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50-467: It supports large populations of Thornicroft's giraffe , and herds of elephants and Cape buffaloes often several hundred strong. It is one of the best-known national parks in Africa for walking safaris . Founded as a game reserve in 1938, it became a national park in 1972 and now covers 9,050 km (3,490 sq mi). The Park is unfenced and bordered to the west by a steep escarpment and to

100-416: A difficult time giving birth to young with the same neck proportions as adults. The giraffe's head and neck are held up by large muscles and a nuchal ligament , which are anchored by long thoracic vertebrae spines, giving them a hump. The giraffe's neck vertebrae have ball and socket joints . The point of articulation between the cervical and thoracic vertebrae of giraffes is shifted to lie between

150-466: A disproportionate lengthening of the cervical vertebrae , not from the addition of more vertebrae. Each cervical vertebra is over 28 cm (11 in) long. They comprise 52–54 per cent of the length of the giraffe's vertebral column , compared with the 27–33 percent typical of similar large ungulates, including the giraffe's closest living relative, the okapi . This elongation largely takes place after birth, perhaps because giraffe mothers would have

200-492: A dry savannah emerged across eastern and northern Africa and western India. Some researchers have hypothesised that this new habitat, coupled with a different diet, including acacia species, may have exposed giraffe ancestors to toxins that caused higher mutation rates and a higher rate of evolution. The coat patterns of modern giraffes may also have coincided with these habitat changes. Asian giraffes are hypothesised to have had more okapi-like colourations. The giraffe genome

250-466: A food shortage. Another theory, the sexual selection hypothesis, proposes that long necks evolved as a secondary sexual characteristic , giving males an advantage in "necking" contests (see below) to establish dominance and obtain access to sexually receptive females. In support of this theory, some studies have stated that necks are longer and heavier for males than females of the same age, and that males do not employ other forms of combat. However,

300-464: A giraffe's skull. However, as males age, their skulls become heavier and more club-like, helping them become more dominant in combat. The occipital condyles at the bottom of the skull allow the animal to tip its head over 90 degrees and grab food on the branches directly above them with the tongue. With eyes located on the sides of the head, the giraffe has a broad visual field from its great height. Compared to other ungulates , giraffe vision

350-592: A medium-sized, lightly built body. Giraffokeryx appeared 15–12 mya on the Indian subcontinent and resembled an okapi or a small giraffe, and had a longer neck and similar ossicones . Giraffokeryx may have shared a clade with more massively built giraffids like Sivatherium and Bramatherium . Giraffids like Palaeotragus , Shansitherium and Samotherium appeared 14 mya and lived throughout Africa and Eurasia. These animals had broader skulls with reduced frontal cavities. Paleotragus resembled

400-519: A vital role in conservation of all subspecies of giraffes, due to their popularity with tourists. Giraffes as a species are classified as vulnerable according to the IUCN, but their populations are rapidly declining, with some subspecies being listed as critically endangered Their primary threats are poaching , human population growth , habitat loss , habitat fragmentation , and habitat degradation . Giraffe See taxonomy The giraffe

450-556: Is 22 years for males and 28 years for females. The ecotype was originally named after Harry Scott Thornicroft , a commissioner in what was then North-Eastern Rhodesia and later Northern Rhodesia . Thornicroft's giraffes are tall with very long necks. They have long, dark tongues and skin-colored horns. Giraffes have a typical coat pattern, with regional differences among subspecies. The pattern consists of large, irregular shaped brown to black patches separated by white to yellow bands. Male giraffes' coats darken with age, particularly

500-551: Is a large African hoofed mammal belonging to the genus Giraffa . It is the tallest living terrestrial animal and the largest ruminant on Earth . Traditionally, giraffes have been thought of as one species , Giraffa camelopardalis , with nine subspecies . Most recently, researchers proposed dividing them into four extant species due to new research into their mitochondrial and nuclear DNA , and individual species can be distinguished by their fur coat patterns. Seven other extinct species of Giraffa are known from

