Thomas Beckington (also spelt Beckynton ; c. 1390 – 14 January 1465) was the Bishop of Bath and Wells and King's Secretary in medieval England under Henry VI .
69-525: Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton , five to seven gill slits on the sides of the head , and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimorpha (or Selachii ) and are the sister group to the Batoidea ( rays and kin). Some sources extend the term "shark" as an informal category including extinct members of Chondrichthyes (cartilaginous fish) with
138-425: A heterocercal caudal fin in which the dorsal portion is usually noticeably larger than the ventral portion. This is because the shark's vertebral column extends into that dorsal portion, providing a greater surface area for muscle attachment. This allows more efficient locomotion among these negatively buoyant cartilaginous fish. By contrast, most bony fish possess a homocercal caudal fin. Tiger sharks have
207-414: A cell similar to hair cells present in the vertebrate ear that interact with the surrounding aquatic environment. This helps sharks distinguish between the currents around them, obstacles off on their periphery, and struggling prey out of visual view. The shark can sense frequencies in the range of 25 to 50 Hz . Shark lifespans vary by species. Most live 20 to 30 years. The spiny dogfish has one of
276-416: A constant supply of oxygenated water. A small number of species have lost the ability to pump water through their gills and must swim without rest. These species are obligate ram ventilators and would presumably asphyxiate if unable to move. Obligate ram ventilation is also true of some pelagic bony fish species. The respiratory and circulatory process begins when deoxygenated venous blood travels to
345-497: A few known exceptions, such as the bull shark and the river sharks , which can be found in both seawater and freshwater, and the Ganges shark , which lives only in freshwater. Sharks have a covering of dermal denticles that protects their skin from damage and parasites in addition to improving their fluid dynamics . They have numerous sets of replaceable teeth. Several species are apex predators , which are organisms that are at
414-422: A large upper lobe , which allows for slow cruising and sudden bursts of speed. The tiger shark must be able to twist and turn in the water easily when hunting to support its varied diet, whereas the porbeagle shark , which hunts schooling fish such as mackerel and herring , has a large lower lobe to help it keep pace with its fast-swimming prey. Other tail adaptations help sharks catch prey more directly, such as
483-436: A long surface area, requiring food to circulate inside the short gut until fully digested, when remaining waste products pass into the cloaca . A few sharks appear fluorescent under blue light, such as the swell shark and the chain catshark , where the fluorophore derives from a metabolite of kynurenic acid . Sharks have keen olfactory senses, located in the short duct (which is not fused, unlike bony fish) between
552-618: A monophyletic superorder within Elasmobranchii that shares a common ancestor with the selachians. [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] Thomas Beckington Beckington was born at Beckington in Somerset , and was educated at Winchester and New College, Oxford . Having entered the Church he held many ecclesiastical appointments, and became dean of
621-648: A number of other extinct chondrichthyans with Elasmobrachii sensu stricto /Neoselachii, to the exclusion of more primitive total group elasmobranchs, which is supported by a number of shared morphological characters of the skeleton. The 5th edition of Fishes of the World sets out the following classification of the Elasmobranchs: Recent molecular studies suggest the Batoidea are not derived selachians as previously thought. Instead, skates and rays are
690-569: A nutrient when food is scarce. The oldest unambigous total group elasmobranch, Phoebodus , has its earliest records in the Middle Devonian (late Givetian ), around 383 million years ago. Several important groups of total group elasmobranchs, including Ctenacanthiformes and Hybodontiformes , had already emerged by the latest Devonian ( Famennian ). During the Carboniferous , some ctenacanths would grow to sizes rivalling
759-444: A row behind the head. A modified slit called a spiracle lies just behind the eye, which assists the shark with taking in water during respiration and plays a major role in bottom–dwelling sharks. Spiracles are reduced or missing in active pelagic sharks. While the shark is moving, water passes through the mouth and over the gills in a process known as "ram ventilation". While at rest, most sharks pump water over their gills to ensure
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#1732772353140828-616: A shark-like morphology, such as hybodonts . Shark-like chondrichthyans such as Cladoselache and Doliodus first appeared in the Devonian Period (419–359 million years), though some fossilized chondrichthyan-like scales are as old as the Late Ordovician (458–444 million years ago). The earliest confirmed modern sharks (selachimorphs) are known from the Early Jurassic around 200 million years ago , with
