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44-565: During this period, mammals and birds continued to evolve into modern forms, while other groups of life remained relatively unchanged. The first humans ( Homo habilis ) appeared in Africa near the end of the period. Some continental movements took place, the most significant event being the connection of North and South America at the Isthmus of Panama , late in the Pliocene. This cut off

88-480: A broad neocortex region of the brain, fur or hair , and three middle ear bones . These characteristics distinguish them from reptiles and birds , from which their ancestors diverged in the Carboniferous Period over 300 million years ago. Around 6,400 extant species of mammals have been described and divided into 27 orders . The largest orders of mammals, by number of species , are

132-767: A decrease in global temperatures termed the Late Miocene Cooling (LMC) ensued, driven by decreases in carbon dioxide concentrations. During the Pliocene, from about 5.3 to 2.7 Ma, another warm interval occurred, being known as the Pliocene Warm Interval (PWI), interrupting the longer-term cooling trend. The Pliocene Thermal Maximum (PTM) occurred between 3.3 and 3.0 Ma. During the Pliocene, Green Sahara phases of wet conditions in North Africa were frequent and occurred about every 21 kyr, being especially intense when Earth's orbit's eccentricity

176-399: A phylogenetic study of crocodyliforms, Benton and Clark (1988) split up Sebecosuchia, finding baurusuchids to be basal notosuchians while sebecids were basal neosuchians . Since that time, most studies have supported a monophyletic Sebecosuchia. In 2007, however, a phylogenetic study placed baurusuchids as basal metasuchians and sebecids as close relatives to a family of notosuchians called

220-553: A positive feedback as sea levels dropped and the ITF diminished and further limited the heat transported southward by the Leeuwin Current . By the end of the period the first of a series of glaciations of the current Ice Age began. Marine and continental flora and fauna have a modern appearance. The reptile group Choristodera went extinct in the early part of the period, while the amphibians known as Allocaudata disappeared at

264-720: Is an extinct group of mesoeucrocodylian crocodyliforms that includes the families Sebecidae and Baurusuchidae . The group was long thought to have first appeared in the Late Cretaceous with the baurusuchids and become extinct in the Miocene with the last sebecids, but Razanandrongobe pushes the origin of Sebecosuchia to the Middle Jurassic. Fossils have been found primarily from South America but have also been found in Europe , North Africa , Madagascar , and

308-480: Is due to the comparatively fine divisibility of time units as time approaches the present, and due to geological preservation that causes the youngest sedimentary geological record to be preserved over a much larger area and to reflect many more environments than the older geological record. By dividing the Cenozoic Era into three (arguably two) periods ( Paleogene , Neogene, Quaternary ) instead of seven epochs,

352-572: Is primarily driven by human poaching and habitat destruction , primarily deforestation . Over 70% of mammal species are in the orders Rodentia , Chiroptera , and Eulipotyphla . Mammal classification has been through several revisions since Carl Linnaeus initially defined the class, and at present , no classification system is universally accepted. McKenna & Bell (1997) and Wilson & Reeder (2005) provide useful recent compendiums. Simpson (1945) provides systematics of mammal origins and relationships that had been taught universally until

396-647: Is subdivided into six ages: In different geophysical regions of the world, other regional names are also used for the same or overlapping ages and other timeline subdivisions. The terms Neogene System (formal) and Upper Tertiary System (informal) describe the rocks deposited during the Neogene Period . The continents in the Neogene were very close to their current positions. The Isthmus of Panama formed, connecting North and South America . The Indian subcontinent continued to collide with Asia , forming

440-426: Is the viviparous placental mammals , so named for the temporary organ ( placenta ) used by offspring to draw nutrition from the mother during gestation . Most mammals are intelligent , with some possessing large brains, self-awareness , and tool use . Mammals can communicate and vocalize in several ways, including the production of ultrasound , scent marking , alarm signals , singing , echolocation ; and, in

484-473: The Cretaceous . The relationships between these three lineages is contentious, and all three possible hypotheses have been proposed with respect to which group is basal . These hypotheses are Atlantogenata (basal Boreoeutheria), Epitheria (basal Xenarthra) and Exafroplacentalia (basal Afrotheria). Boreoeutheria in turn contains two major lineages— Euarchontoglires and Laurasiatheria . Estimates for

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528-516: The Himalayas . Sea levels fell, creating land bridges between Africa and Eurasia and between Eurasia and North America. The global climate became more seasonal and continued an overall drying and cooling trend which began during the Paleogene . The Early Miocene was relatively cool; Early Miocene mid-latitude seawater and continental thermal gradients were already very similar to those of

