95-482: Pomodoro (Italian for " tomato ") may refer to: Arnaldo Pomodoro (born 1926), Italian sculptor Giò Pomodoro (1930–2002), Arnaldo's brother, another sculptor Pappa al pomodoro , an Italian soup dish Pasta al pomodoro , an Italian pasta dish Pasta Pomodoro (restaurant) , American restaurant chain Passata di pomodoro, tomato purée The Pomodoro Technique ,
190-440: A proper noun such as the name of a person or place. Similarly, both parts are italicized in normal text (or underlined in handwriting). Thus the binomial name of the annual phlox (named after botanist Thomas Drummond ) is now written as Phlox drummondii . Often, after a species name is introduced in a text, the generic name is abbreviated to the first letter in subsequent mentions (e.g., P. drummondii ). In scientific works,
285-547: A case of convergent evolution ) in the cultivated tomato. The analysis predicts that var. cerasiforme appeared around 78,000 years ago, while the cultivated tomato originated around 7,000 years ago (5,000 BCE), with substantial uncertainty, making it unclear how humans may have been involved in the process. The Spanish first introduced tomatoes to Europe, where they became used in Spanish food. Elsewhere in Europe, its first use
380-602: A column surrounding the pistil 's style. The anthers bend into a cone-like structure, surrounding the stigma. The flowers are 1–2 cm (0.4–0.8 in) across, yellow, with five pointed lobes on the corolla ; they are borne in a cyme of three to twelve together. The fruit develops from the ovary of the plant after fertilization, its flesh comprising the pericarp walls. The fruit contains locules , hollow spaces full of seeds. These vary among cultivated varieties. Some smaller varieties have two locules; globe-shaped varieties typically have three to five; beefsteak tomatoes have
475-417: A cylindric spike and a terete scape "), which we know today as Plantago media . Such "polynomial names" may sometimes look like binomials, but are significantly different. For example, Gerard's herbal (as amended by Johnson) describes various kinds of spiderwort: "The first is called Phalangium ramosum , Branched Spiderwort; the second, Phalangium non ramosum , Unbranched Spiderwort. The other ...
570-456: A great number of small locules; and plum tomatoes have very few, very small locules. For propagation, the seeds need to come from a mature fruit, and must be lightly fermented to remove the gelatinous outer coating and then dried before use. The tomato has a mutualistic relationship with arbuscular mycorrhizal fungi such as Rhizophagus irregularis . Scientists use the tomato as a model species for investigating such symbioses. Like
665-402: A lighter green, and reducing sugar in the resulting ripe fruit by 10–15%. Perhaps more importantly, the fruit chloroplasts are remodelled during ripening into chlorophyll-free chromoplasts that synthesize and accumulate the carotenoids lycopene , β-carotene , and other metabolites that are sensory and nutritional assets of the ripe fruit. The potent chloroplasts in the dark-green shoulders of
760-978: A name for the fruit. The usual pronunciations of tomato are / t ə ˈ m eɪ t oʊ / (in North American English ) and / t ə ˈ m ɑː t oʊ / (in British English ). The British pronunciation was like the American until the Great Vowel Shift . The word's dual pronunciations were immortalized in Ira and George Gershwin 's 1937 song " Let's Call the Whole Thing Off " ("You like / p ə ˈ t eɪ t oʊ / and I like / p ə ˈ t ɑː t oʊ / / You like / t ə ˈ m eɪ t oʊ / and I like / t ə ˈ m ɑː t oʊ / "). The likely wild ancestor of
855-401: A nomenclature code, it is possible for homonyms (two or more species sharing the same genus name) to happen, and even the same binomial if they occur in different kingdoms. At least 1,258 instances of genus name duplication occur (mainly between zoology and botany). Nomenclature (including binomial nomenclature) is not the same as classification, although the two are related. Classification is
950-481: A number of sources, of which Latin is only one. These include: The first part of the name, which identifies the genus, must be a word that can be treated as a Latin singular noun in the nominative case . It must be unique within the purview of each nomenclatural code , but can be repeated between them. Thus Huia recurvata is an extinct species of plant, found as fossils in Yunnan , China, whereas Huia masonii
1045-423: A paper describing the phylogeny of small benthic freshwater fish called darters, five undescribed putative species (Ozark, Sheltowee, Wildcat, Ihiyo, and Mamequit darters), notable for brightly colored nuptial males with distinctive color patterns, were referred to as " Etheostoma cf. spectabile " because they had been viewed as related to, but distinct from, Etheostoma spectabile (orangethroat darter). This view
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#17327917607681140-410: A personal name is to be converted to a specific epithet. In particular, names ending in a consonant (but not "er") are treated as first being converted into Latin by adding "-ius" (for a man) or "-ia" (for a woman), and then being made genitive (i.e. meaning "of that person or persons"). This produces specific epithets like lecardii for Lecard (male), wilsoniae for Wilson (female), and brauniarum for
