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106-689: The snow leopard ( Panthera uncia ) is a species of large cat in the genus Panthera of the family Felidae . The species is native to the mountain ranges of Central and South Asia . It is listed as Vulnerable on the IUCN Red List because the global population is estimated to number fewer than 10,000 mature individuals and is expected to decline about 10% by 2040. It is mainly threatened by poaching and habitat destruction following infrastructural developments. It inhabits alpine and subalpine zones at elevations of 3,000–4,500 m (9,800–14,800 ft), ranging from eastern Afghanistan ,

212-602: A common ancestor ) or paraphyletic (excluding some descendants), these concepts do not apply to monotypic taxa because they contain only a single member. Monotypic taxa are part of a broader challenge in biological classification known as aphyly – situations where evolutionary relationships are poorly supported by evidence. This includes both monotypic groups and cases where traditional groupings are found to be artificial. Understanding how monotypic taxa fit into this bigger picture helps identify areas needing further research. The German lichenologist Robert Lücking suggests that

318-579: A criminal offence that is punishable by the confiscation of property, a fine and a sentence of at least 10 years in prison. They have been protected in Bhutan since 1995. At the end of 2020, 35 cameras were installed on the outskirts of Almaty, Kazakhstan in hopes to catch footage of snow leopards. In November 2021, it was announced by the Russian World Wildlife Fund (WWF) that snow leopards were spotted 65 times on these cameras in

424-482: A hierarchical system. When taxonomists identify a monotypic taxon, this often reflects uncertainty about its relationships rather than true evolutionary isolation . This uncertainty is evident in many cases across different species. For instance, the diatom Licmophora juergensii is placed in a monotypic genus because scientists have not yet found clear evidence of its relationships to other species. Some taxonomists argue against monotypic taxa because they reduce

530-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles

636-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed

742-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this

848-688: A Russian scientist in 2000 was a dark brown snow leopard skin from the Petrovsk-Zabaykalsky District in southern Transbaikal . The snow leopard was long classified in the monotypic genus Uncia . They were subordinated to the genus Panthera based on results of phylogenetic studies. Until spring 2017, there was no evidence available for the recognition of subspecies . Results of a phylogeographic analysis indicate that three subspecies should be recognised: This view has been both contested and supported by different researchers. An extinct subspecies Panthera uncia pyrenaica

954-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing

1060-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by

1166-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider

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1272-465: A lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from the time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On

1378-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides

1484-553: A natural classification. From a cladistic perspective, which focuses on shared derived characteristics to determine evolutionary relationships, the theoretical status of monotypic taxa is complex. Some argue they can only be justified when relationships cannot be resolved through synapomorphies (shared derived characteristics); otherwise, they would necessarily exclude related species and thus be paraphyletic. However, others contend that while most taxonomic groups can be classified as either monophyletic (containing all descendants of

1590-505: A noticeable increase in marking and calling. Females have a gestation period of 90–100 days, and the cubs are born between April and June. A litter usually consists of two to three cubs, in exceptional cases there can be up to seven. The female gives birth in a rocky den or crevice lined with fur shed from her underside. The cubs are born blind and helpless, although already with a thick coat of fur, and weigh 320 to 567 g (11.3 to 20.0 oz). Their eyes open at around seven days, and

1696-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as

1802-400: A perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that

1908-459: A process known as false splitting . The word once was originally considered to be pronounced as l'once , where l' stands for the elided form of the word la ('the') in French. Once was then understood to be the name of the animal. The word panther derives from the classical Latin panthēra , itself from the ancient Greek πάνθηρ pánthēr , which was used for spotted cats. Felis uncia

2014-618: A role in chemical communication. The snow leopard is a carnivore and actively hunts its prey . Its preferred wild prey species are Himalayan blue sheep ( Pseudois nayaur ), Himalayan tahr ( Hemitragus jemlahicus ), argali ( Ovis ammon ), markhor ( Capra falconeri ) and wild goat ( C. aegagrus ). It also preys on domestic livestock. It prefers prey ranging in weight from 36 to 76 kg (79 to 168 lb), but also hunts smaller mammals such as Himalayan marmot ( Marmota himalayana ), pika and vole species. Its diet depends on prey availability and varies across its range and season. In