572-564: The International Union for Conservation of Nature (IUCN) completed a five-year Global Mammal Assessment for its IUCN Red List , which counted 5,488 species. According to research published in the Journal of Mammalogy in 2018, the number of recognized mammal species is 6,495, including 96 recently extinct. The word " mammal " is modern, from the scientific name Mammalia coined by Carl Linnaeus in 1758, derived from

616-556: The Latin mamma ("teat, pap"). In an influential 1988 paper, Timothy Rowe defined Mammalia phylogenetically as the crown group of mammals, the clade consisting of the most recent common ancestor of living monotremes ( echidnas and platypuses ) and therian mammals ( marsupials and placentals ) and all descendants of that ancestor. Since this ancestor lived in the Jurassic period, Rowe's definition excludes all animals from

660-655: The Peirosauridae . Together, sebecids and peirosaurids made the new clade Sebecia . Below is a cladogram from that study, Larsson and Sues (2007): Notosuchus Malawisuchus Araripesuchus Baurusuchus Neosuchia Pabwehshi Sebecus Bretesuchus Peirosauridae Two years later, Sereno and Larsson (2009) came to the same conclusion, except they placed baurusuchids as advanced notosuchians. More recently however, Turner and Sertich (2010) found support for Sebecosuchia in their analysis of notosuchian relationships. In their study, Sebecosuchia

704-606: The biological classification scheme used, are the primates : apes , monkeys , and lemurs ; the Cetartiodactyla : whales and even-toed ungulates ; and the Carnivora which includes cats , dogs , weasels , bears , seals , and allies. According to Mammal Species of the World , 5,416 species were identified in 2006. These were grouped into 1,229  genera , 153  families and 29 orders. In 2008,

748-406: The dominant terrestrial animal group from 66 million years ago to the present. The basic mammalian body type is quadrupedal , with most mammals using four limbs for terrestrial locomotion ; but in some, the limbs are adapted for life at sea , in the air , in trees or underground . The bipeds have adapted to move using only the two lower limbs, while the rear limbs of cetaceans and

792-588: The mammoths and woolly rhinoceros were common in Pliocene . With lower levels of CO 2 in the atmosphere, C 4 plants expanded and reached ecological dominance in grasslands during the last 10 million years. Also Asteraceae (daisies) went through a significant adaptive radiation . Eucalyptus fossil leaves occur in the Miocene of New Zealand, where the genus is not native today, but have been introduced from Australia. The Neogene traditionally ended at

836-428: The rodents , bats , and Eulipotyphla (including hedgehogs , moles and shrews ). The next three are the primates (including humans , monkeys and lemurs ), the even-toed ungulates (including pigs , camels , and whales ), and the Carnivora (including cats , dogs , and seals ). Mammals are the only living members of Synapsida ; this clade , together with Sauropsida (reptiles and birds), constitutes

880-439: The sea cows are mere internal vestiges . Mammals range in size from the 30–40 millimetres (1.2–1.6 in) bumblebee bat to the 30 metres (98 ft) blue whale —possibly the largest animal to have ever lived. Maximum lifespan varies from two years for the shrew to 211 years for the bowhead whale . All modern mammals give birth to live young, except the five species of monotremes , which lay eggs. The most species-rich group

924-584: The Gelasian Age, which was formerly considered part of the Neogene Period and Pliocene Epoch. Thus the Neogene Period ends bounding the succeeding Quaternary Period at 2.58 Mya. Mammal A mammal (from Latin mamma  'breast') is a vertebrate animal of the class Mammalia ( / m ə ˈ m eɪ l i . ə / ). Mammals are characterized by the presence of milk -producing mammary glands for feeding their young,

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968-535: The Indian subcontinent. Sebecosuchia was first constructed in 1937 by George Gaylord Simpson . In 1946 the concept was again used by American paleontologist Edwin Colbert to include Sebecus and Baurusuchidae. Sebecus , which had been known from South America since 1937, was an unusual crocodyliform with a deep snout and teeth that were ziphodont, or serrated and laterally compressed. The family Baurusuchidae

1012-772: The Neogene and the Pliocene end at 2.58 Ma, that the Gelasian be transferred to the Pleistocene, and the Quaternary be recognized as the third period in the Cenozoic, citing key changes in Earth's climate, oceans, and biota that occurred 2.58 Ma and its correspondence to the Gauss-Matuyama magnetostratigraphic boundary . In 2006 ICS and INQUA reached a compromise that made Quaternary a sub-era, subdividing Cenozoic into

1056-492: The case of humans, complex language . Mammals can organize themselves into fission–fusion societies , harems , and hierarchies —but can also be solitary and territorial . Most mammals are polygynous , but some can be monogamous or polyandrous . Domestication of many types of mammals by humans played a major role in the Neolithic Revolution , and resulted in farming replacing hunting and gathering as