897-403: A similar mechanism for maintaining an elevated body temperature. Larger species, like the whale shark, are able to conserve their body heat through sheer size when they dive to colder depths, and the scalloped hammerhead close its mouth and gills when they dives to depths of around 800 metres, holding its breath till it reach warmer waters again. In contrast to bony fish, with the exception of
966-552: A single type of cone photoreceptor sensitive to green and, seeing only in shades of grey and green, are believed to be effectively colorblind. The study indicates that an object's contrast against the background, rather than colour, may be more important for object detection. Although it is hard to test the hearing of sharks, they may have a sharp sense of hearing and can possibly hear prey from many miles away. The hearing sensitivity for most shark species lies between 20 and 1000 Hz. A small opening on each side of their heads (not
1035-511: A small number of well-developed young as opposed to a large number of poorly developed young. Fecundity in sharks ranges from 2 to over 100 young per reproductive cycle. Sharks mature slowly relative to many other fish. For example, lemon sharks reach sexual maturity at around age 13–15. Elasmobranch See text Elasmobranchs lack swim bladders , and maintain buoyancy with oil that they store in their livers. Some deep sea sharks are targeted by fisheries for this liver oil , including
1104-400: A tissue called tapetum lucidum . This tissue is behind the retina and reflects light back to it, thereby increasing visibility in the dark waters. The effectiveness of the tissue varies, with some sharks having stronger nocturnal adaptations. Many sharks can contract and dilate their pupils , like humans, something no teleost fish can do. Sharks have eyelids, but they do not blink because
1173-437: Is articulated with the upper. Extant elasmobranchs exhibit several archetypal jaw suspensions: amphistyly, orbitostyly, hyostyly, and euhyostyly. In amphistyly, the palatoquadrate has a postorbital articulation with the chondrocranium from which ligaments primarily suspend it anteriorly. The hyoid articulates with the mandibular arch posteriorly, but it appears to provide little support to the upper and lower jaws. In orbitostyly,
1242-647: Is debated. Some believe that electro- and chemoreception are more significant, while others point to the nictating membrane as evidence that sight is important, since presumably the shark would not protect its eyes were they unimportant. The use of sight probably varies with species and water conditions. The shark's field of vision can swap between monocular and stereoscopic at any time. A micro-spectrophotometry study of 17 species of sharks found 10 had only rod photoreceptors and no cone cells in their retinas giving them good night vision while making them colorblind . The remaining seven species had in addition to rods
1311-406: Is depicted twice; one above the other in a two tier arrangement, the bottom effigy depicting his decaying corpse whilst unwrapped from its shroud, and the effigy above depicting him in what is assumed to be his bishop's attire. When his tomb was opened during Victorian times he was found to be buried very simply (as depicted on his tomb) with just one ornament: his bishop's ring . This was removed and
1380-676: Is divided into the superorders Galea (or Galeomorphii ), and Squalea (or Squalomorphii ). The Galeans are the Heterodontiformes , Orectolobiformes , Lamniformes , and Carcharhiniformes . Lamnoids and Carcharhinoids are usually placed in one clade , but recent studies show that Lamnoids and Orectoloboids are a clade. Some scientists now think that Heterodontoids may be Squalean. The Squaleans are divided into Hexanchiformes and Squalomorpha. The former includes cow shark and frilled shark , though some authors propose that both families be moved to separate orders. The Squalomorpha contains
1449-406: Is high, species of sharks from the family Carcharhinidae have smaller olfactory bulbs. Sharks found in deeper waters also have larger olfactory bulbs. Sharks have the ability to determine the direction of a given scent based on the timing of scent detection in each nostril. This is similar to the method mammals use to determine direction of sound. They are more attracted to the chemicals found in
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#17327723531401518-580: Is limited, so sharks employ dynamic lift to maintain depth while swimming. Sand tiger sharks store air in their stomachs, using it as a form of swim bladder. Bottom-dwelling sharks, like the nurse shark , have negative buoyancy, allowing them to rest on the ocean floor. Some sharks, if inverted or stroked on the nose, enter a natural state of tonic immobility . Researchers use this condition to handle sharks safely. Like other fish, sharks extract oxygen from seawater as it passes over their gills . Unlike other fish, shark gill slits are not covered, but lie in
1587-439: Is now in a museum. Beckington played a leading role as architect of the legal aspects of Henry VI's foundation of Eton College in 1440; he is commemorated in the name of the school's central dining hall, 'Bekynton'. In a letter from 1449, Beckington remarked of Bath mentioning that the healing waters of Bath has been turned into abuse by the shamelessness of the inhabitants of the city. Further mentioning: "by ... custom of
