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89-489: Historically, the name Pindos refers to the mountainous territory that separates the greater Epirus region from the regions of Macedonia and Thessaly. According to John Tzetzes (a 12th-century Byzantine writer), the Pindos range was then called Metzovon. When Anastasios Gordios  [ el ] translated (between 1682/83 and 1689) to a more conversational (colloquial) language the initial praise to St. Vissarion, which

178-552: A mated pair accompanied by their offspring. Offspring may leave to form their own packs on the onset of sexual maturity and in response to competition for food within the pack. Wolves are also territorial , and fights over territory are among the principal causes of mortality. The wolf is mainly a carnivore and feeds on large wild hooved mammals as well as smaller animals, livestock, carrion , and garbage. Single wolves or mated pairs typically have higher success rates in hunting than do large packs. Pathogens and parasites, notably

267-530: A taxonomic synonym for the dingo . Wozencraft referred to a 1999 mitochondrial DNA (mtDNA) study as one of the guides in forming his decision, and listed the 38 subspecies of C. lupus under the biological common name of "wolf", the nominate subspecies being the Eurasian wolf ( C. l. lupus ) based on the type specimen that Linnaeus studied in Sweden. Studies using paleogenomic techniques reveal that

356-469: A French encyclopedia of 1756). BY 1825, the traveller John Cam Hobhouse was writing that "…the latter mountains, now known by the name of Metzovo, can be no other than Pindus itself…" while a patriarchal document of 1818 states: "Because the high mountain of Pindos in Epirus, that is commonly called Messovon...". The word Pindos was used more in literary sources, while the folk name for the mountain range from

445-520: A broader snout, shorter ears, a shorter torso and a longer tail. It is slender and powerfully built, with a large, deeply descending rib cage , a sloping back, and a heavily muscled neck. The wolf's legs are moderately longer than those of other canids, which enables the animal to move swiftly, and to overcome the deep snow that covers most of its geographical range in winter, though more short-legged ecomorphs are found in some wolf populations. The ears are relatively small and triangular. The wolf's head

534-635: A considerable amount of social and biographical information, and are full of learned allusions to history, rhetoric, and mythology. These letters became the springboard for what became during the Renaissance perhaps the most influential of his many works, the Book of Histories , usually called Chiliades ("thousands") from the arbitrary division by its first editor (N. Gerbel, 1546) into books each containing 1,000 lines. The work consists of 12,674 lines of political verse , divided into 660 topics, each of which

623-448: A family of eight wolves (two adults, juveniles, and yearlings), or sometimes two or three such families, with examples of exceptionally large packs consisting of up to 42 wolves being known. Cortisol levels in wolves rise significantly when a pack member dies, indicating the presence of stress. During times of prey abundance caused by calving or migration, different wolf packs may join together temporarily. Offspring typically stay in

712-433: A few times and then retreating from a perceived danger. Aggressive or self-assertive wolves are characterized by their slow and deliberate movements, high body posture and raised hackles , while submissive ones carry their bodies low, flatten their fur, and lower their ears and tail. Scent marking involves urine, feces, and preputial and anal gland scents. This is more effective at advertising territory than howling and

801-674: A museum specimen of a wolf from southern China collected in 1963 showed a genome that was 12–14% admixed from this unknown canid. In North America, some coyotes and wolves show varying degrees of past genetic admixture . In more recent times, some male Italian wolves originated from dog ancestry, which indicates female wolves will breed with male dogs in the wild. In the Caucasus Mountains , ten percent of dogs including livestock guardian dogs , are first generation hybrids. Although mating between golden jackals and wolves has never been observed, evidence of jackal-wolf hybridization

890-754: A press release by the U.S. Department of Agriculture in 1921, the infamous Custer Wolf relied on coyotes to accompany him and warn him of danger. Though they fed from his kills, he never allowed them to approach him. Interactions have been observed in Eurasia between wolves and golden jackals, the latter's numbers being comparatively small in areas with high wolf densities. Wolves also kill red , Arctic and corsac foxes , usually in disputes over carcasses, sometimes eating them. Brown bears typically dominate wolf packs in disputes over carcasses, while wolf packs mostly prevail against bears when defending their den sites. Both species kill each other's young. Wolves eat

979-462: A revised edition with marginal notes in prose and verse (ed. T. Kiessling, 1826; on the sources see C. Harder, De J. T. historiarum fontibus quaestiones selectae , diss., Kiel, 1886). Tzetzes supplemented Homer 's Iliad by a work that begins with the birth of Paris and continues the tale to the Achaeans' return home. The Homeric Allegories , in "political" verse and dedicated initially to