1100-757: The clade originating with the last common ancestor of Sinoconodon and living mammals. The earliest-known synapsid satisfying Kemp's definitions is Tikitherium , dated 225 Ma , so the appearance of mammals in this broader sense can be given this Late Triassic date. However, this animal may have actually evolved during the Neogene. As of the early 21st century, molecular studies based on DNA analysis have suggested new relationships among mammal families. Most of these findings have been independently validated by retrotransposon presence/absence data . Classification systems based on molecular studies reveal three major groups or lineages of placental mammals— Afrotheria , Xenarthra and Boreoeutheria —which diverged in

1144-444: The closest thing to an official classification of mammals, despite its known issues. Most mammals, including the six most species-rich orders , belong to the placental group. The three largest orders in numbers of species are Rodentia : mice , rats , porcupines , beavers , capybaras , and other gnawing mammals; Chiroptera : bats; and Eulipotyphla : shrews , moles , and solenodons . The next three biggest orders, depending on

1188-1349: The divergence times between these three placental groups range from 105 to 120 million years ago, depending on the type of DNA used (such as nuclear or mitochondrial ) and varying interpretations of paleogeographic data. Monotremata Marsupialia Xenarthra Afrotheria Glires Euarchonta [REDACTED] Eulipotyphla Chiroptera Pholidota Carnivora [REDACTED] Perissodactyla [REDACTED] Artiodactyla [REDACTED] Monotremata [REDACTED] Paucituberculata [REDACTED] Didelphimorphia [REDACTED] Microbiotheria Notoryctemorphia [REDACTED] Peramelemorphia [REDACTED] Dasyuromorphia [REDACTED] Diprotodontia [REDACTED] Cingulata [REDACTED] Pilosa [REDACTED] Hyracoidea [REDACTED] Sirenia [REDACTED] Proboscidea [REDACTED] Tubulidentata [REDACTED] Macroscelidea [REDACTED] Afrosoricida [REDACTED] Eulipotyphla [REDACTED] Chiroptera [REDACTED] Pholidota [REDACTED] Carnivora [REDACTED] Perissodactyla [REDACTED] Artiodactyla [REDACTED] Scandentia [REDACTED] Lagomorpha [REDACTED] Sebecosuchia Sebecosuchia (meaning " Sobek crocodiles")

1232-503: The dominant terrestrial vertebrates, and took many forms as they adapted to various habitats. An explosive radiation of ursids took place at the Miocene-Pliocene boundary. The first hominins , the ancestors of humans, may have appeared in southern Europe and migrated into Africa. The first humans (belonging to the species Homo habilis ) appeared in Africa near the end of the period. About 20 million years ago gymnosperms in

1276-585: The earlier Triassic , despite the fact that Triassic fossils in the Haramiyida have been referred to the Mammalia since the mid-19th century. If Mammalia is considered as the crown group, its origin can be roughly dated as the first known appearance of animals more closely related to some extant mammals than to others. Ambondro is more closely related to monotremes than to therian mammals while Amphilestes and Amphitherium are more closely related to

1320-404: The end of it. Neogene also marked the end of the reptilian genera Langstonia and Barinasuchus , terrestrial predators that were the last surviving members of Sebecosuchia , a group related to crocodiles. The oceans were dominated by large carnivores like megalodons and livyatans , and 19 million years ago about 70% of all pelagic shark species disappeared. Mammals and birds continued to be

1364-451: The end of the 20th century. However, since 1945, a large amount of new and more detailed information has gradually been found: The paleontological record has been recalibrated, and the intervening years have seen much debate and progress concerning the theoretical underpinnings of systematization itself, partly through the new concept of cladistics . Though fieldwork and lab work progressively outdated Simpson's classification, it remains

Neogene - Misplaced Pages Continue

1408-677: The end of the Pliocene Epoch, just before the older definition of the beginning of the Quaternary Period; many time scales show this division. However, there was a movement amongst geologists (particularly marine geologists ) to also include ongoing geological time (Quaternary) in the Neogene, while others (particularly terrestrial geologists) insist the Quaternary to be a separate period of distinctly different record. The somewhat confusing terminology and disagreement amongst geologists on where to draw what hierarchical boundaries

1452-450: The form of some conifer and cycad groups started to diversify and produce more species due to the changing conditions. In response to the cooler, seasonal climate, tropical plant species gave way to deciduous ones and grasslands replaced many forests. Grasses therefore greatly diversified, and herbivorous mammals evolved alongside it, creating the many grazing animals of today such as horses , antelope , and bison . Ice age mammals like

1496-612: The larger Amniota clade. Early synapsids are referred to as " pelycosaurs ." The more advanced therapsids became dominant during the Guadalupian . Mammals originated from cynodonts , an advanced group of therapsids, during the Late Triassic to Early Jurassic . Mammals achieved their modern diversity in the Paleogene and Neogene periods of the Cenozoic era, after the extinction of non-avian dinosaurs , and have been

