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42-632: 16, see text Sunbirds and spiderhunters make up the family Nectariniidae of passerine birds. They are small, slender passerines from the Old World , usually with downward-curved bills. Many are brightly coloured, often with iridescent feathers, particularly in the males. Many species also have especially long tail feathers. Their range extends through most of Africa to the Middle East, South Asia, South-east Asia and southern China, to Indonesia, New Guinea and northern Australia. Species diversity

84-469: A zeitgeber . Animals active during twilight are crepuscular , those active during the night are nocturnal and animals active at sporadic times during both night and day are cathemeral . Plants that open their flowers during the daytime are described as diurnal, while those that bloom during nighttime are nocturnal. The timing of flower opening is often related to the time at which preferred pollinators are foraging. For example, sunflowers open during

126-435: A balance between them if they are to survive and thrive. Ambient temperature has been shown to affect and even convert nocturnal animals to diurnality as it is a way for them to conserve metabolic energy. Nocturnal animals are often energetically challenged due to being most active in the nighttime when ambient temperatures are lower than through the day, and so they lose a lot of energy in the form of body heat. According to

168-495: A challenge, apart from the obvious increased activity levels during the day time light. Initially, most animals were diurnal, but adaptations that allowed some animals to become nocturnal is what helped contribute to the success of many, especially mammals. This evolutionary movement to nocturnality allowed them to better avoid predators and gain resources with less competition from other animals. This did come with some adaptations that mammals live with today. Vision has been one of

210-700: A gap in their distribution across inland Syria and Iraq, and resuming in Iran , from where the group occurs continuously as far as southern China and Indonesia . In Australasia the family occurs in New Guinea , north eastern Australia and the Solomon Islands . They are generally not found on oceanic islands, with the exception of the Seychelles . The greatest variety of species is found in Africa, where

252-556: A hummingbird, but they usually perch to feed. The family ranges in size from the 5-gram black-bellied sunbird to the spectacled spiderhunter , at about 45 grams. Like the hummingbirds, sunbirds are strongly sexually dimorphic , with the males usually brilliantly plumaged in iridescent colours. In addition to this the tails of many species are longer in the males, and overall the males are larger. Sunbirds have long thin down-curved bills and brush-tipped tubular tongues, both adaptations to their nectar feeding. The spiderhunters , of

294-554: A lack of widespread consensus within the scientific community for extended periods. The continual publication of new data and diverse opinions plays a crucial role in facilitating adjustments and ultimately reaching a consensus over time. The naming of families is codified by various international bodies using the following suffixes: The taxonomic term familia was first used by French botanist Pierre Magnol in his Prodromus historiae generalis plantarum, in quo familiae plantarum per tabulas disponuntur (1689) where he called

336-499: A period of sleeping or other inactivity at night . The common adjective used for daytime activity is " diurnal ". The timing of activity by an animal depends on a variety of environmental factors such as the temperature, the ability to gather food by sight, the risk of predation, and the time of year. Diurnality is a cycle of activity within a 24-hour period; cyclic activities called circadian rhythms are endogenous cycles not dependent on external cues or environmental factors except for

378-407: A state of low activity and responsiveness. The moulting regimes of sunbirds are complex, being different in different species. Many species have no eclipse plumage, but do have juvenile plumage. Some species do show duller plumage in the off-season. In the dry months of June−August, male copper sunbirds and variable sunbirds lose much of their metallic sheen. In some instances different populations of

420-442: A wide range of habitats from sea level to 4900 m. Sunbird are active diurnal birds that generally occur in pairs or occasionally in small family groups. A few species occasionally gather in larger groups, and sunbird will join with other birds to mob potential predators, although sunbirds will also aggressively target other species, even if they are not predators, when defending their territories . Sunbirds that breed outside of

462-455: Is also limited, meaning the risks of predation are less than the risk of freezing or starving to death. Many plants are diurnal or nocturnal, depending on the time period when the most effective pollinators, i.e., insects, visit the plant. Most angiosperm plants are visited by various insects, so the flower adapts its phenology to the most effective pollinators. Thus, the effectiveness of relative diurnal or nocturnal species of insects affects

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504-499: Is commonly referred to as the "walnut family". The delineation of what constitutes a family— or whether a described family should be acknowledged— is established and decided upon by active taxonomists . There are not strict regulations for outlining or acknowledging a family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to

546-535: Is considered an unpleasant song and are tricky to keep alive. Sunbirds are considered attractive birds and readily enter gardens where flowering plants are planted to attract them. There are a few negative interactions, for example the scarlet-chested sunbird is considered a pest in cocoa plantations as it spreads parasitic mistletoes . The family contains 151 species divided into 16 genera: For more detail, see list of sunbird species . Family (biology) Family ( Latin : familia , pl. : familiae )

588-407: Is highest in equatorial regions. There are 151 species in 16 genera . Their family name is from most sunbirds feeding largely on nectar , but they will also catch insects and spiders , especially when feeding their young. Flowers that prevent access to their nectar because of their shape (for example, very long and narrow flowers) are simply punctured at the base near the nectaries, from which

