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128-628: See text Psiloceratoidea is a superfamily of Early Jurassic ammonoid cephalopods proposed by Hyatt in 1867, assigned to the order Ammonitida . They were very successful during Hettangian and Sinemurian . Last of them, family Cymbitidae and genera Hypoxynoticeras and Radstockiceras survived into Early Pliensbachian . Psiloceratoidea is probably derived from the family Ussuritidae , which were Triassic members of Phylloceratoidea . Similar to their ancestors, Psiloceratidae kept smooth, rounded venter for whole of their life. Schlotheimiidae were different, as they had ventral chevrons. Rest of

256-604: A fossil record extending back about 300 million years to the Paleozoic in the late Carboniferous period; even many of the modern genera are recognizable from fossils 60–120 million years old. Other classes and orders, now long extinct, also occur as fossils, particularly from the late Paleozoic and Mesozoic eras. Fossil conifers included many diverse forms, the most dramatically distinct from modern conifers being some herbaceous conifers with no woody stems. Major fossil orders of conifers or conifer-like plants include

384-697: A sister group to Pinales (the 'gnepine' hypothesis) or as being more derived than Pinales but sister to the rest of the group. Most recent studies favor the 'gnepine' hypothesis. The earliest conifers appear in the fossil record during the Late Carboniferous ( Pennsylvanian ), over 300 million years ago. Conifers are thought to be most closely related to the Cordaitales , a group of extinct Carboniferous-Permian trees and clambering plants whose reproductive structures had some similarities to those of conifers. The most primitive conifers belong to

512-711: A 70 km diameter impact structure buried beneath the Kalahari desert in northern South Africa. The impact is dated to the Tithonian, approximately 146.06 ± 0.16 Mya. Another major structure is the Puchezh-Katunki crater , 40 kilometres in diameter, buried beneath Nizhny Novgorod Oblast in western Russia. The impact has been dated to the Sinemurian, 195.9 ± 1.0 Ma. At the beginning of the Jurassic, all of

640-499: A dominant component of Jurassic floras. The Late Triassic and Jurassic was a major time of diversification of conifers, with most modern conifer groups appearing in the fossil record by the end of the Jurassic, having evolved from voltzialean ancestors. Araucarian conifers have their first unambiguous records during the Early Jurassic, and members of the modern genus Araucaria were widespread across both hemispheres by

768-422: A female multicellular gametophyte. The female gametophytes grow to produce two or more archegonia , each of which contains an egg. Upon fertilization, the diploid egg will give rise to the embryo, and a seed is produced. The female cone then opens, releasing the seeds which grow to a young seedling . Conifer reproduction is synchronous with seasonal changes in temperate zones. Reproductive development slows to

896-399: A female cone and are drawn into a tiny opening on the ovule called the micropyle . It is within the ovule that pollen-germination occurs. From here, a pollen tube seeks out the female gametophyte, which contains archegonia each with an egg, and if successful, fertilization occurs. The resulting zygote develops into an embryo , which along with the female gametophyte (nutritional material for

1024-665: A halt during each winter season and then resumes each spring. The male strobilus development is completed in a single year. Conifers are classified by three reproductive cycles that refer to the completion of female strobilus development from initiation to seed maturation. All three types of reproductive cycle have a long gap between pollination and fertilization . One year reproductive cycle : The genera include Abies , Picea , Cedrus , Pseudotsuga , Tsuga , Keteleeria ( Pinaceae ) and Cupressus , Thuja , Cryptomeria , Cunninghamia and Sequoia ( Cupressaceae ) . Female strobili are initiated in late summer or fall of

1152-647: A kink in one of the plate boundaries, resulting in the formation of the Pacific Plate at the centre of the junction. During the Middle to early Late Jurassic, the Sundance Seaway , a shallow epicontinental sea , covered much of northwest North America. The eustatic sea level is estimated to have been close to present levels during the Hettangian and Sinemurian, rising several tens of metres during

1280-413: A maximum of energy from weak sunshine at high latitudes or under forest canopy shade. Conifers from hotter areas with high sunlight levels (e.g. Turkish pine Pinus brutia ) often have yellower-green leaves, while others (e.g. blue spruce , Picea pungens ) may develop blue or silvery leaves to reflect ultraviolet light. In the great majority of genera the leaves are evergreen , usually remaining on

1408-880: A needle-like appearance, but others, including most of the Cupressaceae and some of the Podocarpaceae , have flat, triangular scale-like leaves. Some, notably Agathis in Araucariaceae and Nageia in Podocarpaceae, have broad, flat strap-shaped leaves. Others such as Araucaria columnaris have leaves that are awl-shaped. In the majority of conifers, the leaves are arranged spirally, the exceptions being most of Cupressaceae and one genus in Podocarpaceae, where they are arranged in decussate opposite pairs or whorls of 3 (−4). In many species with spirally arranged leaves, such as Abies grandis (pictured),