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1068-661: A shorter torso and a longer tail. The wolf is nonetheless related closely enough to smaller Canis species, such as the coyote and the golden jackal , to produce fertile hybrids with them. The wolf's fur is usually mottled white, brown, gray, and black, although subspecies in the arctic region may be nearly all white. Of all members of the genus Canis , the wolf is most specialized for cooperative game hunting as demonstrated by its physical adaptations to tackling large prey, its more social nature , and its highly advanced expressive behaviour , including individual or group howling . It travels in nuclear families consisting of

1157-409: A variety of arthropod exoparasites, including fleas , ticks , lice , and mites . The most harmful to wolves, particularly pups, is the mange mite ( Sarcoptes scabiei ), though they rarely develop full-blown mange , unlike foxes. Endoparasites known to infect wolves include: protozoans and helminths ( flukes , tapeworms , roundworms and thorny-headed worms ). Most fluke species reside in

1246-411: A weight of 69–79 kg (152–174 lb). The wolf has very dense and fluffy winter fur, with a short undercoat and long, coarse guard hairs . Most of the undercoat and some guard hairs are shed in spring and grow back in autumn. The longest hairs occur on the back, particularly on the front quarters and neck. Especially long hairs grow on the shoulders and almost form a crest on the upper part of

1335-625: Is a gloss on a literary, historical, or other learned reference in one of his published letters. The first 141 histories serve as poetic footnotes to a verse letter Tzetzes addressed to John Lachanas, an official in Constantinople. This collection of literary, historical, theological, and antiquarian miscellanies provides an important snapshot of the intellectual world of Constantinople in the mid-12th century, and also preserves fragments of more than 200 ancient authors, including many whose works have been lost. The author subsequently brought out

1424-479: Is difficult and ambiguous, with some authors choosing to include C. mosbachensis (which first appeared around 1.4 million years ago) as an early subspecies of C. lupus. Considerable morphological diversity existed among wolves by the Late Pleistocene . Many Late Pleistocene wolf populations had more robust skulls and teeth than modern wolves, often with a shortened snout , a pronounced development of

1513-525: Is dominated by wild medium-sized hoofed mammals and domestic species. The wolf depends on wild species, and if these are not readily available, as in Asia, the wolf is more reliant on domestic species. Across Eurasia, wolves prey mostly on moose , red deer , roe deer and wild boar . In North America, important range-wide prey are elk , moose, caribou , white-tailed deer and mule deer . Prior to their extirpation from North America, wild horses were among

1602-488: Is in a remote area in the northeast of the Pindus mountains, north of the town of Metsovo and south of Perivoli . The park of some 7,000 hectares was established in 1966. There are forests of black pine and beech , and in the higher parts, Bosnian Pine ( Pinus leucodermis ). The park is a refuge for bears , wild cats , and lynxes . The National Park was selected as main motif for two high value euro collectors' coins:

1691-1220: Is itself thought to be derived from the Proto-Germanic * wulfaz . The Proto-Indo-European root * wĺ̥kʷos may also be the source of the Latin word for the animal lupus (* lúkʷos ). The name "gray wolf" refers to the grayish colour of the species. Since pre-Christian times, Germanic peoples such as the Anglo-Saxons took on wulf as a prefix or suffix in their names. Examples include Wulfhere ("Wolf Army"), Cynewulf ("Royal Wolf"), Cēnwulf ("Bold Wolf"), Wulfheard ("Wolf-hard"), Earnwulf ("Eagle Wolf"), Wulfstān ("Wolf Stone") Æðelwulf ("Noble Wolf"), Wolfhroc ("Wolf-Frock"), Wolfhetan ("Wolf Hide"), Scrutolf ("Garb Wolf"), Wolfgang ("Wolf Gait") and Wolfdregil ("Wolf Runner"). Gray wolf [REDACTED] Coyote [REDACTED] African wolf [REDACTED] Ethiopian wolf [REDACTED] Golden jackal [REDACTED] Dhole [REDACTED] African wild dog [REDACTED] Side-striped jackal [REDACTED] Black-backed jackal [REDACTED] In 1758,

1780-399: Is large and heavy, with a wide forehead, strong jaws and a long, blunt muzzle. The skull is 230–280 mm (9–11 in) in length and 130–150 mm (5–6 in) in width. The teeth are heavy and large, making them better suited to crushing bone than those of other canids, though they are not as specialized as those found in hyenas . Its molars have a flat chewing surface, but not to

1869-541: Is largely survivable in wolves, but can be lethal to pups. Bacterial diseases carried by wolves include: brucellosis , Lyme disease , leptospirosis , tularemia , bovine tuberculosis , listeriosis and anthrax . Although lyme disease can debilitate individual wolves, it does not appear to significantly affect wolf populations. Leptospirosis can be contracted through contact with infected prey or urine, and can cause fever , anorexia , vomiting, anemia , hematuria , icterus , and death. Wolves are often infested with