1235-412: A serrated margin; both the stem and leaves are densely glandular-hairy. Tomato flowers are bisexual and are able to self fertilize. As tomatoes were moved from their native areas, their traditional pollinators (probably a species of halictid bee ) did not move with them. The trait of self-fertility became an advantage, and domestic cultivars of tomato have been selected to maximize this trait. This
1330-551: A single genus. The full binomial name must be unique within each code. From the early 19th century onwards it became ever more apparent that a body of rules was necessary to govern scientific names. In the course of time these became nomenclature codes . The International Code of Zoological Nomenclature ( ICZN ) governs the naming of animals, the International Code of Nomenclature for algae, fungi, and plants ( ICNafp ) that of plants (including cyanobacteria ), and
1425-593: A species retains its binomial name if it is moved from one family to another or from one order to another, unless it better fits a different genus in the same or different family, or it is split from its old genus and placed in a newly created genus. The independence is only partial since the names of families and other higher taxa are usually based on genera. Taxonomy includes both nomenclature and classification. Its first stages (sometimes called " alpha taxonomy ") are concerned with finding, describing and naming species of living or fossil organisms. Binomial nomenclature
1520-455: A staple of the peasant population because they were not as filling as other crops. Additionally, both toxic and inedible varieties discouraged many people from attempting to consume or prepare any other varieties. In certain areas of Italy, such as Florence, the fruit was used solely as a tabletop decoration, until it was incorporated into the local cuisine in the late 17th or early 18th century. The earliest discovered cookbook with tomato recipes
1615-655: A strong savoury umami flavor, and is an important ingredient in cuisines around the world. It is used in pizzas , pasta sauces, soups such as gazpacho , curries including dhansak and rogan josh , as juice, and in Bloody Mary cocktails . Tomato festivals are held annually in Buñol , Spain, in Reynoldsburg, Ohio , and in Närpes , Finland. The word tomato comes from the Spanish tomate , which in turn comes from
1710-462: A time management method Topics referred to by the same term [REDACTED] This disambiguation page lists articles associated with the title Pomodoro . If an internal link led you here, you may wish to change the link to point directly to the intended article. Retrieved from " https://en.wikipedia.org/w/index.php?title=Pomodoro&oldid=1123114251 " Category : Disambiguation pages Hidden categories: Short description
1805-403: Is Anthus hodgsoni berezowskii . Informally, in some circumstances, a form may be appended. For example Harmonia axyridis f. spectabilis is the harlequin ladybird in its black or melanic forms having four large orange or red spots. In botany, there are many ranks below species and although the name itself is written in three parts, a "connecting term" (not part of the name) is needed to show
1900-462: Is tobacco mosaic virus . Handling cigarettes and other infected tobacco products can transmit the virus to tomato plants. A serious disease is curly top , carried by the beet leafhopper , which interrupts the lifecycle. As the name implies, it has the symptom of making the top leaves of the plant wrinkle up and grow abnormally. Bacterial wilt is another common disease impacting yield. Wang et al. , 2019 found phage combination therapies to reduce
1995-409: Is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms , although they can be based on words from other languages. Such a name is called a binomial name (which may be shortened to just "binomial"), a binomen , binominal name , or a scientific name ; more informally it is also historically called a Latin name . In
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#17327917607682090-518: Is a species of frog found in Java , Indonesia. The second part of the name, which identifies the species within the genus, is also treated grammatically as a Latin word. It can have one of a number of forms: Whereas the first part of a binomial name must be unique within the purview of each nomenclatural code, the second part is quite commonly used in two or more genera (as is shown by examples of hodgsonii above), but cannot be used more than once within
2185-466: Is aptly termed Phalangium Ephemerum Virginianum , Soon-Fading Spiderwort of Virginia". The Latin phrases are short descriptions, rather than identifying labels. The Bauhins , in particular Caspar Bauhin (1560–1624), took some important steps towards the binomial system by pruning the Latin descriptions, in many cases to two words. The adoption by biologists of a system of strictly binomial nomenclature
2280-439: Is different from Wikidata All article disambiguation pages All disambiguation pages Tomato The tomato ( US : / t ə m eɪ t oʊ / , UK : / t ə m ɑː t oʊ / ), Solanum lycopersicum , is a plant whose fruit is an edible berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco , potato , and chili peppers . It originated from and
2375-461: Is due to Swedish botanist and physician Carl Linnaeus (1707–1778). It was in Linnaeus's 1753 Species Plantarum that he began consistently using a one-word trivial name ( nomen triviale ) after a generic name (genus name) in a system of binomial nomenclature. Trivial names had already appeared in his Critica Botanica (1737) and Philosophia Botanica (1751). This trivial name is what