630-485: Is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy . It is classified between order and genus . A family may be divided into subfamilies , which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae , but that family

672-505: Is one of the strongest influences of the suprachiasmatic nucleus (SCN) which is part of the hypothalamus in the brain that controls the circadian rhythm in most animals. This is what determines whether an animal is diurnal or not. The SCN uses visual information like light to start a cascade of hormones that are released and work on many physiological and behavioural functions. Light can produce powerful masking effects on an animal's circadian rhythm, meaning that it can "mask" or influence

714-402: Is thought to have contributed to plant speciation , including the exceptionally high floral diversity in southern Africa. Overall the family has fared better than many others, with only seven species considered to be threatened with extinction . Most species are fairly resistant to changes in habitat, and while attractive the family is not sought after by the cagebird trade, as they have what

756-530: The Genera Plantarum of George Bentham and Joseph Dalton Hooker this word ordo was used for what now is given the rank of family. Families serve as valuable units for evolutionary, paleontological, and genetic studies due to their relatively greater stability compared to lower taxonomic levels like genera and species. Diurnality Diurnality is a form of plant and animal behavior characterized by activity during daytime , with

798-533: The Gran Chaco in South America showed that increased amounts of moonlight at night increased their activity levels through the night which led to a decrease of daytime activity. Meaning that for this species, ambient moonlight is negatively correlated with diurnal activity. This is also connected with the foraging behaviours of the monkeys, as when there were nights of little to no moonlight, it affected

840-505: The Americas. A key difference is that sunbirds cannot hover, so sunbird-pollinated flowers and inflorescences are typically sturdier than hummingbird-pollinated flowers, with an appropriate landing spot from which the bird can feed. Sunbirds are critical pollinators for many iconic African plants, including proteas , aloes , Erica , Erythrina coral trees, and bird-of-paradise flowers. Specialization on sunbirds vs other pollinators

882-439: The birds sip the nectar. Fruit is also part of the diet of some species. Their flight is fast and direct, thanks to their short wings. The sunbirds have counterparts in two very distantly related groups: the hummingbirds of the Americas and the honeyeaters of Australia. The resemblances are due to convergent evolution brought about by a similar nectar-feeding lifestyle. Some sunbird species can take nectar by hovering like

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924-454: The circadian thermos-energetics (CTE) hypothesis, animals that are expending more energy than they are taking in (through food and sleep) will be more active in the light cycle, meaning they will be more active in the day. This has been shown in studies done on small nocturnal mice in a laboratory setting. When they were placed under a combination of enough cold and hunger stress, they converted to diurnality through temporal niche switching, which

966-463: The day to attract bees, whereas the night-blooming cereus opens at night to attract large sphinx moths . Many types of animals are classified as being diurnal, meaning they are active during the day time and inactive or have periods of rest during the night time. Commonly classified diurnal animals include mammals , birds , and reptiles . Most primates are diurnal, including humans . Scientifically classifying diurnality within animals can be

1008-400: The day, and therefore conserve more energy, especially when colder seasonal temperatures hit. Light is one of the most defining environmental factors that determines an animal's activity pattern. Photoperiod or a light dark cycle is determined by the geographical location, with day time being associated with much ambient light, and night time being associated with little ambient light. Light

1050-420: The diurnal or nocturnal nature of the plants they pollinate, causing in some instances an adjustment of the opening and closing cycles of the plants. For example, the baobab is pollinated by fruit bats and starts blooming in late afternoon; the flowers are dead within twenty-four hours. Services that alternate between high and low utilization in a daily cycle are described as being diurnal. Many websites have

1092-403: The equatorial regions are mostly seasonal breeders, with the majority of them breeding in the wet season . This timing reflects the increased availability of insect prey for the growing young. Where species, like the buff-throated sunbird , breed in the dry season, it is thought to be associated with the flowering of favoured food plants. Species of sunbird in the equatorial areas breed throughout

1134-439: The evolution of diurnality. With about 20 transitions counted for the gecko lineages, it shows the significance of diurnality. Strong environmental influences like climate change, predation risk, and competition for resources are all contributing factors. Using the example of geckos, it is thought that species like Mediodactylus amictopholis that live at higher altitudes have switched to diurnality to help gain more heat through

1176-540: The family as a rank intermediate between order and genus was introduced by Pierre André Latreille in his Précis des caractères génériques des insectes, disposés dans un ordre naturel (1796). He used families (some of them were not named) in some but not in all his orders of "insects" (which then included all arthropods ). In nineteenth-century works such as the Prodromus of Augustin Pyramus de Candolle and

1218-442: The genus Arachnothera , are distinct in appearance from the other members of the family. They are typically larger than the other sunbirds, with drab brown plumage that is the same for both sexes, and long, down-curved beaks. In metabolic behaviour similar to that of Andes hummingbirds, species of sunbirds that live at high altitudes or latitudes will enter torpor while roosting at night, lowering their body temperature and entering