1540-420: The last common ancestor and all descendants of baurusuchids and sebecids. More recently, other crocodyliforms have been assigned to Sebecosuchia that cannot be placed into either family. These include the genera Eremosuchus , named in 1989, and Pehuenchesuchus , named in 2005. They are usually considered to be more basal sebecosuchians than the sebecids and baurusuchids. Below is a cladogram showing

1584-609: The old classical Tertiary and Quaternary, a compromise that was rejected by International Union of Geological Sciences because it split both Neogene and Pliocene in two. Following formal discussions at the 2008 International Geological Congress in Oslo, Norway, the ICS decided in May 2009 to make the Quaternary the youngest period of the Cenozoic Era with its base at 2.58 Mya and including

1628-627: The periods are more closely comparable to the duration of periods in the Mesozoic and Paleozoic Eras. The International Commission on Stratigraphy (ICS) once proposed that the Quaternary be considered a sub-era (sub-erathem) of the Neogene, with a beginning date of 2.58 Ma, namely the start of the Gelasian Stage . In the 2004 proposal of the ICS, the Neogene would have consisted of the Miocene and Pliocene Epochs. The International Union for Quaternary Research (INQUA) counterproposed that

1672-609: The possible phylogenetic position of Sebecosuchia modified from Turner and Calvo (2005). In this cladogram, Sebecidae is a paraphyletic assemblage of basal sebecosuchians while Baurusuchidae is monophyletic and includes the more derived sebecosuchians. Hsisosuchus Thalattosuchia Araripesuchus Trematochampsidae Peirosauridae Mahajangasuchidae Atoposauridae Goniopholididae Bernissartia Crocodylia Libycosuchus Notosuchia Pehuenchesuchus Eremosuchus Sebecus Iberosuchus Bretesuchus Baurusuchus Pabwehshi In

1716-751: The present. During the Middle Miocene , Earth entered a warm phase known as the Middle Miocene Climatic Optimum (MMCO), which was driven by the emplacement of the Columbia River Basalt Group . Around 11 Ma, the Middle Miocene Warm Interval gave way to the much cooler Late Miocene. The ice caps on both poles began to grow and thicken, a process enhanced by positive feedbacks from increased formation of sea ice. Between 7 and 5.3 Ma,

1760-716: The primary source of food for humans. This led to a major restructuring of human societies from nomadic to sedentary, with more co-operation among larger and larger groups, and ultimately the development of the first civilizations . Domesticated mammals provided, and continue to provide, power for transport and agriculture, as well as food ( meat and dairy products ), fur , and leather . Mammals are also hunted and raced for sport, kept as pets and working animals of various types, and are used as model organisms in science. Mammals have been depicted in art since Paleolithic times, and appear in literature, film, mythology, and religion. Decline in numbers and extinction of many mammals

1804-548: The therians; as fossils of all three genera are dated about 167 million years ago in the Middle Jurassic , this is a reasonable estimate for the appearance of the crown group. T. S. Kemp has provided a more traditional definition: " Synapsids that possess a dentary – squamosal jaw articulation and occlusion between upper and lower molars with a transverse component to the movement" or, equivalently in Kemp's view,

Neogene - Misplaced Pages Continue

1848-678: The warm ocean currents from the Pacific to the Atlantic Ocean, leaving only the Gulf Stream to transfer heat to the Arctic Ocean . The global climate cooled considerably throughout the Neogene, culminating in a series of continental glaciations in the Quaternary Period that followed. In ICS terminology, from upper (later, more recent) to lower (earlier): The Pliocene Epoch is subdivided into two ages: The Miocene Epoch

1892-552: Was high. The PWI had similar levels of atmospheric carbon dioxide to contemporary times and is often seen as an analogous climate to the projected climate of the near future as a result of anthropogenic global warming . Towards the end of the Pliocene, decreased heat transport towards the Antarctic resulting from a weakening of the Indonesian Throughflow (ITF) cooled the Earth, a process that exacerbated itself in

1936-721: Was named the year before and included the newly described Baurusuchus , which was also a South American deep-snouted form. The clade Sebecosuchia was given a phylogenetic definition in the PhyloCode by Juan Leardi and colleagues in 2024 as "the least inclusive clade containing Sebecus icaeorhinus and Baurusuchus pachecoi provided that it doesn’t include Araripesuchus gomesii , Montealtosuchus arrudacamposi , or Crocodylus niloticus (the Nile crocodile)". This definition ensures that Sebecosuchia self destructs if peirosaurids , uruguaysuchids , or modern crocodylians fall within

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