2470-486: Is increasing, providing fruit during those times of the year when field-grown fruit is not readily available. Smaller fruit (cherry and grape), or cluster tomatoes (fruit-on-the-vine) are the fruit of choice for the large commercial greenhouse operators while the beefsteak varieties are the choice of owner-operator growers. Tomatoes are also grown using hydroponics . To facilitate transportation and storage, tomatoes are often picked unripe (green) and ripened in storage with
2565-614: Is not in sight. (There is also a published code for a different system of biotic nomenclature, which does not use ranks above species, but instead names clades . This is called PhyloCode .) As noted above, there are some differences between the codes in how binomials can be formed; for example the ICZN allows both parts to be the same, while the ICNafp does not. Another difference is in how personal names are used in forming specific names or epithets. The ICNafp sets out precise rules by which
2660-401: Is not the same as self-pollination , despite the common claim that tomatoes do so. That tomatoes pollinate themselves poorly without outside aid is clearly shown in greenhouse situations, where pollination must be aided by artificial wind, vibration of the plants, or by cultured bumblebees . The flowers develop on the apical meristem . They have the anthers fused along the edges, forming
2755-494: Is not used because it violates the International Code of Nomenclature barring the use of tautonyms in botanical nomenclature. The corrected name Lycopersicon lycopersicum (Nicolson 1974) was technically valid, because Miller's genus name and Linnaeus's species name differ in exact spelling. As Lycopersicon esculentum has become so well known, it was officially listed as a nomen conservandum in 1983, and would be
2850-433: Is now known as a specific epithet ( ICNafp ) or specific name ( ICZN ). The Bauhins' genus names were retained in many of these, but the descriptive part was reduced to a single word. Linnaeus's trivial names introduced the important new idea that the function of a name could simply be to give a species a unique label, meaning that the name no longer needed to be descriptive. Both parts could, for example, be derived from
2945-426: Is thus an important part of taxonomy as it is the system by which species are named. Taxonomists are also concerned with classification, including its principles, procedures and rules. A complete binomial name is always treated grammatically as if it were a phrase in the Latin language (hence the common use of the term "Latin name" for a binomial name). However, the two parts of a binomial name can each be derived from
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3040-436: Is usually followed by the "authority" – a way of designating the scientist(s) who first published the name. The authority is written in slightly different ways in zoology and botany. For names governed by the ICZN the surname is usually written in full together with the date (normally only the year) of publication. One example of author citation of scientific name is: " Amabela Möschler, 1880 ." The ICZN recommends that
3135-481: Is when several species from the same genus are being listed or discussed in the same paper or report, or the same species is mentioned repeatedly; in which case the genus is written in full when it is first used, but may then be abbreviated to an initial (and a period/full stop). For example, a list of members of the genus Canis might be written as " Canis lupus , C. aureus , C. simensis ". In rare cases, this abbreviated form has spread to more general use; for example,
3230-427: The International Code of Nomenclature for algae, fungi, and plants ( ICNafp or ICN ). Although the general principles underlying binomial nomenclature are common to these two codes, there are some differences in the terminology they use and their particular rules. In modern usage, the first letter of the generic name is always capitalized in writing, while that of the specific epithet is not, even when derived from
3325-487: The International Code of Nomenclature of Bacteria ( ICNB ) that of bacteria (including Archaea ). Virus names are governed by the International Committee on Taxonomy of Viruses ( ICTV ), a taxonomic code, which determines taxa as well as names. These codes differ in certain ways, e.g.: Unifying the different codes into a single code, the " BioCode ", has been suggested , although implementation
3420-695: The Columbian exchange ; certainly the tomato was being grown in Europe within a few years of that event. The earliest discussion of the tomato in European literature appeared in Pietro Andrea Mattioli 's 1544 herbal . He suggested that a new type of eggplant had been brought to Italy. He stated that it was blood red or golden color when mature, and could be divided into segments and eaten like an eggplant—that is, cooked and seasoned with salt, black pepper, and oil. Ten years later Mattioli named
3515-464: The ICZN is more protective of the form used by the original author. By tradition, the binomial names of species are usually typeset in italics; for example, Homo sapiens . Generally, the binomial should be printed in a font style different from that used in the normal text; for example, " Several more Homo sapiens fossils were discovered ." When handwritten, a binomial name should be underlined; for example, Homo sapiens . The first part of