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1958-703: Is most likely used mainly to induce vomiting to rid themselves of intestinal parasites or long guard hairs. They are known to eat the berries of mountain-ash , lily of the valley , bilberries , cowberries , European black nightshade , grain crops, and the shoots of reeds. In times of scarcity, wolves will readily eat carrion . In Eurasian areas with dense human activity, many wolf populations are forced to subsist largely on livestock and garbage. As prey in North America continue to occupy suitable habitats with low human density, North American wolves eat livestock and garbage only in dire circumstances. Cannibalism

2047-562: Is not uncommon in wolves during harsh winters, when packs often attack weak or injured wolves and may eat the bodies of dead pack members. Wolves typically dominate other canid species in areas where they both occur. In North America, incidents of wolves killing coyotes are common, particularly in winter, when coyotes feed on wolf kills. Wolves may attack coyote den sites, digging out and killing their pups, though rarely eating them. There are no records of coyotes killing wolves, though coyotes may chase wolves if they outnumber them. According to

2136-677: Is now extirpated (locally extinct) from much of its range in Western Europe, the United States and Mexico, and completely in the British Isles and Japan. In modern times, the wolf occurs mostly in wilderness and remote areas. The wolf can be found between sea level and 3,000 m (9,800 ft). Wolves live in forests, inland wetlands , shrublands , grasslands (including Arctic tundra ), pastures , deserts, and rocky peaks on mountains. Habitat use by wolves depends on

2225-458: Is often used in combination with scratch marks. Wolves increase their rate of scent marking when they encounter the marks of wolves from other packs. Lone wolves will rarely mark, but newly bonded pairs will scent mark the most. These marks are generally left every 240 m (260 yd) throughout the territory on regular travelways and junctions. Such markers can last for two to three weeks, and are typically placed near rocks, boulders, trees, or

2314-471: Is that elucidating the obscure Cassandra or Alexandra of the Hellenistic poet Lycophron , usually called "On Lycophron" (edited by K.O. Müller , 1811), in the production of which his brother Isaac is generally associated with him. Mention may also be made of a dramatic sketch in iambic verse, in which the caprices of fortune and the wretched lot of the learned are described; and of an iambic poem on

2403-595: Is usually constructed not more than 500 m (550 yd) away from a water source. It typically faces southwards where it can be better warmed by sunlight exposure, and the snow can thaw more quickly. Resting places, play areas for the pups, and food remains are commonly found around wolf dens. The odor of urine and rotting food emanating from the denning area often attracts scavenging birds like magpies and ravens . Though they mostly avoid areas within human sight, wolves have been known to nest near homes, paved roads and railways . During pregnancy, female wolves remain in

2492-435: Is usually founded by an unrelated dispersing male and female, travelling together in search of an area devoid of other hostile packs. Wolf packs rarely adopt other wolves into their fold and typically kill them. In the rare cases where other wolves are adopted, the adoptee is almost invariably an immature animal of one to three years old, and unlikely to compete for breeding rights with the mated pair. This usually occurs between

2581-659: The Allegoriai on the Iliad and the Odyssey , which are long didactic poems containing interpretations of Homeric theology. Tzetzes described himself as pure Greek on his father's side and part Iberian ( Georgian ) on his mother's side. In his works, Tzetzes states that his grandmother was a relative of the Georgian Bagratid princess Maria of Alania who came to Constantinople with her and later became

2670-744: The Antehomerica , Tzetzes recalls the events taking place before Homer's Iliad . This work was followed by the Homerica , covering the events of the Iliad , and the Posthomerica , reporting the events taking place between the Iliad and the Odyssey . All three are currently available in English translations. Tzetzes also wrote commentaries on a number of Greek authors, the most important of which

2759-601: The Eurasian lynx , which may feed on smaller prey where wolves are present and may be suppressed by large wolf populations. Wolves encounter cougars along portions of the Rocky Mountains and adjacent mountain ranges. Wolves and cougars typically avoid encountering each other by hunting at different elevations for different prey ( niche partitioning ). This is more difficult during winter. Wolves in packs usually dominate cougars and can steal their kills or even kill them, while one-to-one encounters tend to be dominated by

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2848-462: The gray wolf or grey wolf , is a canine native to Eurasia and North America . More than thirty subspecies of Canis lupus have been recognized, including the dog and dingo , though gray wolves, as popularly understood, only comprise naturally-occurring wild subspecies. The wolf is the largest wild extant member of the family Canidae , and is further distinguished from other Canis species by its less pointed ears and muzzle, as well as

