ( C4 plants , unlike the plants used by bees, and also sugar beet , which are predominantly C3 plants ) skews the isotopic ratio of sugars present in honey, but does not influence the isotopic ratio of proteins. In an unadulterated honey, the carbon isotopic ratios of sugars and proteins should match. Levels as low as 7% of addition can be detected. In 2020, global production of honey
3969-537: The chapel partially in front of the Krasiński Palace in the far distance can be seen on the left side the Lelewel Palace , another work of Efraim Szreger. 52°14′50″N 21°0′35″E / 52.24722°N 21.00972°E / 52.24722; 21.00972 Honey Honey is a sweet and viscous substance made by several species of bees , the best-known of which are honey bees . Honey
4050-427: The colony to a site for a new hive. Bees generally swarm before a suitable location for another hive has been discovered by scouts sent out for this purpose. Until such a location is found the swarm will simply conglomerate near the former hive, often from tree branches. These swarms are unusually docile and amenable to transport by humans. When provided with a suitable nesting site, such as a commercial Langstroth hive ,
4131-524: The court painter for Poland 's last king Stanisław August Poniatowski , painted with meticulous details the streets and architecture of 18th century Warsaw. Canaletto painted a view of the street with all the hustle and bustle of its traffic. The buildings are, on the right, the palace of Crown Marshal Jan Klemens Branicki (built in 1740) and, on the left, the palace of the Bishops of Cracow (built in 1622, rebuilt between 1760-1762 by Jakub Fontana ) and
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#17327718463674212-461: The diet, such as organic acids , polyols , and ethanol (drinking alcohol) may contribute to the energy input. Some diet components that provide little or no food energy, such as water , minerals , vitamins , cholesterol , and fiber , may still be necessary to health and survival for other reasons. Some organisms have instead anaerobic respiration , which extracts energy from food by reactions that do not require oxygen. The energy contents of
4293-493: The dry weight of food. Ruminants can extract food energy from the respiration of cellulose because of bacteria in their rumens that decompose it into digestible carbohydrates. Other minor components of the human diet that contribute to its energy content are organic acids such as citric and tartaric , and polyols such as glycerol , xylitol , inositol , and sorbitol . Some nutrients have regulatory roles affected by cell signaling , in addition to providing energy for
4374-497: The flora also have an effect on honey aroma properties, such as a "fruity" or "grassy" aroma from longan honey, or a "waxy" aroma from sunflower honey. Dominant VOCs in one study were linalool oxide , trans-linalool oxide, 2- phenylacetaldehyde , benzyl ethanol , isophorone , and methyl nonanoate . VOCs can also be introduced from the bodies of the bees, be produced by the enzymatic actions of digestion, or from chemical reactions that occur between different substances within
4455-871: The greatest amount of food energy per unit mass, 37 and 29 kJ/g (9 and 7 kcal/g), respectively. Proteins and most carbohydrates have about 17 kJ/g (4 kcal/g), though there are differences between different kinds. For example, the values for glucose, sucrose, and starch are 15.57, 16.48 and 17.48 kilojoules per gram (3.72, 3.94 and 4.18 kcal/g) respectively. The differing energy density of foods (fat, alcohols, carbohydrates and proteins) lies mainly in their varying proportions of carbon, hydrogen, and oxygen atoms. Carbohydrates that are not easily absorbed, such as fibre, or lactose in lactose-intolerant individuals , contribute less food energy. Polyols (including sugar alcohols ) and organic acids contribute 10 kJ/g (2.4 kcal/g) and 13 kJ/g (3.1 kcal/g) respectively. The energy contents of
4536-475: The hive and after its removal by a beekeeper . Honey bees are not the only eusocial insects to produce honey. All non-parasitic bumblebees and stingless bees produce honey. Some wasp species, such as Brachygastra lecheguana and Brachygastra mellifica , found in South and Central America, are known to feed on nectar and produce honey. Other wasps, such as Polistes versicolor , also consume honey. In
4617-466: The honey during storage, and therefore may change, increase, or decrease over long periods of time. VOCs may be produced, altered, or greatly affected by temperature and processing. Some VOCs are heat labile, and are destroyed at elevated temperatures, while others can be created during non-enzymatic reactions, such as the Maillard reaction . VOCs are responsible for nearly all of the aroma produced by
4698-404: The honey stomach holds about 40 mg of liquid. This is about half the weight of an unladen bee. Collecting this quantity in nectar can require visits to more than a thousand flowers. When nectar is plentiful, it can take a bee more than an hour of ceaseless work to collect enough nectar to fill its honey crop. Salivary enzymes and proteins from the bee's hypopharyngeal gland are secreted into