3610-488: The International Code of Zoological Nomenclature (ICZN), the system is also called binominal nomenclature , with an "n" before the "al" in "binominal", which is not a typographic error, meaning "two-name naming system". The first part of the name – the generic name – identifies the genus to which the species belongs, whereas the second part – the specific name or specific epithet – distinguishes
3705-473: The Nahuatl word tomatl [ˈtomat͡ɬ] pronunciation . The specific name lycopersicum , meaning "wolf peach", originated with Galen , who used it to denote a plant that has never been identified. Luigi Anguillara speculated in the 16th century that Galen's lycopersicum might be the tomato, and despite the impossibility of this identification, lycopersicum entered scientific use as
3800-432: The authority for a binomial name is usually given, at least when it is first mentioned, and the year of publication may be specified. The word binomial is composed of two elements: bi- ( Latin prefix meaning 'two') and nomial (the adjective form of nomen , Latin for 'name'). In Medieval Latin, the related word binomium was used to signify one term in a binomial expression in mathematics. In fact,
3895-654: The potato , tomatoes belong to the genus Solanum , which is a member of the nightshade family, the Solanaceae . That is a diverse family of flowering plants, often poisonous, that includes the mandrake ( Mandragora ), deadly nightshade ( Atropa ), and tobacco ( Nicotiana ), as shown in the outline phylogenetic tree (many branches omitted). many garden flowers and other species Nicotiana (tobacco) Atropa (nightshades) Mandragora (mandrake) (sweet and bell peppers) S. lycopersicum (tomato) S. tuberosum (potato) In 1753, Linnaeus placed
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3990-412: The tomato bug , stink bugs , cutworms , tomato hornworms and tobacco hornworms , aphids , cabbage loopers , whiteflies , tomato fruitworms , flea beetles , red spider mite , slugs , and Colorado potato beetles . The tomato russet mite , Aculops lycopersici , feeds on foliage and young fruit of tomato plants, causing shrivelling and necrosis of leaves, flowers, and fruit, possibly killing
4085-697: The "U" phenotype are beneficial here, but have the disadvantage of leaving green shoulders near the stems of the ripe fruit, and even cracked yellow shoulders. This is apparently because of oxidative stress due to overload of the photosynthetic chain in direct sunlight at high temperatures. Hence, genetic design of a commercial variety that combines the advantages of types "u" and "U" requires fine tuning, but may be feasible. Breeders strive to produce tomato plants with improved yield, shelf life, size, and resistance to environmental pressures, including disease. These efforts have yielded unintended negative consequences on various fruit attributes. For instance, linkage drag ,
4180-465: The "cf." qualifier vary. In paleontology, it is typically used when the identification is not confirmed. For example, " Corvus cf. nasicus " was used to indicate "a fossil bird similar to the Cuban crow but not certainly identified as this species". In molecular systematics papers, "cf." may be used to indicate one or more undescribed species assumed to be related to a described species. For example, in
4275-404: The "original author and date of a name should be cited at least once in each work dealing with the taxon denoted by that name." For names governed by the ICNafp the name is generally reduced to a standard abbreviation and the date omitted. The International Plant Names Index maintains an approved list of botanical author abbreviations. Historically, abbreviations were used in zoology too. When
4370-523: The 1540s. It was probably eaten shortly after it was introduced, and was certainly being used as food by the early 17th century in Spain, as documented in the 1618 play La octava maravilla by Lope de Vega with "lovelier than ... a tomato in season". The tomato was introduced to China, likely via the Philippines or Macau, in the 16th century. It was given the name 番茄 fānqié (foreign eggplant), as
4465-491: The Braun sisters. By contrast, the ICZN does not require the intermediate creation of a Latin form of a personal name, allowing the genitive ending to be added directly to the personal name. This explains the difference between the names of the plant Magnolia hodgsonii and the bird Anthus hodgsoni . Furthermore, the ICNafp requires names not published in the form required by the code to be corrected to conform to it, whereas
4560-619: The Chinese named many foodstuffs introduced from abroad, but referring specifically to early introductions. In 1548, when the house steward of Cosimo de' Medici , the grand duke of Tuscany , wrote to the Medici private secretary informing him that the basket of tomatoes sent from the grand duke's Florentine estate at Torre del Gallo "had arrived safely". Tomatoes were grown mainly as ornamentals early on after their arrival in Italy. For example,
4655-499: The Florentine aristocrat Giovanvettorio Soderini wrote how they "were to be sought only for their beauty", and were grown only in gardens or flower beds. The tomato's ability to mutate and create new and different varieties helped contribute to its success and spread throughout Italy. However, in areas where the climate supported growing tomatoes, their habit of growing close to the ground suggested low status. They were not adopted as