1656-433: Is observed in the cookiecutter shark . Tooth shape depends on the shark's diet: those that feed on mollusks and crustaceans have dense and flattened teeth used for crushing, those that feed on fish have needle-like teeth for gripping, and those that feed on larger prey such as mammals have pointed lower teeth for gripping and triangular upper teeth with serrated edges for cutting. The teeth of plankton-feeders such as
1725-410: Is one of the two subclasses of cartilaginous fish in the class Chondrichthyes , the other being Holocephali ( chimaeras ). Members of the elasmobranchii subclass have no swim bladders , five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins , and small placoid scales . The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw
1794-494: Is still evidential in several species termed " dogfish ," or the porbeagle . The etymology of the word shark is uncertain. The most likely etymology states that the original sense of the word was that of "predator, one who preys on others" from the Dutch schurk , meaning 'villain, scoundrel' ( cf. card shark , loan shark , etc.), which was later applied to the fish due to its predatory behaviour. A now disproven theory
1863-774: Is that it derives from the Yucatec Maya word xook ( pronounced [ʃoːk] ), meaning 'shark'. Evidence for this etymology came from the Oxford English Dictionary , which notes that shark first came into use after Sir John Hawkins ' sailors exhibited one in London in 1569 and posted " sharke " to refer to the large sharks of the Caribbean Sea . However, the Middle English Dictionary records an isolated occurrence of
1932-622: The Devonian . Anachronistidae , the oldest probable representatives of Neoselachii, the group containing modern sharks (Selachimorpha) and rays (Batoidea) to the exclusion of most extinct elasmobranch groups, date to the Carboniferous . Selachiimorpha and Batoidea are suggested by some to have diverged during the Triassic . Fossils of the earliest true sharks may have appeared during the Permian , based on remains of " synechodontiforms " found in
2001-733: The Squaliformes and the Hypnosqualea. The Hypnosqualea may be invalid. It includes the Squatiniformes , and the Pristorajea, which may also be invalid, but includes the Pristiophoriformes and the Batoidea . There are more than 500 species of sharks split across thirteen orders , including several orders of sharks that have gone extinct: Shark teeth are embedded in the gums rather than directly affixed to
2070-460: The bull shark , which has developed a way to change its kidney function to excrete large amounts of urea. When a shark dies, the urea is broken down to ammonia by bacteria, causing the dead body to gradually smell strongly of ammonia. Research in 1930 by Homer W. Smith showed that sharks' urine does not contain sufficient sodium to avoid hypernatremia , and it was postulated that there must be an additional mechanism for salt secretion. In 1960 it
2139-508: The clade Selachimorpha in the subclass Elasmobranchii in the class Chondrichthyes . The Elasmobranchii also include rays and skates ; the Chondrichthyes also include Chimaeras . It was thought that the sharks form a polyphyletic group: some sharks are more closely related to rays than they are to some other sharks, but current molecular studies support monophyly of both groups of sharks and batoids. The clade Selachimorpha
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2208-439: The coelacanth , the blood and other tissue of sharks and Chondrichthyes is generally isotonic to their marine environments because of the high concentration of urea (up to 2.5%) and trimethylamine N-oxide (TMAO), allowing them to be in osmotic balance with the seawater. This adaptation prevents most sharks from surviving in freshwater, and they are therefore confined to marine environments. A few exceptions exist, such as
2277-460: The cranium . The jaw's surface (in comparison to the shark's vertebrae and gill arches) needs extra support due to its heavy exposure to physical stress and its need for strength. It has a layer of tiny hexagonal plates called " tesserae ", which are crystal blocks of calcium salts arranged as a mosaic. This gives these areas much of the same strength found in the bony tissue found in other animals. Generally sharks have only one layer of tesserae, but
2346-454: The electric fields they produce. Ocean currents moving in the magnetic field of the Earth also generate electric fields that sharks can use for orientation and possibly navigation. This system is found in most fish, including sharks. It is a tactile sensory system which allows the organism to detect water speed and pressure changes near by. The main component of the system is the neuromast,
2415-414: The great white shark ) are homeothermic and maintain a higher body temperature than the surrounding water. In these sharks, a strip of aerobic red muscle located near the center of the body generates the heat, which the body retains via a countercurrent exchange mechanism by a system of blood vessels called the rete mirabile ("miraculous net"). The common thresher and bigeye thresher sharks have