2937-510: The incubation period for rabies is eight to 21 days, and results in the host becoming agitated, deserting its pack, and travelling up to 80 km (50 mi) a day, thus increasing the risk of infecting other wolves. Although canine distemper is lethal in dogs, it has not been recorded to kill wolves, except in Canada and Alaska. The canine parvovirus, which causes death by dehydration , electrolyte imbalance , and endotoxic shock or sepsis ,

3026-537: The rabies virus , may infect wolves. The global wild wolf population was estimated to be 300,000 in 2003 and is considered to be of Least Concern by the International Union for Conservation of Nature (IUCN). Wolves have a long history of interactions with humans, having been despised and hunted in most pastoral communities because of their attacks on livestock, while conversely being respected in some agrarian and hunter-gatherer societies. Although

3115-610: The temporalis muscle, and robust premolars . It is proposed that these features were specialized adaptations for the processing of carcass and bone associated with the hunting and scavenging of Pleistocene megafauna . Compared with modern wolves, some Pleistocene wolves showed an increase in tooth breakage similar to that seen in the extinct dire wolf . This suggests they either often processed carcasses, or that they competed with other carnivores and needed to consume their prey quickly. The frequency and location of tooth fractures in these wolves indicates they were habitual bone crackers like

3204-628: The Egyptian Sinai Peninsula showed admixture with Middle Eastern gray wolves and dogs. There is evidence of gene flow between golden jackals and Middle Eastern wolves, less so with European and Asian wolves, and least with North American wolves. This indicates the golden jackal ancestry found in North American wolves may have occurred before the divergence of the Eurasian and North American wolves. The common ancestor of

3293-399: The Eurasian wolf is a mixture of ochreous (yellow to orange) and rusty ochreous (orange/red/brown) colours with light gray. The muzzle is pale ochreous gray, and the area of the lips, cheeks, chin, and throat is white. The top of the head, forehead, under and between the eyes, and between the eyes and ears is gray with a reddish film. The neck is ochreous. Long, black tips on the hairs along

3382-578: The German-born empress Irene and then to Constantine Cotertzes , are two didactic poems, the first based on the Iliad and the second based on the Odyssey, in which Homer and the Homeric theology are set forth and then explained by means of three kinds of allegory : euhemeristic ( πρακτική ), anagogic ( ψυχική ) and physic ( στοιχειακή ). These works were translated into English in 2015 and 2019 by Adam J. Goldwyn and Dimitra Kokkini. In

3471-521: The Middle Ages up to the 19th century was either "Metsovo" or "the mountains of Metsovo". Most probably this name was not meant to indicate the whole range as it is meant today, but only its central part between the area of Aspropotamos and the springs of the Aoös River. This part coincides with the mountainous region which the ancient Greeks used to call Pindos. The most notable mountains in

3560-513: The Middle East, and Asia is estimated to be fairly recent at around 1,600 years ago. Among New World wolves, the Mexican wolf diverged around 5,400 years ago. In the distant past, there was gene flow between African wolves , golden jackals , and gray wolves. The African wolf is a descendant of a genetically admixed canid of 72% wolf and 28% Ethiopian wolf ancestry. One African wolf from

3649-574: The Pindus, one of them being Samarina , which boasts one of the highest elevations in Greece. The area had a traditional pastoral economy in which sheep were raised by shepherds who were ethnically Sarakatsani and Aromanian . Many of the villages such as Perivoli and Smixi include communities of Aromanians ( Vlachs ), originally shepherds and farmers. In recent decades, a number of villages, such as Metsovo , have developed into tourist resorts with ski facilities. The A2 motorway (Egnatia Odos) serves

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3738-518: The Swedish botanist and zoologist Carl Linnaeus published in his Systema Naturae the binomial nomenclature . Canis is the Latin word meaning " dog ", and under this genus he listed the doglike carnivores including domestic dogs, wolves, and jackals . He classified the domestic dog as Canis familiaris , and the wolf as Canis lupus . Linnaeus considered the dog to be a separate species from

3827-442: The abundance of prey, snow conditions, livestock densities, road densities, human presence and topography . Like all land mammals that are pack hunters , the wolf feeds predominantly on ungulates that can be divided into large size 240–650 kg (530–1,430 lb) and medium size 23–130 kg (51–287 lb), and have a body mass similar to that of the combined mass of the pack members. The wolf specializes in preying on

3916-481: The age of the pack's pups. They tend to increase in size in areas with low prey populations, or when the pups reach the age of six months when they have the same nutritional needs as adults. Wolf packs travel constantly in search of prey, covering roughly 9% of their territory per day, on average 25 km/d (16 mi/d). The core of their territory is on average 35 km (14 sq mi) where they spend 50% of their time. Prey density tends to be much higher on