4750-486: The Latin word binomium may validly refer to either of the epithets in the binomial name, which can equally be referred to as a binomen (pl. binomina ). Prior to the adoption of the modern binomial system of naming species, a scientific name consisted of a generic name combined with a specific name that was from one to several words long. Together they formed a system of polynomial nomenclature. These names had two separate functions. First, to designate or label
4845-547: The bacterium Escherichia coli is often referred to as just E. coli , and Tyrannosaurus rex is perhaps even better known simply as T. rex , these two both often appearing in this form in popular writing even where the full genus name has not already been given. The abbreviation "sp." is used when the actual specific name cannot or need not be specified. The abbreviation "spp." (plural) indicates "several species". These abbreviations are not italicised (or underlined). For example: " Canis sp." means "an unspecified species of
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#17327917607684940-414: The binomial, the genus name, is always written with an initial capital letter. Older sources, particularly botanical works published before the 1950s, used a different convention: if the second part of the name was derived from a proper noun, e.g., the name of a person or place, a capital letter was used. Thus, the modern form Berberis darwinii was written as Berberis Darwinii . A capital was also used when
5035-452: The biochemistry (many of the alkaloids common to other Solanum species are conspicuously absent from the tomato). On the other hand, hybrids of tomato and diploid potato can be created in the lab by somatic fusion , and are partially fertile, providing evidence of the close relationship between these species. An international consortium of researchers from 10 countries began sequencing the tomato genome in 2004. A prerelease version of
5130-415: The brightest red to the deepest yellow. Sahagún mentioned Aztecs cooking various sauces, some with tomatoes of different sizes, serving them in city markets: "foods sauces, hot sauces; ... with tomatoes, ... sauce of large tomatoes, sauce of ordinary tomatoes, ..." The Spanish conquistador Hernán Cortés 's capture of Tenochtitlan in 1521 initiated the widespread cultural and biological interchange called
5225-475: The correct name for the tomato in classifications which do not place the tomato in the genus Solanum . Genetic evidence shows that Linnaeus was correct to put the tomato in the genus Solanum , making S. lycopersicum the correct name. Both names, however, will probably be found in the literature for some time. Two of the major reasons for considering the genera separate are the leaf structure (tomato leaves are markedly different from any other Solanum ), and
5320-440: The culinary use of the tomato included Michele Felice Cornè and Robert Gibbon Johnson . Many Americans considered tomatoes to be poisonous at this time and, in general, they were grown more as ornamental plants than as food. In 1897, W. H. Garrison stated, "The belief was once transmitted that the tomato was sinisterly dangerous." He recalled in his youth tomatoes were dubbed "love-apples or wolf-apples" and shunned as "globes of
5415-502: The devil". When Alexander W. Livingston (1821–1898) began developing the tomato as a commercial crop, his aim had been to grow tomatoes smooth in contour, uniform in size, and sweet in flavor. He eventually developed over seventeen varieties. The U.S. Department of Agriculture's 1937 yearbook declared that "half of the major varieties were a result of the abilities of the Livingstons to evaluate and perpetuate superior material in
5510-525: The direct ancestor of some modern cultivated tomatoes. The Aztecs raised several varieties of tomato, with red tomatoes called xitomatl and green tomatoes (physalis) called tomatl ( tomatillo ). Bernardino de Sahagún reported seeing a great variety of tomatoes in the Aztec market at Tenochtitlán (Mexico City): "large tomatoes, small tomatoes, leaf tomatoes, sweet tomatoes, large serpent tomatoes, nipple-shaped tomatoes", and tomatoes of all colors from
5605-431: The early blight pathogen Alternaria solani . However, this tactic has limitations, since selection for traits such as pathogen resistance can negatively impact other favorable traits such as fruit production. The tomato is grown worldwide for its edible fruits , with thousands of cultivars . Greenhouse tomato production in large-acreage commercial greenhouses and owner-operator stand-alone or multiple-bay greenhouses
5700-407: The endings used differ between zoology and botany. Ranks below species receive three-part names, conventionally written in italics like the names of species. There are significant differences between the ICZN and the ICNafp . In zoology, the only formal rank below species is subspecies and the name is written simply as three parts (a trinomen). Thus, one of the subspecies of the olive-backed pipit
5795-453: The first part of the name of the species is a genus name. In a classification system based on ranks, there are also ways of naming ranks above the level of genus and below the level of species. Ranks above genus (e.g., family, order, class) receive one-part names, which are conventionally not written in italics. Thus, the house sparrow, Passer domesticus , belongs to the family Passeridae . Family names are normally based on genus names, although