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1536-402: A protective cone called a strobilus . The cones take from four months to three years to reach maturity, and vary in size from 2 to 600 millimetres ( 1 ⁄ 8 to 23 + 5 ⁄ 8  in) long. In Pinaceae , Araucariaceae , Sciadopityaceae and most Cupressaceae , the cones are woody , and when mature the scales usually spread open allowing the seeds to fall out and be dispersed by

1664-453: A result of high ocean acidity and temperature inhibiting its mineralisation into apatite; the abundance of phosphorus in marine environments caused further eutrophication and consequent anoxia in a positive feedback loop. The end-Jurassic transition was originally considered one of eight mass extinctions, but is now considered to be a complex interval of faunal turnover, with the increase in diversity of some groups and decline in others, though

1792-487: A serious environmental issue causing problems for pastoral farming and for conservation . Radiata pine was introduced to Australia in the 1870s. It is "the dominant tree species in the Australian plantation estate" – so much so that many Australians are concerned by the resulting loss of native wildlife habitat. The species is widely regarded as an environmental weed across southeastern and southwestern Australia and

1920-438: A single extant class , Pinopsida . All extant conifers are perennial woody plants with secondary growth . The great majority are trees , though a few are shrubs . Examples include cedars , Douglas-firs , cypresses , firs , junipers , kauri , larches , pines , hemlocks , redwoods , spruces , and yews . As of 2002, Pinophyta contained seven families, 60 to 65 genera, and more than 600 living species. Although

2048-406: A single white spruce tree from 1926 to 1961. Apical growth of the stem was slow from 1926 through 1936 when the tree was competing with herbs and shrubs and probably shaded by larger trees. Lateral branches began to show reduced growth and some were no longer in evidence on the 36-year-old tree. Apical growth totaling about 340 m, 370 m, 420 m, 450 m, 500 m, 600 m, and 600 m was made by the tree in

2176-469: A useful guide by which to assess the adequacy of particular nutrients, and the ratios among the major nutrients are helpful guides to nutritional imbalances. The softwood derived from conifers is of great economic value, providing about 45% of the world's annual lumber production. Other uses of the timber include the production of paper and plastic from chemically treated wood pulp. Some conifers also provide foods such as pine nuts and juniper berries ,

2304-591: A year, then they overwinter. Female strobili emerge followed by pollination in the following spring. Fertilization takes place in summer of the following year, only 3–4 months after pollination. Cones mature and seeds are then shed by the end of that same year. Pollination and fertilization occur in a single growing season. Two-year reproductive cycle : The genera includes Widdringtonia , Sequoiadendron ( Cupressaceae ) and most species of Pinus . Female strobilus initials are formed in late summer or fall then overwinter. Female strobili emerge and receive pollen in

2432-492: Is Austrohamia minuta from the Early Jurassic (Pliensbachian) of Patagonia, known from many parts of the plant. The reproductive structures of Austrohamia have strong similarities to those of the primitive living cypress genera Taiwania and Cunninghamia . By the Middle to Late Jurassic Cupressaceae were abundant in warm temperate–tropical regions of the Northern Hemisphere, most abundantly represented by

2560-446: Is a coast redwood ( Sequoia sempervirens ), with a height of 115.55 metres (although one mountain ash, Eucalyptus regnans , allegedly grew to a height of 140 metres, the tallest living angiosperms are significantly smaller at around 100 metres. ) The thickest (that is, the tree with the greatest trunk diameter ) is a Montezuma cypress ( Taxodium mucronatum ), 11.42 metres in diameter. The largest tree by three-dimensional volume

2688-418: Is a giant sequoia ( Sequoiadendron giganteum ), with a volume 1486.9 cubic metres. The smallest is the pygmy pine ( Lepidothamnus laxifolius ) of New Zealand, which is seldom taller than 30 cm when mature. The oldest non-clonal living tree is a Great Basin bristlecone pine ( Pinus longaeva ), 4,700 years old. Since most conifers are evergreens, the leaves of many conifers are long, thin and have

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2816-494: Is divided into three epochs : Early, Middle, and Late. Similarly, in stratigraphy , the Jurassic is divided into the Lower Jurassic , Middle Jurassic , and Upper Jurassic series . Geologists divide the rocks of the Jurassic into a stratigraphic set of units called stages , each formed during corresponding time intervals called ages. Stages can be defined globally or regionally. For global stratigraphic correlation,