550-598: Is around 2.9 billion base pairs in length, compared to the 3.3 billion base pairs of the okapi. Of the proteins in giraffe and okapi genes, 19.4% are identical. The divergence of giraffe and okapi lineages dates to around 11.5 mya. A small group of regulatory genes in the giraffe appears responsible for the animal's height and associated circulatory adaptations. The International Union for Conservation of Nature (IUCN) currently recognises only one species of giraffe with nine subspecies. Carl Linnaeus originally classified living giraffes as one species in 1758. He gave it

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600-404: Is correlated with calf survival. The skin under the blotches may regulate the animal's body temperature, being sites for complex blood vessel systems and large sweat glands . Spotless or solid-color giraffes are very rare, but have been observed. The fur may give the animal chemical defense, as its parasite repellents give it a characteristic scent. At least 11 main aromatic chemicals are in

650-482: Is flexible and hairy to protect against sharp prickles. The upper jaw has a hard palate instead of front teeth. The molars and premolars are wide with low crowns on the surface. The giraffe has an extremely elongated neck, which can be up to 2.4 m (7 ft 10 in) in length. Along the neck is a mane made of short, erect hairs. The neck typically rests at an angle of 50–60 degrees, though juveniles are closer to 70 degrees. The long neck results from

700-477: Is leaves, fruits, and flowers of woody plants, primarily acacia species, which they browse at heights most other ground-based herbivores cannot reach. Lions , leopards , spotted hyenas , and African wild dogs may prey upon giraffes. Giraffes live in herds of related females and their offspring or bachelor herds of unrelated adult males but are gregarious and may gather in large groups. Males establish social hierarchies through "necking", combat bouts where

750-439: Is more binocular and the eyes are larger with a greater retinal surface area. Giraffes may see in colour, and their senses of hearing and smell are sharp. The ears are movable. The nostrils are slit-shaped, possibly to withstand blowing sand. The giraffe's tongue is about 45 cm (18 in) long. It is black, perhaps to protect against sunburn, and can grasp foliage and delicately pick off leaves. The upper lip

800-671: Is one of only two living genera of the family Giraffidae in the order Artiodactyla , the other being the okapi . They are ruminants of the clade Pecora , along with Antilocapridae ( pronghorns ), Cervidae (deer), Bovidae (cattle, antelope, goats and sheep) and Moschidae (musk deer). A 2019 genome study (cladogram below) finds that Giraffidae are a sister taxon to Antilocapridae, with an estimated split of over 20 million years ago. Tragulidae [REDACTED] Antilocapridae [REDACTED] Giraffidae [REDACTED] Cervidae [REDACTED] Bovidae [REDACTED] Moschidae [REDACTED] The family Giraffidae

850-511: Is relatively short. The skin is mostly gray or tan, and can reach a thickness of 20 mm (0.79 in). The 80–100 cm (31–39 in) long tail ends in a long, dark tuft of hair and is used as a defense against insects. The coat has dark blotches or patches, which can be orange, chestnut , brown, or nearly black, surrounded by light hair, usually white or cream coloured. Male giraffes become darker as they grow old. The coat pattern has been claimed to serve as camouflage in

900-450: Is subsumed into G. camelopardalis camelopardalis . G. camelopardalis antiquorum (Kordofan giraffe) G. c. camelopardalis (Nubian giraffe) G. c. peralta (West African giraffe) (no subspecies) G. tippelskirchi tippelskirchi (Masai giraffe sensu stricto ) G. t. thornicrofti (Luangwa or Thornicroft's giraffe) G. giraffa angolensis (Angolan giraffe) G. g. giraffa (South African giraffe) The following table compares

950-537: The Pleistocene . Some biologists suggest the modern giraffes descended from G. jumae ; others find G. gracilis a more likely candidate. G. jumae was larger and more robust, while G. gracilis was smaller and more slender. The changes from extensive forests to more open habitats , which began 8 mya, are believed to be the main driver for the evolution of giraffes. During this time, tropical plants disappeared and were replaced by arid C4 plants , and

1000-568: The Rhodesian giraffe or Luangwa giraffe , is a subspecies of giraffe . It is sometimes considered a species in its own right (as Giraffa thornicrofti ) or a subspecies of the Masai giraffe (as Giraffa tippelskirchi thornicrofti ). It is geographically isolated, occurring only in Zambia ’s South Luangwa Valley . An estimated 550 live in the wild, with no captive populations. Its lifespan