4779-412: The honey's sugars until it is thick and viscous. Honey bees stockpile honey in the hive . Within the hive is a structure made from wax called honeycomb . The honeycomb is made up of hundreds or thousands of hexagonal cells, into which the bees regurgitate honey for storage. Other honey-producing species of bee store the substance in different structures, such as the pots made of wax and resin used by
4860-431: The honey-storage areas. This temperature is regulated either by generating heat with their bodies or removing it through water evaporation. The evaporation removes water from the stored honey, drawing heat from the colony. The bees use their wings to govern hive cooling. Coordinated wing beating moves air across the wet honey, drawing out water and heat. Ventilation of the hive eventually expels both excess water and heat into
4941-447: The individual sugars allow on their own, which may be due to other ingredients it contains. Fermentation of honey usually occurs after crystallization, because without the glucose, the liquid portion of the honey primarily consists of a concentrated mixture of fructose, acids, and water, providing the yeast with enough of an increase in the water percentage for growth. Honey that is to be stored at room temperature for long periods of time
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#17327718463675022-575: The mansion of the Warsaw banker, Piotr Tepper (built in 1774 according to design by Efraim Szreger , destroyed during the Warsaw Uprising and not rebuilt after the war). Further back can be seen the roof of the 17th-century church of the Capuchins, founded by King John III Sobieski (built between 1683-1694 by Tylman van Gameren ), which bas a chapel containing the urn with his heart. Behind
5103-494: The middle of their life cycles they alternate between feeding on protein-rich pollen and feeding on honey, which is a far denser source of food energy . Human beings have semi-domesticated several species of honey bee by taking advantage of their swarming stage. Swarming is the means by which new colonies are established when there is no longer space for expansion in the colony's present hive. The old queen lays eggs that will develop into new queens and then leads as many as half
5184-415: The minor substances in honey can be affected greatly by heating, changing the flavor, aroma, or other properties, so heating is usually done at the lowest temperature and for the shortest time possible. The average pH of honey is 3.9, but can range from 3.4 to 6.1. Honey contains many kinds of acids, both organic and amino . However, the different types and their amounts vary considerably, depending on
5265-567: The mixture of VOCs in different honeys in one review, longan honey had a higher amount of volatiles (48 VOCs), while sunflower honey had the lowest number of volatiles (8 VOCs). VOCs are primarily introduced into the honey from the nectar, where they are excreted by the flowers imparting individual scents. The specific types and concentrations of certain VOCs can be used to determine the type of flora used to produce monofloral honeys. The specific geography, soil composition and acidity used to grow
5346-402: The nectar once it is in the bee's honey stomach. These substances begin cleaving complex sugars like sucrose and starches into simpler sugars such as glucose and fructose . This process slightly raises the water content and the acidity of the partially digested nectar. Once filled, the forager bees return to the hive. There they regurgitate and transfer nectar to hive bees. Once it
5427-402: The outside world. The process of evaporating continues until the honey reaches its final water content of between 15.5% to 18%. This concentrates the sugars far beyond the saturation point of water, which is to say there is far more sugar dissolved in what little water remains in honey than ever could be dissolved in an equivalent volume of water. Honey, even at hive temperatures, is therefore
5508-605: The packaging and processing used. Regional honeys are also identified. In the US, honey is also graded on its color and optical density by USDA standards, graded on the Pfund scale, which ranges from 0 for "water white" honey to more than 114 for "dark amber" honey. Food energy Most animals derive most of their energy from aerobic respiration , namely combining the carbohydrates , fats , and proteins with oxygen from air or dissolved in water . Other smaller components of
5589-456: The polarization plane. The fructose gives a negative rotation, while the glucose gives a positive one. The overall rotation can be used to measure the ratio of the mixture. Honey is generally pale yellow and dark brown in color, but other colors can occur, depending on the sugar source. Bee colonies that forage on Kudzu ( Pueraria montana var. lobata ) flowers, for example, produce honey that varies in color from red to purple. Honey has
5670-431: The processed nectar to bubbling and then passing the refined liquid on to others. It can take as long as 20 minutes of continuous regurgitation, digestion and evaporation until the product reaches storage quality. The new honey is then placed in honeycomb cells, which are left uncapped. This honey still has a very high water content, up to 70%, depending on the concentration of nectar gathered. At this stage of its refinement