2120-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,

2226-497: A shoulder height of 56 cm (22 in), and ranging in head to body size from 75 to 150 cm (30 to 59 in). Its tail is 80 to 105 cm (31 to 41 in) long. Males average 45 to 55 kg (99 to 121 lb), and females 35 to 40 kg (77 to 88 lb). Occasionally, large males reaching 75 kg (165 lb) have been recorded, and small females under 25 kg (55 lb). Its canine teeth are 28.6 mm (1.13 in) long and are more slender than those of

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2332-407: A snow leopard cannot roar , as its 9 mm (0.35 in) short vocal folds provide little resistance to airflow. Its nasal openings are large in relation to the length of its skull and width of its palate ; thanks to their size the volume of air inhaled with each breath is optimised, and the cold dry air becomes warmer. It is not especially adapted to high-altitude hypoxia . The snow leopard

2438-498: A snow leopard was photographed carrying a freshly killed woolly flying squirrel ( Eupetaurus cinereus ) near Gangotri National Park . In Mongolia, domestic sheep comprises less than 20% of its diet, although wild prey has been reduced and interactions with people are common. It is capable of killing most ungulates in its habitat, with the probable exception of the adult male wild yak . It also eats grass and twigs. The snow leopard actively pursues prey down steep mountainsides, using

2544-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of

2650-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in

2756-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as

2862-491: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there

2968-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define

3074-740: A survey of Chinese websites revealed 15 advertisements for 44 snow leopard products; the dealers offered skins, canine teeth, claws and a tongue. In September 2014, nine snow leopard skins were found during a market survey in Afghanistan. Greenhouse gas emissions will likely cause a shift of the treeline in the Himalayas and a shrinking of the alpine zone , which may reduce snow leopard habitat by an estimated 30%. Where snow leopards prey on domestic livestock , they are subject to human–wildlife conflict . The loss of natural prey due to overgrazing by livestock, poaching, and defense of livestock are

3180-528: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including

3286-517: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines

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3392-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species

3498-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,

3604-639: Is solitary and mostly active at dawn till early morning, and again in afternoons and early evenings. They mostly rest near cliffs and ridges that provide vantage points and shade. In Nepal's Shey Phoksundo National Park , the home ranges of five adult radio-collared snow leopards largely overlapped, though they rarely met. Their individual home ranges ranged from 12 to 39 km (4.6 to 15.1 sq mi). Males moved between 0.5 and 5.45 km (0.31 and 3.39 mi) per day, and females between 0.2 and 2.25 km (0.12 and 1.40 mi), measured in straight lines between survey points. Since they often zigzagged in

3710-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that

3816-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if

3922-455: Is a political symbol of the Tatars , Kazakhs , and Bulgars . A mythical winged Aq Bars is depicted on the national coat of arms of Tatarstan , the seal of the city of Samarqand , Uzbekistan and the old coat of arms of Astana . A snow leopard is depicted on the official seal of Almaty and on the former 10,000 Kazakhstani tenge banknote. In Kyrgyzstan, it is used in highly stylized form in

4028-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This

4134-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation

4240-412: Is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example,

4346-958: Is considered good habitat, and 14.4% is protected. In the beginning of the 1990s, the Indian snow leopard population was estimated at 200–600 individuals living across about 25 protected areas. The Snow Leopard Population Assessment in India (SPAI) Programme counted the number of snow leopards between 2019 and 2023 and found their number to be 718, with 477 in Ladakh , 124 in Uttarakhand , 51 in Himachal Pradesh, 36 in Arunachal Pradesh, 21 in Sikkim, and nine in Jammu and Kashmir. In summer,

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4452-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when