4005-414: The age of two years and sexually mature from the age of three years. The age of first breeding in wolves depends largely on environmental factors: when food is plentiful, or when wolf populations are heavily managed, wolves can rear pups at younger ages to better exploit abundant resources. Females are capable of producing pups every year, one litter annually being the average. Oestrus and rut begin in

4094-437: The back form a broad stripe, with black hair tips on the shoulders, upper chest and rear of the body. The sides of the body, tail, and outer limbs are a pale dirty ochreous colour, while the inner sides of the limbs, belly, and groin are white. Apart from those wolves which are pure white or black, these tones vary little across geographical areas, although the patterns of these colours vary between individuals. In North America,

4183-655: The brown bears they kill, while brown bears seem to eat only young wolves. Wolf interactions with American black bears are much rarer because of differences in habitat preferences. Wolves have been recorded on numerous occasions actively seeking out American black bears in their dens and killing them without eating them. Unlike brown bears, American black bears frequently lose against wolves in disputes over kills. Wolves also dominate and sometimes kill wolverines , and will chase off those that attempt to scavenge from their kills. Wolverines escape from wolves in caves or up trees. Wolves may interact and compete with felids , such as

4272-645: The cat, who likewise will kill wolves. Wolves more broadly affect cougar population dynamics and distribution by dominating territory and prey opportunities and disrupting the feline's behaviour. Wolf and Siberian tiger interactions are well-documented in the Russian Far East , where tigers significantly depress wolf numbers, sometimes to the point of localized extinction . In Israel, Palestine, Central Asia and India wolves may encounter striped hyenas , usually in disputes over carcasses. Striped hyenas feed extensively on wolf-killed carcasses in areas where

4361-665: The coat colours of wolves follow Gloger's rule , wolves in the Canadian arctic being white and those in southern Canada, the U.S., and Mexico being predominantly gray. In some areas of the Rocky Mountains of Alberta and British Columbia, the coat colour is predominantly black, some being blue-gray and some with silver and black. Differences in coat colour between sexes is absent in Eurasia; females tend to have redder tones in North America. Black-coloured wolves in North America acquired their colour from wolf-dog admixture after

4450-460: The coyote and the wolf admixed with a ghost population of an extinct unidentified canid. This canid was genetically close to the dhole and evolved after the divergence of the African hunting dog from the other canid species. The basal position of the coyote compared to the wolf is proposed to be due to the coyote retaining more of the mitochondrial genome of this unidentified canid. Similarly,

4539-575: The critically endangered Balkan lynx, of which only 15 are estimated to remain. In the Greek section of the Pindus mountains there are two national parks. The Vikos-Aoos National Park is south of the town of Konitsa , in the west part of Zagori region. It includes Mount Tymfi , the Vikos Gorge and the Aoos Gorge. It was created in 1973. The Pindus National Park (also known as Valia Kalda)

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4628-755: The death of the emperor Manuel I Komnenos , noticeable for introducing at the beginning of each line the last word of the line preceding it (both in Pietro Matranga , Anecdota Graeca 1850). For the other works of Tzetzes see J. A. Fabricius , Bibliotheca graeca (ed. Harles ), xi.228, and Karl Krumbacher , Geschichte der byz. Litt. (2nd edition, 1897); monograph by G. Hart, "De Tzetzarum nomine, vitis, scriptis," in Jahn 's Jahrbucher für classische Philologie. Supplementband xii (Leipzig, 1881). Wolf See Subspecies of Canis lupus The wolf ( Canis lupus ; pl. : wolves ), also known as

4717-534: The extant wolf C. lupus from the earlier C. mosbachensis (which in turn descended from C. etruscus ) is widely accepted. Among the oldest fossils of the modern grey wolf is from Ponte Galeria in Italy, dating to 406,500 ± 2,400 years ago. Remains from Cripple Creek Sump in Alaska may be considerably older, around 1 million years old, though differentiating between the remains of modern wolves and C. mosbachensis

4806-482: The fear of wolves exists in many human societies, the majority of recorded attacks on people have been attributed to animals suffering from rabies . Wolf attacks on humans are rare because wolves are relatively few, live away from people, and have developed a fear of humans because of their experiences with hunters, farmers, ranchers, and shepherds. The English "wolf" stems from the Old English wulf , which

4895-465: The first arrival of dogs across the Bering Strait 12,000 to 14,000 years ago. Research into the inheritance of white colour from dogs into wolves has yet to be undertaken. Wolves occur across Eurasia and North America. However, deliberate human persecution because of livestock predation and fear of attacks on humans has reduced the wolf's range to about one-third of its historic range; the wolf