5751-415: The quality of honey in terms of ash content. The effect honey has on light is useful for determining the type and quality. Variations in its water content alter its refractive index . Water content can easily be measured with a refractometer . Typically, the refractive index for honey ranges from 1.504 at 13% water content to 1.474 at 25%. Honey also has an effect on polarized light , in that it rotates
5832-435: The reaction can also be slowed by storing the honey at colder temperatures. Unlike many other liquids, honey has very poor thermal conductivity of 0.5 W/(m⋅K) at 13% water content (compared to 401 W/(m⋅K) of copper ), taking a long time to reach thermal equilibrium . Due to its high kinematic viscosity honey does not transfer heat through momentum diffusion ( convection ) but rather through thermal diffusion (more like
5913-853: The respective food energy contents, previously obtained by measurement of metabolic heat released by the body. In particular, the fibre content is excluded. This method is known as the Modified Atwater system, after Wilbur Atwater who pioneered these measurements in the late 19th century. The system was later improved by Annabel Merrill and Bernice Watt of the USDA , who derived a system whereby specific calorie conversion factors for different foods were proposed. The typical human diet consists chiefly of carbohydrates, fats, proteins, water, ethanol, and indigestible components such as bones , seeds , and fibre (mostly cellulose ). Carbohydrates, fats, and proteins typically comprise ninety percent of
5994-425: The same level of calories as the older generation. [1] Older people and those with sedentary lifestyles require less energy; children and physically active people require more. Recognizing these factors, Australia's National Health and Medical Research Council recommends different daily energy intakes for each age and gender group. Notwithstanding, nutrition labels on Australian food products typically recommend
6075-503: The sap on which those insects feed and is usually somewhat more dilute than nectar. One source describes the water content of honeydew as around 89%. Whether it is feeding on nectar or honeydew, the bee sucks these runny fluids through its proboscis , which delivers the liquid to the bee's honey stomach or "honey crop". This cavity lies just above its food stomach, the latter of which digests pollen and sugars consumed by an individual honey bee for its own nourishment. In Apis mellifera ,
6156-441: The swarm will readily form a new colony in artificial surroundings. These semi-domesticated colonies are then looked after by humans practicing apiculture or meliponiculture. Captured bees are encouraged to forage, often in agricultural settings such as orchards, where pollinators are highly valued. The honey, pollen , wax and resins the bees produce are all harvested by humans for a variety of uses. The term "semi-domesticated"
6237-425: The type of flora used to produce it (pasturage), temperature, and the proportion of the specific sugars it contains. Fresh honey is a supersaturated liquid, containing more sugar than the water can typically dissolve at ambient temperatures. At room temperature, honey is a supercooled liquid, in which the glucose precipitates into solid granules. This forms a semisolid solution of precipitated glucose crystals in
6318-717: The type of honey. These acids may be aromatic or aliphatic (nonaromatic). The aliphatic acids contribute greatly to the flavor of honey by interacting with the flavors of other ingredients. Organic acids comprise most of the acids in honey, accounting for 0.17–1.17% of the mixture, with gluconic acid formed by the actions of glucose oxidase as the most prevalent. Minor amounts of other organic acids are present, consisting of formic , acetic , butyric , citric , lactic , malic , pyroglutamic , propionic , valeric , capronic , palmitic , and succinic , among many others. Individual honeys from different plant sources contain over 100 volatile organic compounds (VOCs), which play
6399-500: The water content of the honey is high enough that ubiquitous yeast spores can reproduce in it, a process which, if left unchecked, would rapidly consume the new honey's sugars. To combat this, bees use an ability rare among insects: the endogenous generation of heat. Bees are among the few insects that can create large amounts of body heat. They use this ability to produce a constant ambient temperature in their hives. Hive temperatures are usually around 35 °C (95 °F) in
6480-613: The winter depends on the variety of bees and on the length and severity of local winters. Many animal species are attracted to wild or domestic sources of honey. Because of its composition and chemical properties, honey is suitable for long-term storage, and is easily assimilated even after long preservation. Honey, and objects immersed in honey, have been preserved for centuries. (However, no edible honey has been found in Egyptian tombs; all such cases have been proven to be other substances or only chemical traces. ) The key to preservation
6561-401: Was 1.8 million tonnes , led by China with 26% of the world total (table). Other major producers were Turkey , Iran , Argentina , and Ukraine . Over its history as a food, the main uses of honey are in cooking, baking, desserts, as a spread on bread, as an addition to various beverages such as tea, and as a sweetener in some commercial beverages. Due to its energy density, honey
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