4558-884: Is distributed from the west of Lake Baikal through southern Siberia, in the Kunlun Mountains , Altai Mountains, Sayan and Tannu-Ola Mountains , in the Tian Shan , through Tajikistan , Kyrgyzstan , Uzbekistan and Kazakhstan to the Hindu Kush in eastern Afghanistan, Karakoram in northern Pakistan , in the Pamir Mountains , the Tibetan Plateau and in the high elevations of the Himalayas in India , Nepal and Bhutan . In Mongolia, they inhabit

4664-486: Is easily driven away from livestock and readily abandons kills, often without defending itself. Only two attacks on humans have been reported, both near Almaty in Kazakhstan, and neither were fatal. In 1940, a rabid snow leopard attacked two men; and an old, toothless emaciated individual attacked a person passing by. Snow leopards become sexually mature at two to three years, and normally live for 15–18 years in

4770-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as

4876-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,

4982-402: Is known as "Gregg's Paradox": if a single species is the only member of multiple hierarchical levels (for example, being the only species in its genus, which is the only genus in its family), then each level needs a distinct definition to maintain logical structure. Otherwise, the different taxonomic ranks become effectively identical, which creates problems for organizing biological diversity in

5088-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept

5194-478: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with

5300-407: Is particularly associated with island species. Among 25 documented extinct monotypic genera studied, 22 occurred on islands, with flightless animals being particularly vulnerable to human impacts. Just as the term monotypic is used to describe a taxon including only one subdivision, the contained taxon can also be referred to as monotypic within the higher-level taxon, e.g. a genus monotypic within

5406-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,

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5512-488: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. Monotypic genus Monotypic taxa present several important theoretical challenges in biological classification . One key issue

5618-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,

5724-416: Is whitish to grey with black spots on the head and neck, with larger rosettes on the back, flanks and bushy tail. Its muzzle is short, its forehead domed, and its nasal cavities are large. The fur is thick with hairs measuring 5 to 12 cm (2.0 to 4.7 in) in length, and its underbelly is whitish. They are stocky, short-legged, and slightly smaller than other cats of the genus Panthera , reaching

5830-692: The Dolpo region of the Nepal Himalayas in search of the snow leopard. Species A species ( pl. : species) is a population of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use

5936-559: The Himalayas and the Tibetan Plateau to southern Siberia , Mongolia and western China . In the northern part of its range, it also lives at lower elevations. Taxonomically , the snow leopard was long classified in the monotypic genus Uncia . Since phylogenetic studies revealed the relationships among Panthera species, it has since been considered a member of that genus . Two subspecies were described based on morphological differences, but genetic differences between

6042-530: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as

6148-630: The Red Data Book of the Russian Federation as threatened with extinction. Hunting snow leopards is only permitted for the purposes of conservation and monitoring, and to eliminate a threat to the life of humans and livestock. Smuggling of snow leopard body parts is punished with imprisonment and a fine . Hunting snow leopards has been prohibited in Afghanistan since 1986. In China, they have been protected by law since 1989; hunting and trading snow leopards or their body parts constitute

6254-1096: The Snow Leopard Trust , and the Nature and Biodiversity Conservation Union . Also supporting the initiative were the Snow Leopard Network, the World Bank 's Global Tiger Initiative , the United Nations Development Programme , the World Wild Fund for Nature , the United States Agency for International Development , and Global Environment Facility . The Moscow Zoo exhibited the first captive snow leopard in 1872 that had been caught in Turkestan . In Kyrgyzstan, 420 live snow leopards were caught between 1936 and 1988 and exported to zoos around

6360-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is

6466-507: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen

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6572-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept

6678-524: The 170 protected areas in their range countries are smaller than the home range of a single male snow leopard. Snow leopards leave scent marks to indicate their territories and common travel routes. They scrape the ground with the hind feet before depositing urine or feces , but also spray urine onto rocks. Their urine contains many characteristic low molecular weight compounds with diverse functional groups including pentanol , hexanol , heptanol , 3-octanone , nonanal and indole , which possibly play