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#17327917607685890-625: The fresh market and for canning and processing. The University of California, Davis 's C.M. Rick Tomato Genetics Resource Center maintains a gene bank of wild relatives, monogenic mutants and genetic stocks. Research on processing tomatoes is also conducted by the California Tomato Research Institute in Escalon, California . In California, growers have used a method of cultivation called dry-farming , especially with Early Girl tomatoes. This technique encourages
5985-702: The fruits in print as pomi d'oro , or "golden apples". After the Spanish colonization of the Americas , the Spanish distributed the tomato throughout their colonies in the Caribbean . They brought it to the Philippines , from where it spread to southeast Asia and then the whole of Asia. The Spanish brought the tomato to Europe, where it grew easily in Mediterranean climates ; cultivation began in
6080-595: The genome was made available in December 2009. The complete genome for the cultivar Heinz 1706 was published on 31 May 2012 in Nature . The latest reference genome published in 2021 had 799 MB and encodes 34,384 (predicted) proteins, spread over 12 chromosomes. The first commercially available genetically modified food was a tomato called Flavr Savr , which was engineered to have a longer shelf life. It could be vine ripened without compromising shelf life . However,
6175-513: The genus Canis ", while " Canis spp." means "two or more species of the genus Canis ". (These abbreviations should not be confused with the abbreviations "ssp." (zoology) or "subsp." (botany), plurals "sspp." or "subspp.", referring to one or more subspecies . See trinomen (zoology) and infraspecific name .) The abbreviation " cf. " (i.e., confer in Latin) is used to compare individuals/taxa with known/described species. Conventions for use of
6270-747: The impact of bacterial wilt, sometimes by reducing bacterial abundance and sometimes by selecting for resistant but slow growing genetics. Tomatoes, with their umami flavor, are extensively used in Mediterranean cuisine as a key ingredient in pizza and many pasta sauces. Tomatoes are used in Spanish gazpacho and Catalan pa amb tomàquet . The tomato is a crucial and ubiquitous part of Middle Eastern cuisine , served fresh in salads (e.g., Arab salad , Israeli salad , Shirazi salad and Turkish salad ), grilled with kebabs and other dishes, made into sauces, and so on. Binomial nomenclature In taxonomy , binomial nomenclature ("two-term naming system"), also called binary nomenclature ,
6365-551: The introduction of an undesired trait during backcrossing , has altered the metabolism of the fruit. This trait is physically close to the desired allele along the chromosome. Breeding for traits like larger fruit has thus unintentionally altered nutritional value and flavor. Breeders have turned to wild tomato species as a source of alleles to introduce beneficial traits into modern varieties. For example, wild relatives may possess higher amounts of fruit solids (associated with greater sugar content), or resistance to diseases such as to
6460-642: The name 'Love Apple' as a French misreading of the Italian pomo dei Mori ("the Moors' apple") as pomme d'amour , ("apple of love"). The tomato was introduced to cultivation in the Middle East by John Barker , British consul in Aleppo c. 1799 to 1825 . Nineteenth century descriptions of its consumption are uniformly as an ingredient in a cooked dish. In 1881, it is described as only eaten in
6555-451: The name is formed by two nouns in apposition, e.g., Panthera Leo or Centaurea Cyanus . In current usage, the second part is never written with an initial capital. When used with a common name, the scientific name often follows in parentheses, although this varies with publication. For example, "The house sparrow ( Passer domesticus ) is decreasing in Europe." The binomial name should generally be written in full. The exception to this
6650-552: The names of people. Thus Gerard's Phalangium ephemerum virginianum became Tradescantia virginiana , where the genus name honoured John Tradescant the Younger , an English botanist and gardener. A bird in the parrot family was named Psittacus alexandri , meaning "Alexander's parrot", after Alexander the Great , whose armies introduced eastern parakeets to Greece. Linnaeus's trivial names were much easier to remember and use than
6745-495: The ordering of items into groups based on similarities or differences; in biological classification , species are one of the kinds of item to be classified. In principle, the names given to species could be completely independent of their classification. This is not the case for binomial names, since the first part of a binomial is the name of the genus into which the species is placed. Above the rank of genus, binomial nomenclature and classification are partly independent; for example,
6840-422: The original name is changed, e.g., the species is moved to a different genus, both codes use parentheses around the original authority; the ICNafp also requires the person who made the change to be given. In the ICNafp , the original name is then called the basionym . Some examples: Binomial nomenclature, as described here, is a system for naming species. Implicitly, it includes a system for naming genera, since