4984-506: The forests. The forests of this region have faced many threats over the course of human history, including overgrazing , agriculture, and deforestation . The greatest threats now come from the development of mountain tourism and ski resorts . Because of the instability of the soil on steep mountains, road-building and clear-cutting operations have led to dangerous landslides and the collapse of mountain slopes. Mining for bauxite , overgrazing, and over-collection of plants are also threatening

5073-466: The great biodiversity of this ecoregion. In the Albanian section of the Pindus mountains is Albania's newest national park. The Vjosa River is Europe's first Wild River National Park that was designated on 15 March 2023. The river valley is considered Albania's biodiversity hotspot, offering ideal aquatic habitats for over 1,100 species of wildlife, including otters, the endangered Egyptian vulture and

5162-834: The hind feet are 220–250 mm ( 8 + 5 ⁄ 8 – 9 + 7 ⁄ 8  in). The size and weight of the modern wolf increases proportionally with latitude in accordance with Bergmann's rule . The mean body mass of the wolf is 40 kg (88 lb), the smallest specimen recorded at 12 kg (26 lb) and the largest at 79.4 kg (175 lb). On average, European wolves weigh 38.5 kg (85 lb), North American wolves 36 kg (79 lb), and Indian and Arabian wolves 25 kg (55 lb). Females in any given wolf population typically weigh 2.3–4.5 kg (5–10 lb) less than males. Wolves weighing over 54 kg (119 lb) are uncommon, though exceptionally large individuals have been recorded in Alaska and Canada. In central Russia, exceptionally large males can reach

5251-478: The lineage that led to dogs from other Old World wolves around 11,100–12,300 years ago. An extinct Late Pleistocene wolf may have been the ancestor of the dog, with the dog's similarity to the extant wolf being the result of genetic admixture between the two. The dingo, Basenji , Tibetan Mastiff and Chinese indigenous breeds are basal members of the domestic dog clade. The divergence time for wolves in Europe,

5340-424: The middle of the back is 60–70 mm ( 2 + 3 ⁄ 8 – 2 + 3 ⁄ 4  in), and the guard hairs on the shoulders generally do not exceed 90 mm ( 3 + 1 ⁄ 2  in), but can reach 110–130 mm ( 4 + 3 ⁄ 8 – 5 + 1 ⁄ 8  in). A wolf's coat colour is determined by its guard hairs. Wolves usually have some hairs that are white, brown, gray and black. The coat of

5429-527: The modern spotted hyena . Genomic studies suggest modern wolves and dogs descend from a common ancestral wolf population. A 2021 study found that the Himalayan wolf and the Indian plains wolf are part of a lineage that is basal to other wolves and split from them 200,000 years ago. Other wolves appear to share most of their common ancestry much more recently, within the last 23,000 years (around

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5518-593: The modern wolf and the dog are sister taxa , as modern wolves are not closely related to the population of wolves that was first domesticated . In 2019, a workshop hosted by the IUCN /Species Survival Commission's Canid Specialist Group considered the New Guinea singing dog and the dingo to be feral Canis familiaris , and therefore should not be assessed for the IUCN Red List . The phylogenetic descent of

5607-400: The months of February and May. Adoptee males may mate with an available pack female and then form their own pack. In some cases, a lone wolf is adopted into a pack to replace a deceased breeder. Wolves are territorial and generally establish territories far larger than they require to survive assuring a steady supply of prey. Territory size depends largely on the amount of prey available and

5696-958: The most frequently consumed prey of North American wolves. Wolves can digest their meal in a few hours and can feed several times in one day, making quick use of large quantities of meat. A well-fed wolf stores fat under the skin, around the heart, intestines, kidneys, and bone marrow, particularly during the autumn and winter. Nonetheless, wolves are not fussy eaters. Smaller-sized animals that may supplement their diet include rodents , hares , insectivores and smaller carnivores. They frequently eat waterfowl and their eggs. When such foods are insufficient, they prey on lizards , snakes , frogs , and large insects when available. Wolves in some areas may consume fish and even marine life. Wolves also consume some plant material. In Europe, they eat apples, pears, figs , melons, berries and cherries . In North America, wolves eat blueberries and raspberries . They also eat grass, which may provide some vitamins, but

5785-417: The neck. The hairs on the cheeks are elongated and form tufts. The ears are covered in short hairs and project from the fur. Short, elastic and closely adjacent hairs are present on the limbs from the elbows down to the calcaneal tendons . The winter fur is highly resistant to the cold. Wolves in northern climates can rest comfortably in open areas at −40 °C (−40 °F) by placing their muzzles between