6784-554: The 1950s. In India, the snow leopard is granted the highest level of protection under the Wildlife Protection Act, 1972 , and hunting is sentenced with imprisonment of 3–7 years. In Nepal, they have been legally protected since 1973, with penalties of 5–15 years in prison and a fine for poaching and trading them. Since 1978, they have been listed in the Soviet Union ’s Red Book and is still inscribed today in

6890-792: The Himalayas, it preys mostly on Himalayan blue sheep, Siberian ibex ( C. sibirica ), white-bellied musk deer ( Moschus leucogaster ) and wild boar ( Sus scrofa ). In the Karakoram , Tian Shan , Altai and Mongolia's Tost Mountains, its main prey consists of Siberian ibex, Thorold's deer ( Cervus albirostris ), Siberian roe deer ( Capreolus pygargus ) and argali. Snow leopard feces collected in northern Pakistan also contained remains of rhesus macaque ( Macaca mulatta ), masked palm civet ( Paguma larvata ), Cape hare ( Lepus capensis ), house mouse ( Mus musculus ), Kashmir field mouse ( Apodemus rusiges ), grey dwarf hamster ( Cricetulus migratorius ) and Turkestan rat ( Rattus pyctoris ). In 2017,

6996-890: The Mongolian and Gobi Altai Mountains and the Khangai Mountains . In Tibet , they occur up to the Altyn-Tagh in the north. They inhabit alpine and subalpine zones at elevations of 3,000 to 4,500 m (9,800 to 14,800 ft), but also lives at lower elevations in the northern part of their range. Potential snow leopard habitat in the Indian Himalayas is estimated at less than 90,000 km (35,000 sq mi) in Jammu and Kashmir , Ladakh , Uttarakhand , Himachal Pradesh , Sikkim and Arunachal Pradesh , of which about 34,000 km (13,000 sq mi)

7102-626: The Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for

7208-618: The State Agency on Environmental Protection and Forestry under the government of Kyrgyzstan. The meeting was held in Bishkek , and all countries agreed that the snow leopard and the high mountain habitat need trans-boundary support to ensure a viable future for snow leopard populations, and to safeguard its fragile environment. The event brought together many partners, including NGOs like the Snow Leopard Conservancy ,

7314-564: The Trans-Ili Alatau mountains since the cameras were installed. In 2013, government leaders and officials from all 12 countries encompassing the snow leopard's range (Afghanistan, Bhutan, China, India, Kazakhstan, Kyrgyzstan, Mongolia, Nepal, Pakistan, Russia, Tajikistan, and Uzbekistan) came together at the Global Snow Leopard Forum (GSLF) initiated by the then-President of Kyrgyzstan Almazbek Atambayev , and

7420-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as

7526-523: The age of about 22 months, but stayed in her vicinity for a month and moved out of his natal range at 23 months of age. The snow leopard has a generation length of eight years. Major threats to the population include poaching and illegal trade of its skins and body parts. Between 1999 and 2002, three live snow leopard cubs and 16 skins were confiscated , 330 traps were destroyed and 110 poachers were arrested in Kyrgyzstan. Undercover operations in

7632-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to

7738-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species

7844-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,

7950-428: The boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in a ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be

8056-423: The cat to balance in rocky terrain. The tail is very thick due to fat storage, and is covered in a thick layer of fur, which allows the cat to use it like a blanket to protect its face when asleep. The snow leopard differs from the other Panthera species by a shorter muzzle, an elevated forehead , a vertical chin and a less developed posterior process of the lower jaw . Despite its partly ossified hyoid bone ,