2484-490: The hybodonts (Order Hybodontiformes), xenacanths (order Xenacanthformes) and Ctenacanthiformes . These are also often referred to as "sharks" in reference to their similar anatomy and ecology to modern sharks. The name Elasmobranchii comes from the Ancient Greek words elasmo- ("plate") and bránchia ("gill"), referring to the broad, flattened gills which are characteristic of these fishes. Elasmobranchii
2553-482: The school , gulper and basking sharks (pictured) . All three of these species have been assessed by the IUCN as vulnerable due to overfishing . From a practical point of view the life-history pattern of elasmobranchs makes this group of animals extremely susceptible to over fishing. It is no coincidence that the commercially exploited marine turtles and baleen whales, which have life-history patterns similar to
2622-615: The Arches in 1423; and Archdeacon of Buckingham in 1424. After that he devoted his time to secular affairs and was sent on an embassy to Calais in 1439 and to John IV, Count of Armagnac in 1442. At this time Beckington was acting as secretary to Henry VI, and soon after his return in 1443 he was appointed Lord Privy Seal , an office he held until 1444. He was consecrated Bishop of Bath and Wells on 13 October 1443. The bishop erected many buildings in Wells and elsewhere, probably altering
2691-1062: The Early Permian of Russia, but if remains of "synechodontiformes" from the Permian and Triassic are true sharks, they only had low diversity. Modern shark orders first appeared during the Early Jurassic, and during the Jurassic true sharks underwent great diversification. Selachimorphs largely replaced the hybodonts , which had previously been a dominant group of shark-like fish during the Triassic and Early Jurassic. Batoidea [REDACTED] Heterodontiformes [REDACTED] Orectolobiformes [REDACTED] Carcharhiniformes [REDACTED] Lamniformes [REDACTED] Hexanchiformes [REDACTED] Squatiniformes [REDACTED] Pristiophoriformes [REDACTED] Squaliformes [REDACTED] Sharks belong to
2760-436: The Triassic and Early Jurassic . Hybodonts were extensively present in both marine and freshwater environments. While Neoselachii/Elasmobranchi sensu stricto (the group of modern sharks and rays) had already appeared by the Triassic, they only had low diversity during this period would and only begin to extensively diversify from the Early Jurassic onwards, when modern orders of sharks and rays appeared. This co-incided with
2829-414: The anterior and posterior nasal openings, with some species able to detect as little as one part per million of blood in seawater. The size of the olfactory bulb varies across different shark species, with size dependent on how much a given species relies on smell or vision to find their prey. In environments with low visibility, shark species generally have larger olfactory bulbs. In reefs, where visibility
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2898-565: The basking shark are small and non-functional. Shark skeletons are very different from those of bony fish and terrestrial vertebrates . Sharks and other cartilaginous fish ( skates and rays ) have skeletons made of cartilage and connective tissue . Cartilage is flexible and durable, yet is about half the normal density of bone. This reduces the skeleton's weight, saving energy. Because sharks do not have rib cages, they can easily be crushed under their own weight on land. The jaws of sharks, like those of rays and skates, are not attached to
2967-415: The body then flows through the posterior cardinal veins and enters the posterior cardinal sinuses . From there venous blood re-enters the heart ventricle and the cycle repeats. Most sharks are "cold-blooded" or, more precisely, poikilothermic , meaning that their internal body temperature matches that of their ambient environment. Members of the family Lamnidae (such as the shortfin mako shark and
3036-640: The city, [the people] shamelessly strip them of their said garments and reveal to them to the gaze of bystanders." Bekynton's architecture is marked with a symbol depicting a barrel and a flame. This is a pun on his name, being "beacon tun". One of the easiest to be viewed is in Penniless Porch Wells. Beckington's own journal was published in the Proceedings of the Privy Council , vol. v., edited by Nicholas Harris Nicolas (1835); and
3105-531: The clade is unclear with respect to fossil chondrichthyans. Some authors consider it as equivalent to Neoselachii (the crown group clade including modern sharks, rays, and all other descendants of their last common ancestor ). Other authors use the name Elasmobranchii for a broader branch-based group of all chondrichthyans more closely related to modern sharks and rays than to Holocephali (the clade containing chimaeras and their extinct relatives). Important extinct groups of elasmobranchs sensu lato include
3174-648: The cranium. Instead, the hyomandibular cartilages provide the only means of jaw support, while the ceratohyal and basihyal elements articulate with the lower jaw, but are disconnected from the rest of the hyoid. The eyes have a tapetum lucidum . The inner margin of each pelvic fin in the male fish is grooved to constitute a clasper for the transmission of sperm . These fish are widely distributed in tropical and temperate waters. Many fish maintain buoyancy with swim bladders . However elasmobranchs lack swim bladders, and maintain buoyancy instead with large livers that are full of oil. This stored oil may also function as