5874-478: The number of worms and the age of the host. The wolf is a social animal . Its populations consist of packs and lone wolves, most lone wolves being temporarily alone while they disperse from packs to form their own or join another one. The wolf's basic social unit is the nuclear family consisting of a mated pair accompanied by their offspring. The average pack size in North America is eight wolves and 5.5 in Europe. The average pack across Eurasia consists of

5963-451: The pack for 10–54 months before dispersing. Triggers for dispersal include the onset of sexual maturity and competition within the pack for food. The distance travelled by dispersing wolves varies widely; some stay in the vicinity of the parental group, while other individuals may travel great distances of upwards of 206 km (128 mi), 390 km (240 mi), and 670 km (420 mi) from their natal (birth) packs. A new pack

6052-459: The pack usually before and after hunts, to pass on an alarm particularly at a den site, to locate each other during a storm, while crossing unfamiliar territory, and to communicate across great distances. Wolf howls can under certain conditions be heard over areas of up to 130 km (50 sq mi). Other vocalizations include growls , barks and whines. Wolves do not bark as loudly or continuously as dogs do in confrontations, rather barking

6141-466: The pads come in contact with ice and snow. In warm climates, the fur is coarser and scarcer than in northern wolves. Female wolves tend to have smoother furred limbs than males and generally develop the smoothest overall coats as they age. Older wolves generally have more white hairs on the tip of the tail, along the nose, and on the forehead. Winter fur is retained longest by lactating females, although with some hair loss around their teats. Hair length on

6230-477: The peak and the end of the Last Glacial Maximum ), originating from Siberia or Beringia . While some sources have suggested that this was a consequence of a population bottleneck , other studies have suggested that this a result of gene flow homogenising ancestry. A 2016 genomic study suggests that Old World and New World wolves split around 12,500 years ago followed by the divergence of

6319-580: The principal causes of wolf mortality, one study concluding that 14–65% of wolf deaths in Minnesota and the Denali National Park and Preserve were due to other wolves. Wolves communicate using vocalizations, body postures, scent, touch, and taste. The phases of the moon have no effect on wolf vocalization, and despite popular belief, wolves do not howl at the Moon. Wolves howl to assemble

6408-625: The range are Mount Grammos , Mount Smolikas , Mount Vasilitsa , Tymfi or Gamila , Lakmos or Peristeri , the Athamanika or Tzoumerka mountains , Vardousia , and the Agrafa mountains. Some mountains in Southern Greece are also considered part of the extended Pindus range. Its highest peak is on Mount Smolikas at an altitude of 2,637 meters. The main groups of Pindus are Epirotes , Roumeliotes , and Vlachs . There are many villages in

6497-445: The range. The Pindus region covers a wide range of elevations and habitats, from deep canyons to steep mountains. The wide range in altitude results in two major forest zones: Large breeding colonies of herons , spoonbills , egrets , and pelicans fish the waters of the mountain lakes of the Pindus. This is one of the few areas in Europe where the rare Dalmatian pelican can be found. Wolves , jackals , and bears are found in

6586-410: The rear legs and covering their faces with their tail. Wolf fur provides better insulation than dog fur and does not collect ice when warm breath is condensed against it. In cold climates, the wolf can reduce the flow of blood near its skin to conserve body heat. The warmth of the foot pads is regulated independently from the rest of the body and is maintained at just above tissue-freezing point where

6675-575: The region and connects it with the rest of Greece. Besides the imposing mountainous terrain of the range, two significant gorges in Europe are located in the area: the Vikos Gorge and the Aoos Gorge . Together with the mountain valley of Valia Kalda they have been declared protected regions and constitute the National Park of Northern Pindos. Furthermore, many mountain settlements with long history and unique architecture are located throughout

6764-474: The same extent as the coyote, whose diet contains more vegetable matter. Females tend to have narrower muzzles and foreheads, thinner necks, slightly shorter legs, and less massive shoulders than males. Adult wolves measure 105–160 cm (41–63 in) in length and 80–85 cm (31–33 in) at shoulder height. The tail measures 29–50 cm (11–20 in) in length, the ears 90–110 mm ( 3 + 1 ⁄ 2 – 4 + 3 ⁄ 8  in) in height, and

6853-552: The second half of winter and lasts for two weeks. Dens are usually constructed for pups during the summer period. When building dens, females make use of natural shelters like fissures in rocks, cliffs overhanging riverbanks and holes thickly covered by vegetation. Sometimes, the den is the appropriated burrow of smaller animals such as foxes, badgers or marmots. An appropriated den is often widened and partly remade. On rare occasions, female wolves dig burrows themselves, which are usually small and short with one to three openings. The den