8162-745: The common application of the term monotypic is frequently misleading, "since each taxon by definition contains exactly one type and is hence "monotypic", regardless of the total number of units", and suggests using "monospecific" for a genus with a single species, and "monotaxonomic" for a taxon containing only one unit. Species in monotypic genera tend to be more threatened with extinction than average species. Studies have found this pattern particularly pronounced in amphibians , where about 6.56% of monotypic genera are critically endangered , compared to birds and mammals where around 4.54% and 4.02% of monotypic genera face critical endangerment respectively. Studies have found that extinction of monotypic genera

8268-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered

8374-416: The concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given a two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part

8480-1039: The country revealed an illegal trade network with links to Russia and China via Kazakhstan. The major skin trade center in the region is the city of Kashgar in Xinjiang . In Tibet and Mongolia, skins are used for traditional dresses, and meat in traditional Tibetan medicine to cure kidney problems; bones are used in traditional Chinese and Mongolian medicine for treating rheumatism , injuries and pain of human bones and tendons . Between 1996 and 2002, 37 skins were found in wildlife markets and tourist shops in Mongolia. Between 2003 and 2016, 710 skins were traded, of which 288 skins were confiscated. In China, an estimated 103 to 236 animals are poached every year, in Mongolia between 34 and 53, in Pakistan between 23 and 53, in India from 21 to 45, and in Tajikistan 20 to 25. In 2016,

8586-450: The cubs can walk at five weeks and are fully weaned by 10 weeks. The cubs leave the den when they are around two to four months of age. Three radio-collared snow leopards in Mongolia's Tost Mountains gave birth between late April and late June. Two female cubs started to part from their mothers at the age of 20 to 21 months, but reunited with them several times for a few days over a period of 4–7 months. One male cub separated from his mother at

8692-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising

8798-427: The information content of biological classifications. As taxonomists Backlund and Bremer explain in their critique, "'Monotypic' taxa do not provide any information about the relationships of the immediately subordinate taxon". When monotypic taxa are sister to a single larger group, they might be merged into that group; however, when they are sister to multiple other groups, they may need to remain separate to maintain

8904-702: The major drivers for the ever decreasing snow leopard population. Livestock also cause habitat degradation, which, alongside the increasing use of forests for fuel, reduces snow leopard habitat. The snow leopard is listed in CITES Appendix I . They have been listed as threatened with extinction in Schedule I of the Convention on the Conservation of Migratory Species of Wild Animals since 1985. Hunting snow leopards has been prohibited in Kyrgyzstan since

9010-665: The modern emblem of the capital Bishkek , and the same art has been integrated into the badge of the Kyrgyzstan Girl Scouts Association . It is also considered to be a sacred creature by the Kyrgyz people . A crowned snow leopard features in the arms of Shushensky District in Russia. It is the state animal of Ladakh and Himachal Pradesh in India. The 1978 book The Snow Leopard is an account by Peter Matthiessen about his two-month journey through

9116-440: The momentum of its initial leap to chase animals for up to 300 m (980 ft). Then it drags the prey to a safe location and consumes all edible parts of the carcass. It can survive on a single Himalayan blue sheep for two weeks before hunting again, and one adult individual apparently needs 20–30 adult blue sheep per year. Snow leopards have been recorded to hunt successfully in pairs, especially mating pairs. The snow leopard

9222-483: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting

9328-400: The other Panthera species. The snow leopard shows several adaptations for living in cold, mountainous environments. Its small rounded ears help to minimize heat loss, and its broad paws effectively distribute the body weight for walking on snow. Fur on the undersides of the paws enhances its grip on steep and unstable surfaces, and helps to minimize heat loss. Its long and flexible tail helps

9434-593: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using

9540-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species

9646-650: The precipitous terrain, they actually moved up to 7 km (4.3 mi) in a single night. Up to 10 individuals inhabit an area of 100 km (39 sq mi); in habitats with sparse prey, an area of 1,000 km (390 sq mi) usually supports only five individuals. A study in the Gobi Desert from 2008 to 2014 revealed that adult males used a mean home range of 144–270 km (56–104 sq mi), while adult females ranged in areas of 83–165 km (32–64 sq mi). Their home ranges overlapped less than 20%. These results indicate that about 40% of