6935-470: The parallel polynomial names, and eventually replaced them. The value of the binomial nomenclature system derives primarily from its economy, its widespread use, and the uniqueness and stability of names that the Codes of Zoological and Botanical , Bacterial and Viral Nomenclature provide: Binomial nomenclature for species has the effect that when a species is moved from one genus to another, sometimes
7030-403: The plant hormone ethylene . At industrial scale, such as for canning, tomatoes are picked mechanically. The machine cuts the whole vine and uses sensors to separate ripe tomatoes from the rest of the plant, which is returned to the farm for use either as green manure or to be grazed by livestock . In 2022, world production of tomatoes was 186 million tonnes , with China accounting for 37% of
7125-422: The plant to send roots deep to find existing moisture. Tomato plants are vines, becoming decumbent , and can grow up to 3 m (9.8 ft); bush varieties are generally no more than 100 cm (3 ft 3 in) tall. They are tender perennials, often grown as annuals. Tomato plants are dicots . They grow as a series of branching stems, with a terminal bud at the tip that does the actual growing. When
7220-430: The plant. After an insect attack tomato plants produce systemin , a plant peptide hormone . This activates defensive mechanisms, such as the production of protease inhibitors to slow the growth of insects. The hormone was first identified in tomatoes. Tomato cultivars vary widely in their resistance to disease. Modern hybrids focus on improving disease resistance over the heirloom plants . A common tomato disease
7315-408: The product was not commercially successful, and was sold only until 1997. The poor taste and lack of sugar in modern garden and commercial tomato varieties resulted from breeding tomatoes to ripen uniformly red. This change occurred after discovery of a mutant "u" phenotype in the mid-20th century, so named because the fruits ripened uniformly. This was widely cross-bred to produce red fruit without
7410-722: The region "within the last forty years". The earliest reference to tomatoes being grown in British North America is from 1710, when herbalist William Salmon saw them in what is today South Carolina , perhaps introduced from the Caribbean. By the mid-18th century, they were cultivated on some Carolina plantations, and probably in other parts of the Southeast. Thomas Jefferson , who ate tomatoes in Paris, sent some seeds back to America. Some early American advocates of
7505-589: The species within the genus. For example, modern humans belong to the genus Homo and within this genus to the species Homo sapiens . Tyrannosaurus rex is likely the most widely known binomial. The formal introduction of this system of naming species is credited to Carl Linnaeus , effectively beginning with his work Species Plantarum in 1753. But as early as 1622, Gaspard Bauhin introduced in his book Pinax theatri botanici (English, Illustrated exposition of plants ) containing many names of genera that were later adopted by Linnaeus. Binomial nomenclature
7600-491: The species, and second, to be a diagnosis or description; however, these two goals were eventually found to be incompatible. In a simple genus, containing only two species, it was easy to tell them apart with a one-word genus and a one-word specific name; but as more species were discovered, the names necessarily became longer and unwieldy, for instance, Plantago foliis ovato-lanceolatus pubescentibus, spica cylindrica, scapo tereti ("plantain with pubescent ovate-lanceolate leaves,
7695-483: The specific name or epithet must be changed as well. This may happen because the specific name is already used in the new genus, or to agree in gender with the new genus if the specific epithet is an adjective modifying the genus name. Some biologists have argued for the combination of the genus name and specific epithet into a single unambiguous name, or for the use of uninomials (as used in nomenclature of ranks above species). Because genus names are unique only within
7790-590: The tip eventually stops growing, whether because of pruning or flowering, lateral buds take over and grow into new, fully functional, vines. Tomato vines are typically pubescent, meaning covered with fine short hairs. The hairs facilitate the vining process, turning into roots wherever the plant is in contact with the ground and moisture, especially if the vine's connection to its original root has been damaged or severed. The leaves are 10–25 cm (4–10 in) long, odd pinnate , with five to nine leaflets on petioles , each leaflet up to 8 cm (3 in) long, with
7885-518: The tomato in England. Gerard knew the tomato was eaten in Spain and Italy. Nonetheless, he believed it was poisonous. Gerard's views were influential, and the tomato was considered unfit for eating for many years in Britain and its North American colonies . By 1820, tomatoes were described as "to be seen in great abundance in all our vegetable markets" and to be "used by all our best cooks", reference
7980-411: The tomato in the genus Solanum (alongside the potato) as Solanum lycopersicum . In 1768, Philip Miller moved it to its own genus, naming it Lycopersicon esculentum . The name came into wide use, but was technically in breach of the plant naming rules because Linnaeus's species name lycopersicum still had priority. Although the name Lycopersicum lycopersicum was suggested by Karsten (1888), it