6942-420: The second wife of the sebastos Constantine Keroularios , megas droungarios and nephew of the patriarch Michael Keroularios . He worked as a secretary to a provincial governor for a time and later began to earn a living by teaching and writing. He was described as vain, seems to have resented any attempt at rivalry, and violently attacked his fellow grammarians. Owing to a lack of written material, he

7031-433: The skeletons of large animals. Raised leg urination is considered to be one of the most important forms of scent communication in the wolf, making up 60–80% of all scent marks observed. Wolves are monogamous , mated pairs usually remaining together for life. Should one of the pair die, another mate is found quickly. With wolves in the wild, inbreeding does not occur where outbreeding is possible. Wolves become mature at

7120-551: The territory's periphery. Wolves tend to avoid hunting on the fringes of their range to avoid fatal confrontations with neighbouring packs. The smallest territory on record was held by a pack of six wolves in northeastern Minnesota, which occupied an estimated 33 km (13 sq mi), while the largest was held by an Alaskan pack of ten wolves encompassing 6,272 km (2,422 sq mi). Wolf packs are typically settled, and usually leave their accustomed ranges only during severe food shortages. Territorial fights are among

7209-476: The two species interact. One-to-one, hyenas dominate wolves, and may prey on them, but wolf packs can drive off single or outnumbered hyenas. There is at least one case in Israel of a hyena associating and cooperating with a wolf pack. Viral diseases carried by wolves include: rabies , canine distemper , canine parvovirus , infectious canine hepatitis , papillomatosis , and canine coronavirus . In wolves,

7298-404: The vulnerable individuals of large prey, with a pack of 15 able to bring down an adult moose . The variation in diet between wolves living on different continents is based on the variety of hoofed mammals and of available smaller and domesticated prey. In North America, the wolf's diet is dominated by wild large hoofed mammals (ungulates) and medium-sized mammals. In Asia and Europe, their diet

7387-541: The wolf because of its "cauda recurvata" (upturning tail) which is not found in any other canid . In the third edition of Mammal Species of the World published in 2005, the mammalogist W. Christopher Wozencraft listed under C. lupus 36 wild subspecies, and proposed two additional subspecies: familiaris (Linnaeus, 1758) and dingo (Meyer, 1793). Wozencraft included hallstromi —the New Guinea singing dog —as

7476-468: The wolf's intestines. Tapeworms are commonly found in wolves, which they get though their prey, and generally cause little harm in wolves, though this depends on the number and size of the parasites, and the sensitivity of the host. Symptoms often include constipation , toxic and allergic reactions , irritation of the intestinal mucosa , and malnutrition . Wolves can carry over 30 roundworm species, though most roundworm infections appear benign, depending on

7565-403: The €10 Greek Birds and Flowers and Black Pine Trees commemorative coins, minted in 2007. On the obverse of the latter there is a panoramic view of the common black pine trees that are prevalent in the park. John Tzetzes John Tzetzes ( ‹See Tfd› Greek : Ἰωάννης Τζέτζης , translit.   Iōánnēs Tzétzēs ; c.  1110 , Constantinople – 1180, Constantinople)

7654-511: Was a Byzantine poet and grammarian who lived at Constantinople in the 12th century. He is known for making significant contributions in preserving much valuable information from ancient Greek literature and scholarship. Of his numerous works, the most important one is the Book of Histories , also known as Chiliades ('Thousands'). The work is a long poem containing knowledge that is unavailable elsewhere and serves as commentary on Tzetzes's own letters. Two of his other important works are

7743-536: Was discovered through mitochondrial DNA analysis of jackals living in the Caucasus Mountains and in Bulgaria. In 2021, a genetic study found that the dog's similarity to the extant gray wolf was the result of substantial dog-into-wolf gene flow , with little evidence of the reverse. The wolf is the largest extant member of the family Canidae , and is further distinguished from coyotes and jackals by

7832-654: Was drafted in 1552 by Pahomios Rousanos  [ el ] , he wrote: “A mountain called by the Greeks Pindos is the same mountain which is called Metzovon in Barbarian” and further down the same text he adds “this mountain, Metzovon, separates the Ioannina region from the Thessaloniki region.” By the eighteenth century, there had been identification of the name Metsovo with the Pindos mountain range (in

7921-401: Was obliged to trust to his memory; therefore caution has to be exercised in reading his work. However, he was learned, and made a great contribution to the furtherance of the study of ancient Greek literature . Tzetzes published a collection of 107 of his Letters addressed partly to fictitious/unidentified personages, and partly to influential men and women of the writer's time. They contain

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