1050-478: The binomial name Cervus camelopardalis . Mathurin Jacques Brisson coined the generic name Giraffa in 1762. During the 1900s, various taxonomies with two or three species were proposed. A 2007 study on the genetics of giraffes using mitochondrial DNA suggested at least six lineages could be recognised as species. A 2011 study using detailed analyses of the morphology of giraffes, and application of

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1100-461: The fossil record. The giraffe's distinguishing characteristics are its extremely long neck and legs, horn-like ossicones , and spotted coat patterns. It is classified under the family Giraffidae , along with its closest extant relative, the okapi . Its scattered range extends from Chad in the north to South Africa in the south and from Niger in the west to Somalia in the east. Giraffes usually inhabit savannahs and woodlands . Their food source

1150-454: The phylogenetic species concept , described eight species of living giraffes. A 2016 study also concluded that living giraffes consist of multiple species. The researchers suggested the existence of four species, which have not exchanged genetic information between each other for 1 to 2 million years. A 2020 study showed that depending on the method chosen, different taxonomic hypotheses recognizing from two to six species can be considered for

1200-553: The 1590s. The modern English form developed around 1600 from the French girafe . "Camelopard" / k ə ˈ m ɛ l ə ˌ p ɑːr d / is an archaic English name for the giraffe; it derives from the Ancient Greek καμηλοπάρδαλις ( kamēlopárdalis ), from κάμηλος ( kámēlos ), " camel ", and πάρδαλις ( párdalis ), " leopard ", referring to its camel-like shape and leopard-like colouration. The giraffe

1250-678: The 1950s, he persuaded the Paramount Chief to set aside a portion of tribal land as a Game Reserve and built the first game viewing camp open to the public in Northern Rhodesia (now Zambia). Guests shot with cameras and not rifles; thus the South Luangwa became the home of the photographic and walking safari. Profits from this remote photographic camp in the bush went back into community. The park spans two eco-regions, both of them woodland savannah, distinguished by

1300-522: The Lower Zambezi and Lilongwe in Malawi. Since 2005, the protected area is considered a Lion Conservation Unit together with North Luangwa National Park . [REDACTED] South Luangwa National Park travel guide from Wikivoyage Rhodesian giraffe G. thornicrofti G. tippelskirchi thornicrofti Thornicroft's giraffe ( Giraffa camelopardalis thornicrofti ), also known as

1350-432: The canopy. However, scientists disagree about just how much time giraffes spend feeding at levels beyond the reach of other browsers, and a 2010 study found that adult giraffes with longer necks actually suffered higher mortality rates under drought conditions than their shorter-necked counterparts. This study suggests that maintaining a longer neck requires more nutrients, which puts longer-necked giraffes at risk during

1400-496: The different hypotheses for giraffe species. The description column shows the traditional nine subspecies in the one-species hypothesis. Also known as Baringo giraffe or Ugandan giraffe Also known as Niger giraffe or Nigerian giraffe Also known as Somali giraffe Also known as Namibian giraffe Also known as Cape giraffe Also known as Kilimanjaro giraffe Also known as Luangwa giraffe or Rhodesian giraffe The first extinct species to be described

1450-469: The dominant tree: Southern Miombo woodlands cover the higher slopes of the valley, while Zambezian and Mopane woodlands cover the bottom of the valley. The Mopane tree tolerates the higher temperatures and lower rainfall found at lower elevations than miombo trees which are found on the higher plateau. Within these woodland savannahs are larger patches of grassland, so that grazers such as zebra and leaf browsers such as giraffe are found in profusion in

1500-812: The east by the Luangwa River. The Luangwa Valley lies at the tail end of the Great African Rift Valley system, which extends 4,000 km (2,500 mi) all the way from the Red Sea down to the Pungwe River mouth in Mozambique. The Muchinga Escarpment in Muchinga and Central Provinces forms the park's western or north-western boundary. It slopes down from there to the river, lying mostly on its western bank. The eastern bank of