8075-588: The tomato, the red-fruited Solanum pimpinellifolium , is native to western South America, where it was probably first domesticated. The resulting domesticated plant, ancestral to the modern large-fruited tomato varieties, was probably the cherry tomato, S. lycopersicum var. cerasiforme . However, genomic analysis suggests that the domestication process may have been more complex than this. S. lycopersicon var. cerasiforme may have existed before domestication, while traits supposedly typical of domestication may have been reduced in that variety and then reselected (in
8170-773: The tomato." Livingston's first breed of tomato, the Paragon, was introduced in 1870. In 1875, he introduced the Acme, said to be in the parentage of most cultivars for the next twenty-five years. Other early breeders included Henry Tilden in Iowa and a Dr. Hand in Baltimore. Because of the tomato's need for heat and a long growing season, several states in the Sun Belt became major producers, particularly Florida and California . In California, tomatoes are grown under irrigation for both
8265-666: The total, followed by India , Turkey , and the United States as major producers (table). The world dedicated 4.8 million hectares in 2012 for tomato cultivation and the total production was about 161.8 million tonnes . The average world farm yield for tomato was 33.6 tonnes per hectare in 2012. Tomato farms in the Netherlands were the most productive in 2012, with a nationwide average of 476 tonnes per hectare, followed by Belgium (463 tonnes per hectare) and Iceland (429 tonnes per hectare). Common tomato pests include
8360-424: The typical green ring around the stem on un-crossbred varieties. Before this, most tomatoes produced more sugar during ripening, and were sweeter and more flavorful. 10–20% of the total carbon fixed in the fruit can be produced by photosynthesis in the developing fruit of the normal U phenotype. The u mutation encodes a factor that produces defective chloroplasts with lower density in developing fruit, making them
8455-399: The well known and highly prized San Marzano tomato grown in that region, with a European protected designation of origin certification. Tomatoes were not grown in England until the 1590s. One of the earliest cultivators was John Gerard , a barber-surgeon . Gerard's Herbal , published in 1597, and largely plagiarized from continental sources, is also one of the earliest discussions of
8550-713: Was domesticated in western South America . It was introduced to the Old World by the Spanish in the Columbian exchange in the 16th century. Tomato plants are vines, largely annual and vulnerable to frost, though sometimes living longer in greenhouses. The flowers are able to self-fertilise. Modern varieties have been bred to ripen uniformly red, in a process that has impaired the fruit's sweetness and flavor. There are thousands of cultivars, varying in size, color, shape, and flavor. Tomatoes are attacked by many insect pests and nematodes, and are subject to diseases caused by viruses and by mildew and blight fungi. The tomato has
8645-428: Was introduced in order to provide succinct, relatively stable and verifiable names that could be used and understood internationally, unlike common names which are usually different in every language. The application of binomial nomenclature is now governed by various internationally agreed codes of rules, of which the two most important are the International Code of Zoological Nomenclature ( ICZN ) for animals and
8740-565: Was made to their cultivation in gardens still "for the singularity of their appearance", while their use in cooking was associated with exotic Italian or Jewish cuisine. For example, in Elizabeth Blackwell 's A Curious Herbal , it is described under the name "Love Apple ( Amoris Pomum )" as being consumed with oil and vinegar in Italy, similar to consumption of cucumbers in the UK. In 1963, The New York Times gave an explanation of
8835-514: Was ornamental, not least because it was understood to be related to the nightshades and assumed to be poisonous. The exact date of domestication is unknown; by 500 BC, it was already being cultivated in southern Mexico and probably other areas. The Pueblo people believed that tomato seeds could confer powers of divination . The large, lumpy variety of tomato, a mutation from a smoother, smaller fruit, originated in Mesoamerica, and may be
8930-536: Was published in Naples in 1692, though the author had apparently obtained these recipes from Spanish sources. Varieties were developed over the following centuries for drying, for sauce, for pizzas, and for long-term storage. These varieties are usually known for their place of origin as much as by a variety name. For example, there is the Pomodorino del Piennolo del Vesuvio , the "hanging tomato of Vesuvius", and
9025-410: Was supported to varying degrees by DNA analysis. The somewhat informal use of taxa names with qualifying abbreviations is referred to as open nomenclature and it is not subject to strict usage codes. In some contexts, the dagger symbol ("†") may be used before or after the binomial name to indicate that the species is extinct. In scholarly texts, at least the first or main use of the binomial name
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