3243-619: The decline of the hybodonts, which had become minor components of marine environments by the Late Jurassic, but would remain common in freshwater environments into the Cretaceous . The youngest remains of hybodonts date to the very end of the Cretaceous. Elasmobranchii was first coined in 1838 by Charles Lucien Bonaparte . Bonaparte's original definition of Elasmobranchii was effectively identical to modern Chondrichthyes , and
3312-1162: The horny keratin in hair and feathers. Most sharks have eight fins. Sharks can only drift away from objects directly in front of them because their fins do not allow them to move in the tail-first direction. Unlike bony fish, sharks have a complex dermal corset made of flexible collagenous fibers and arranged as a helical network surrounding their body. This works as an outer skeleton, providing attachment for their swimming muscles and thus saving energy. Their dermal teeth give them hydrodynamic advantages as they reduce turbulence when swimming. Some species of shark have pigmented denticles that form complex patterns like spots (e.g. Zebra shark ) and stripes (e.g. Tiger shark ). These markings are important for camouflage and help sharks blend in with their environment, as well as making them difficult for prey to detect. For some species, dermal patterning returns to healed denticles even after they have been removed by injury. Tails provide thrust, making speed and acceleration dependent on tail shape. Caudal fin shapes vary considerably between shark species, due to their evolution in separate environments. Sharks possess
3381-417: The inner ear has been lost. The ampullae of Lorenzini are the electroreceptor organs. They number in the hundreds to thousands. Sharks use the ampullae of Lorenzini to detect the electromagnetic fields that all living things produce. This helps sharks (particularly the hammerhead shark ) find prey. The shark has the greatest electrical sensitivity of any animal. Sharks find prey hidden in sand by detecting
3450-402: The intestines of many species, and as a result often linger near or in sewage outfalls. Some species, such as nurse sharks , have external barbels that greatly increase their ability to sense prey. Shark eyes are similar to the eyes of other vertebrates , including similar lenses , corneas and retinas , though their eyesight is well adapted to the marine environment with the help of
3519-451: The jaw, and are constantly replaced throughout life. Multiple rows of replacement teeth grow in a groove on the inside of the jaw and steadily move forward in comparison to a conveyor belt ; some sharks lose 30,000 or more teeth in their lifetime. The rate of tooth replacement varies from once every 8 to 10 days to several months. In most species, teeth are replaced one at a time as opposed to the simultaneous replacement of an entire row, which
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#17327723531403588-451: The jaws of large specimens, such as the bull shark, tiger shark, and the great white shark, have two to three layers or more, depending on body size. The jaws of a large great white shark may have up to five layers. In the rostrum (snout), the cartilage can be spongy and flexible to absorb the power of impacts. Fin skeletons are elongated and supported with soft and unsegmented rays named ceratotrichia, filaments of elastic protein resembling
3657-544: The last common ancestor of modern sharks and rays, a grouping which had previously been named Neoselachii by Compagno (1977). Other recent authors have used Elasmobranchii in a broad sense to include all chondrichthyans more closely related to modern sharks and rays than to chimaeras. The total group of Elasmobranchii includes the Cohort Euselachii Hay, 1902, which groups the Hybodontiformes and
3726-539: The longest lifespans at more than 100 years. Whale sharks ( Rhincodon typus ) may also live over 100 years. Earlier estimates suggested the Greenland shark ( Somniosus microcephalus ) could reach about 200 years, but a recent study found that a 5.02-metre-long (16.5 ft) specimen was 392 ± 120 years old (i.e., at least 272 years old), making it the longest-lived vertebrate known. Unlike most bony fish , sharks are K-selected reproducers, meaning that they produce
3795-472: The modern great white shark with bodies in the region of 7 metres (23 ft) in length. During the Carboniferous and Permian , the xenacanths were abundant in both freshwater and marine environments, and would continue to exist into the Triassic with reduced diversity. The hybodonts had achieved a high diversity by the Permian, and would end up becoming the dominant group of elasmobranchs during
3864-551: The oldest known member being Agaleus , though records of true sharks may extend back as far as the Permian . Sharks range in size from the small dwarf lanternshark ( Etmopterus perryi ), a deep sea species that is only 17 centimetres (6.7 in) in length, to the whale shark ( Rhincodon typus ), the largest fish in the world, which reaches approximately 12 metres (40 ft) in length. They are found in all seas and are common to depths up to 2,000 metres (6,600 ft). They generally do not live in freshwater, although there are