1550-472: The elongation of the neck, as it enabled giraffes to reach food that competitors could not. This advantage is real, as giraffes can and do feed up to 4.5 m (15 ft) high, while even quite large competitors, such as kudu, can feed up to only about 2 m (6 ft 7 in) high. There is also research suggesting that browsing competition is intense at lower levels, and giraffes feed more efficiently (gaining more leaf biomass with each mouthful) high in

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1600-420: The existence of four distinct species and seven subspecies. A 2024 study found a higher amount of ancient gene flow than expected between populations. The cladogram below shows the phylogenetic relationship between the four proposed species and seven subspecies based on a 2021 genome analysis. The eight lineages correspond to eight traditional subspecies in the one-species hypothesis. The Rothschild giraffe

1650-538: The first and second thoracic vertebrae (T1 and T2), unlike in most other ruminants, where the articulation is between the seventh cervical vertebra (C7) and T1. This allows C7 to contribute directly to increased neck length and has given rise to the suggestion that T1 is actually C8, and that giraffes have added an extra cervical vertebra. However, this proposition is not generally accepted, as T1 has other morphological features, such as an articulating rib , deemed diagnostic of thoracic vertebrae, and because exceptions to

1700-425: The fur, although indole and 3-methylindole are responsible for most of the smell. Because males have a stronger odour than females, it may also have a sexual function . Both sexes have prominent horn-like structures called ossicones , which can reach 13.5 cm (5.3 in). They are formed from ossified cartilage , covered in skin, and fused to the skull at the parietal bones . Being vascularised ,

1750-482: The genus Giraffa . That study also found that multi-species coalescent methods can lead to taxonomic over-splitting, as those methods delimit geographic structures rather than species. The three-species hypothesis, which recognises G . camelopardalis , G . giraffa , and G . tippelskirchi , is highly supported by phylogenetic analyses and also corroborated by most population genetic and multi-species coalescent analyses. A 2021 whole genome sequencing study suggests

1800-486: The light and shade patterns of savannah woodlands. When standing among trees and bushes, they are hard to see at even a few metres distance. However, adult giraffes move about to gain the best view of an approaching predator, relying on their size and ability to defend themselves rather than on camouflage, which may be more important for calves. Each giraffe has a unique coat pattern. Calves inherit some coat pattern traits from their mothers, and variation in some spot traits

1850-470: The mammalian limit of seven cervical vertebrae are generally characterised by increased neurological anomalies and maladies. There are several hypotheses regarding the evolutionary origin and maintenance of elongation in giraffe necks. Charles Darwin originally suggested the "competing browsers hypothesis", which has been challenged only recently. It suggests that competitive pressure from smaller browsers, like kudu , steenbok and impala , encouraged

1900-689: The neck is used as a weapon. Dominant males gain mating access to females, which bear sole responsibility for rearing the young. The giraffe has intrigued various ancient and modern cultures for its peculiar appearance and has often been featured in paintings, books, and cartoons. It is classified by the International Union for Conservation of Nature (IUCN) as vulnerable to extinction. It has been extirpated from many parts of its former range. Giraffes are still found in many national parks and game reserves , but estimates as of 2016 indicate there are approximately 97,500 members of Giraffa in

1950-452: The okapi and may have been its ancestor. Others find that the okapi lineage diverged earlier, before Giraffokeryx . Samotherium was a particularly important transitional fossil in the giraffe lineage, as the length and structure of its cervical vertebrae were between those of a modern giraffe and an okapi, and its neck posture was likely similar to the former's. Bohlinia , which first appeared in southeastern Europe and lived 9–7 mya,

2000-484: The ossicones may have a role in thermoregulation , and are used in combat between males. Appearance is a reliable guide to the sex or age of a giraffe: the ossicones of females and young are thin and display tufts of hair on top, whereas those of adult males tend to be bald and knobbed on top. A lump, which is more prominent in males, emerges in the middle of the skull. Males develop calcium deposits that form bumps on their skulls as they age. Multiple sinuses lighten