9752-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean

9858-586: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be

9964-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that

10070-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered

10176-637: The snow leopard usually lives above the tree line on alpine meadows and in rocky regions at elevations of 2,700 to 6,000 m (8,900 to 19,700 ft). In winter, they descend to elevations around 1,200 to 2,000 m (3,900 to 6,600 ft). They prefer rocky, broken terrain, and can move in 85 cm (33 in) deep snow, but prefers to use existing trails made by other animals. Snow leopards were recorded by camera traps at 16 locations in northeastern Afghanistan's isolated Wakhan Corridor . The snow leopard's vocalizations include meowing, grunting, prusten and moaning. They can purr when exhaling. It

10282-506: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike

10388-535: The species likely belongs to a different genus. The mitochondrial genomes of the snow leopard, the leopard and the lion ( P. leo ) are more similar to each other than their nuclear genomes , indicating that their ancestors hybridised at some point in their evolution . Fossils of the snow leopard found in the Pabbi Hills of Pakistan were dated to the Early Pleistocene . The snow leopard's fur

10494-427: The tiger probably diverged between 3.7 to 2.7 million years ago . Panthera originates most likely in northern Central Asia. Panthera blytheae excavated in western Tibet's Ngari Prefecture has been initially described the oldest known Panthera species and exhibits skull characteristics similar to the snow leopard, though its taxonomic placement has been disputed by other researchers who suggest that

10600-564: The two have not yet been confirmed. It is therefore regarded as a monotypic species . The species is widely depicted in Kyrgyz culture . The Old French word once , which was intended to be used for the Eurasian lynx ( Lynx lynx ), is where the Latin name uncia and the English word ounce both originate. Once is believed to have originated from a previous form of the word lynx through

10706-540: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of

10812-444: The wild. In captivity they can live for up to 25 years. Oestrus typically lasts five to eight days, and males tend not to seek out another partner after mating, probably because the short mating season does not allow sufficient time. Paired snow leopards mate in the usual felid posture , from 12 to 36 times a day. They are unusual among large cats in that they have a well-defined birth peak. They usually mate in late winter, marked by

10918-565: The world. The Bronx Zoo housed a live snow leopard in 1903; this was the first ever specimen exhibited in a North American zoo. The first captive bred snow leopard cubs were born in the 1990s in the Beijing Zoo . The Snow Leopard Species Survival Plan was initiated in 1984; by 1986, American zoos held 234 individuals. The snow leopard is widely used in heraldry and as an emblem in Central Asia. The Aq Bars ('White Leopard')

11024-777: Was a skin of a female snow leopard collected in the Altai Mountains . He also clarified that several leopard ( P. pardus ) skins were previously misidentified as snow leopard skins. Felis uncioides proposed by Thomas Horsfield in 1855 was a snow leopard skin from Nepal in the collection of the Museum of the East India Company . Uncia uncia was used by Reginald Innes Pocock in 1930 when he reviewed skins and skulls of Panthera species from Asia. He also described morphological differences between snow leopard and leopard skins. Panthera baikalensis-romanii proposed by

11130-562: Was described in 2022 based on fossil material found in France that was dated to the early Middle Pleistocene around 0.57 to 0.53 million years ago . Based on the phylogenetic analysis of the DNA sequence sampled across the living Felidae , the snow leopard forms a sister group with the tiger ( P. tigris ). The genetic divergence time of this group is estimated at 4.62 to 1.82 million years ago . The snow leopard and

11236-578: Was the scientific name used by Johann Christian Daniel von Schreber in 1777 who described a snow leopard based on an earlier description by Georges-Louis Leclerc, Comte de Buffon , assuming that the cat occurred along the Barbary Coast , in Persia , East India and China . The genus name Uncia was proposed by John Edward Gray in 1854 for Asian cats with a long and thick tail. Felis irbis, proposed by Christian Gottfried Ehrenberg in 1830,

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