3933-412: The orbital process hinges with the orbital wall and the hyoid provides the majority of suspensory support. In contrast, hyostyly involves an ethmoid articulation between the upper jaw and the cranium, while the hyoid most likely provides vastly more jaw support compared to the anterior ligaments. Finally, in euhyostyly, also known as true hyostyly, the mandibular cartilages lack a ligamentous connection to
4002-504: The rectory at Sutton Courtenay in Berkshire (now Oxfordshire ), an early preferment. The most important results of Beckington's missions to France were one Latin journal, written by himself, referring to the embassy to Calais; and another, written by one of his attendants, relating to the journey to Armagnac . He died at Wells on 14 January 1465. Beckington is buried at Wells Cathedral and has an unusual monument there: his effigy
4071-403: The shark's two-chambered heart . Here, the shark pumps blood to its gills via the ventral aorta where it branches into afferent branchial arteries . Gas exchange takes place in the gills and the reoxygenated blood flows into the efferent branchial arteries, which come together to form the dorsal aorta . The blood flows from the dorsal aorta throughout the body. The deoxygenated blood from
4140-453: The sharks, are also in trouble. Elasmobranchii ( / ɪ ˌ l æ z m ə ˈ b r æ ŋ k i aɪ / ) is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks (superorder Selachii), rays , skates , and sawfish (superorder Batoidea ). Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on
4209-447: The skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades . The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil. The definition of
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#17327723531404278-413: The spiracle) leads directly into the inner ear through a thin channel. The lateral line shows a similar arrangement, and is open to the environment via a series of openings called lateral line pores . This is a reminder of the common origin of these two vibration- and sound-detecting organs that are grouped together as the acoustico-lateralis system. In bony fish and tetrapods the external opening into
4347-413: The stomach, and instead the shark either vomits or turns its stomachs inside out and ejects unwanted items from its mouth. One of the biggest differences between the digestive systems of sharks and mammals is that sharks have much shorter intestines. This short length is achieved by the spiral valve with multiple turns within a single short section instead of a long tube-like intestine. The valve provides
4416-423: The surrounding water cleans their eyes. To protect their eyes some species have nictitating membranes . This membrane covers the eyes while hunting and when the shark is being attacked. However, some species, including the great white shark ( Carcharodon carcharias ), do not have this membrane, but instead roll their eyes backwards to protect them when striking prey. The importance of sight in shark hunting behavior
4485-401: The thresher shark's usage of its powerful, elongated upper lobe to stun fish and squid. Unlike bony fish, sharks do not have gas-filled swim bladders for buoyancy. Instead, sharks rely on a large liver filled with oil that contains squalene , and their cartilage, which is about half the normal density of bone. Their liver constitutes up to 30% of their total body mass. The liver's effectiveness
4554-453: The top of their food chain . Select examples include the bull shark , tiger shark , great white shark , mako sharks , thresher sharks , and hammerhead sharks . Sharks are caught by humans for shark meat or shark fin soup . Many shark populations are threatened by human activities. Since 1970, shark populations have been reduced by 71%, mostly from overfishing . Until the 16th century, sharks were known to mariners as "sea dogs". This
4623-545: The word shark (referring to a sea fish) in a letter written by Thomas Beckington in 1442, which rules out a New World etymology. The oldest total-group chondrichthyans, known as acanthodians or "spiny sharks", appeared during the Early Silurian , around 439 million years ago. The oldest confirmed members of Elasmobranchii sensu lato (the group containing all cartilaginous fish more closely related to modern sharks and rays than to chimaeras ) appeared during
4692-432: Was based around gill architecture shared by all 3 living major cartilaginous fish groups. During the 20th century it became standard to exclude chimaeras from Elasmobranchii; along with including many fossil chondrichthyans within the group. The definition of Elasmobranchii has since been subject to much confusion with regard to fossil chondrichthyans. Maisey (2012) suggested that Elasmobranchii should exclusively be used for
4761-574: Was discovered at the Mount Desert Island Biological Laboratory in Salsbury Cove, Maine that sharks have a type of salt gland located at the end of the intestine, known as the "rectal gland", whose function is the secretion of chlorides. Digestion can take a long time. The food moves from the mouth to a J-shaped stomach, where it is stored and initial digestion occurs. Unwanted items may never get past
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