1260-614: The group probably arose. Most species are sedentary or short-distance seasonal migrants . Sunbirds occur over the entire family's range, whereas the spiderhunters are restricted to Asia . The sunbirds and spiderhunters occupy a wide range of habitats, with a majority of species being found in primary rainforest, but other habitats used by the family including disturbed secondary forest, open woodland, open scrub and savannah, coastal scrub and alpine forest. Some species have readily adapted to human modified landscapes such as plantations, gardens and agricultural land. Many species are able to occupy

1302-416: The internal clock, changing the activity patterns of an animal, either temporarily or over the long term if exposed to enough light over a long period of time. Masking can be referred to either as positive masking or negative masking, with it either increasing an diurnal animals activity or decreasing a nocturnal animal's activity, respectively. This can be depicted when exposing different types of rodents to

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1344-604: The monkey's ability to forage efficiently, so they were forced to be more active in the day to find food. Diurnality has shown to be an evolutionary trait in many animal species, with diurnality mostly reappearing in many lineages. Other environmental factors like ambient temperature, food availability, and predation risk can all influence whether an animal will evolve to be diurnal, or if their effects are strong enough, then mask over their circadian rhythm, changing their activity patterns to becoming diurnal. All three factors often involve one another, and animals need to be able to find

1386-572: The most common transitions. Still today, diurnality seems to be reappearing in many lineages of other animals, including small rodent mammals like the Nile grass rat and golden mantle squirrel and reptiles. More specifically, geckos, which were thought to be naturally nocturnal have shown many transitions to diurnality, with about 430 species of geckos now showing diurnal activity. With so many diurnal species recorded, comparative analysis studies using newer lineages of gecko species have been done to study

1428-767: The most greatly affected senses from switching back and forth from diurnality to nocturnality, and this can be seen using biological and physiological analysis of rod nuclei from primate eyes. This includes losing two of four cone opsins that assists in colour vision , making many mammals dichromats . When early primates converted back to diurnality, better vision that included trichromatic colour vision became very advantageous, making diurnality and colour vision adaptive traits of simiiformes , which includes humans. Studies using chromatin distribution analysis of rod nuclei from different simian eyes found that transitions between diurnality and nocturnality occurred several times within primate lineages, with switching to diurnality being

1470-512: The nestlings. In the spiderhunters both sexes help to incubate the eggs. The nests of sunbirds and spiderhunters are often targeted by brood parasites such as cuckoos and honeyguides . As nectar is a primary food source for sunbirds, they are important pollinators in African ecosystems. Sunbird-pollinated flowers are typically long, tubular, and red-to-orange in colour, showing convergent evolution with many hummingbird -pollinated flowers in

1512-439: The same photoperiods. When a diurnal Nile grass rat and nocturnal mouse are exposed to the same photoperiod and light intensity, increased activity occurred within the grass rat (positive masking), and decreased activity within the mouse (negative masking). Even small amounts of environmental light change have shown to have an effect on the activity of mammals. An observational study done on the activity of nocturnal owl monkeys in

1554-648: The same species can display variation in different molting regimes. Sunbirds are a tropical Old World family, with representatives in Africa , Asia and Australasia . In Africa they are found mostly in sub-Saharan Africa and Madagascar but are also distributed in Egypt . In Asia the group occurs along the coasts of the Red Sea as far north as Israel , and along the Mediterranean as far north as Beirut , with

1596-575: The seventy-six groups of plants he recognised in his tables families ( familiae ). The concept of rank at that time was not yet settled, and in the preface to the Prodromus Magnol spoke of uniting his families into larger genera , which is far from how the term is used today. In his work Philosophia Botanica published in 1751, Carl Linnaeus employed the term familia to categorize significant plant groups such as trees , herbs , ferns , palms , and so on. Notably, he restricted

1638-400: The underside of large leaves; that of the yellow-eared spiderhunter is similarly attached but is a long tube. The nests of spiderhunters are inconspicuous, in contrast to those of the other sunbirds which are more visible. In most species the female alone constructs the nest. Up to four eggs are laid. The female builds the nest and incubates the eggs alone, although the male assists in rearing

1680-549: The use of this term solely within the book's morphological section, where he delved into discussions regarding the vegetative and generative aspects of plants. Subsequently, in French botanical publications, from Michel Adanson 's Familles naturelles des plantes (1763) and until the end of the 19th century, the word famille was used as a French equivalent of the Latin ordo (or ordo naturalis ). In zoology ,

1722-423: The year. They are generally monogamous and often territorial, although a few species of sunbirds have lekking behaviour. The nests of sunbirds are generally purse-shaped, enclosed, suspended from thin branches with generous use of spiderweb. The nests of the spiderhunters are different, both from the sunbirds and in some cases from each other. Some, like the little spiderhunter , are small woven cups attached to

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1764-407: Was expected. Another similar study that involved energetically challenging small mammals showed that diurnality is most beneficial when the animal has a sheltered location to rest in, reducing heat loss. Both studies concluded that nocturnal mammals do change their activity patterns to be more diurnal when energetically stressed (due to heat loss and limited food availability), but only when predation

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