2050-527: The patches. The darkening of the coat has not been studied extensively enough to indicate absolute age; however, it can estimate relative age of male Thornicroft's giraffes. Giraffes occur in arid and dry-savannah zones in sub-Saharan Africa, provided trees are available as a food source. Thornicroft's giraffe is endemic to Zambia. Giraffes are herd animals with a flexible social system. Giraffes are exclusively browsers that primarily feed on leaves and shoots of trees and shrubs. They consume deciduous plants in

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2100-647: The river is in Eastern Province , and as access to the park is only from that side, it is usually thought of as being wholly in Eastern Province. It was initially created as Luangwa Game Reserve in 1904. British conservationist Norman Carr was influential in setting up the South Luangwa National Park. A man ahead of his time, Norman Carr broke the mould of track-and-hunt safari and created conservation based tourism. In

2150-533: The same areas. Patches of flooded grassland habitats (floodplains) are found close to the river, on which hippopotamus graze at night. Their dung released into the river fertilises its waters and sustains the fish population which in turn sustains the crocodiles. The Luangwa valley, continued to the west by the Lunsemfwa River valley, contains some varieties of animals such as Cookson's wildebeest and Crawshay's zebra which are endemic or near-endemic to

2200-478: The species. Thornicroft's giraffes breed throughout the year. They reach sexual maturity at approximately six years, and then produce offspring approximately every 677 days. About half of all calves die before one year of age, mostly due to predation. Giraffes can become pregnant while lactating, an unusual characteristic. Giraffa camelopardalis thornicrofti is endemic to Zambia with a population of less than 550. There are none in captivity. Ecotourism has played

2250-476: The valley. It also represents something of a natural barrier to human migration and transport, no roads cross it and this has helped conserve its wildlife. Although this park is generally well-protected from poaching , its black rhinos were extirpated by 1987, and the elephant population has been under serious pressure at times. The main settlement of the park is actually outside its eastern boundary at Mfuwe , and it has an airport which has flights to Lusaka,

2300-491: The wet season and transition to evergreen and semi-evergreen species in the dry season, choosing flowers, fruits, and pods when they are available. They are true ruminants with forestomach fermentation. Their food intake is approximately 2.1% of the body mass of females and 1.6% for males. They can obtain their water through the foliage they consume, but drink regularly when water is available. Giraffes seek out acacia species when browsing. Their feeding stimulates shoot production of

2350-683: The wild. More than 1,600 were kept in zoos in 2010. The name "giraffe" has its earliest known origins in the Arabic word zarāfah ( زرافة ), ultimately from Persian زُرنَاپَا ( zurnāpā ), a compound of زُرنَا ( zurnā , "flute, zurna") and پَا ( pā , "leg"). In early Modern English the spellings jarraf and ziraph were used, probably directly from the Arabic, and in Middle English jarraf and ziraph , gerfauntz . The Italian form giraffa arose in

2400-520: Was Giraffa sivalensis Falconer and Cautley 1843, a reevaluation of a vertebra that was initially described as a fossil of the living giraffe. While taxonomic opinion may be lacking on some names, the extinct species that have been published include: Fully grown giraffes stand 4.3–5.7 m (14–19 ft) tall, with males taller than females. The average weight is 1,192 kg (2,628 lb) for an adult male and 828 kg (1,825 lb) for an adult female. Despite its long neck and legs, its body

2450-682: Was likely a direct ancestor of the giraffe. Bohlinia closely resembled modern giraffes, having a long neck and legs and similar ossicones and dentition. Bohlinia colonised China and northern India and produced the Giraffa , which, around 7  million years ago , reached Africa. Climate changes led to the extinction of the Asian giraffes, while the African giraffes survived and radiated into new species. Living giraffes appear to have arisen around 1  million years ago in eastern Africa during

2500-631: Was once much more extensive, with over 10 fossil genera described. The elongation of the neck appears to have started early in the giraffe lineage . Comparisons between giraffes and their ancient relatives suggest vertebrae close to the skull lengthened earlier, followed by lengthening of vertebrae further down. One early giraffid ancestor was Canthumeryx , which has been dated variously to have lived 25 to 20 million years ago , 17–15 mya or 18–14.3 mya and whose deposits have been found in Libya. This animal resembled an antelope and had

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