2944-565: Is followed by the Kimmeridgian Warm Interval (KWI) between 164 and 150 million years ago. Based on fossil wood distribution, this was one of the wettest intervals of the Jurassic. The Pangaean interior had less severe seasonal swings than in previous warm periods as the expansion of the Central Atlantic and Western Indian Ocean provided new sources of moisture. A prominent drop in temperatures occurred during

3072-470: Is fourth order, with a periodicity of approximately 410,000 years. During the Early Jurassic the world's oceans transitioned from an aragonite sea to a calcite sea chemistry, favouring the dissolution of aragonite and precipitation of calcite . The rise of calcareous plankton during the Middle Jurassic profoundly altered ocean chemistry, with the deposition of biomineralized plankton on

3200-618: Is linked to the Jura Mountains , a forested mountain range that mainly follows the France–Switzerland border . The name "Jura" is derived from the Celtic root * jor via Gaulish *iuris "wooded mountain", which was borrowed into Latin as a name of a place and evolved into Juria and finally Jura . During a tour of the region in 1795, German naturalist Alexander von Humboldt recognized carbonate deposits within

3328-475: Is marked by a globally documented high amplitude negative carbon isotope excursion, as well as the deposition of black shales and the extinction and collapse of carbonate-producing marine organisms, associated with a major rise in global temperatures. The TOAE is often attributed to the eruption of the Karoo-Ferrar large igneous provinces and the associated increase of carbon dioxide concentration in

3456-577: Is named after the city of Oxford in England and was named by Alcide d'Orbigny in 1844 in reference to the Oxford Clay . The base of the Oxfordian lacks a defined GSSP. W. J. Arkell in studies in 1939 and 1946 placed the lower boundary of the Oxfordian as the first appearance of the ammonite Quenstedtoceras mariae (then placed in the genus Vertumniceras ). Subsequent proposals have suggested

3584-402: Is probably: first to apical growth and new needle formation, then to buds for the next year's growth, with the cambium in the older parts of the branches receiving sustenance last. In the white spruce studied by Fraser et al. (1964), the needles constituted 17.5% of the over-day weight. Undoubtedly, the proportions change with time. Wind and animal dispersals are two major mechanisms involved in

3712-432: Is the whitespotted sawyer ( Monochamus scutellatus ). Adults are found in summer on newly fallen or recently felled trees chewing tiny slits in the bark in which they lay eggs. The eggs hatch in about two weeks and the tiny larvae tunnel to the wood and score its surface with their feeding channels. With the onset of cooler weather, they bore into the wood, making oval entrance holes and tunnelling deeply. Feeding continues

3840-707: Is the setting of the world's largest oil reserves, including the Ghawar Field , the world's largest oil field. The Jurassic-aged Sargelu and Naokelekan formations are major source rocks for oil in Iraq . Over 1500 gigatons of Jurassic coal reserves are found in north-west China, primarily in the Turpan-Hami Basin and the Ordos Basin . Major impact structures include the Morokweng impact structure ,

3968-622: Is used to mark the beginnings of stages, as well as smaller timespans within stages, referred to as "ammonite zones"; these, in turn, are also sometimes subdivided further into subzones. Global stratigraphy is based on standard European ammonite zones, with other regions being calibrated to the European successions. The oldest part of the Jurassic Period has historically been referred to as the Lias or Liassic, roughly equivalent in extent to

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4096-623: The Cephalotaxaceae may be better included within the Taxaceae, and some authors additionally recognize Phyllocladaceae as distinct from Podocarpaceae (in which it is included here). The family Taxodiaceae is here included in the family Cupressaceae, but was widely recognized in the past and can still be found in many field guides. A new classification and linear sequence based on molecular data can be found in an article by Christenhusz et al. The conifers are an ancient group, with

4224-583: The Cordaitales , Vojnovskyales , Voltziales and perhaps also the Czekanowskiales (possibly more closely related to the Ginkgophyta ). Pinaceae Araucariaceae Podocarpaceae Sciadopityaceae Cupressaceae Cephalotaxaceae Taxaceae Multiple studies also indicate that the Gnetophyta belong within the conifers despite their distinct appearances, either placing them as

4352-513: The International Commission on Stratigraphy (ICS) ratify global stages based on a Global Boundary Stratotype Section and Point (GSSP) from a single formation (a stratotype ) identifying the lower boundary of the stage. The ages of the Jurassic from youngest to oldest are as follows: Jurassic stratigraphy is primarily based on the use of ammonites as index fossils . The first appearance datum of specific ammonite taxa

4480-743: The Mesozoic era. Modern groups of conifers emerged from the Voltziales during the Late Permian through Jurassic . Conifers underwent a major decline in the Late Cretaceous corresponding to the explosive adaptive radiation of flowering plants . All living conifers are woody plants, and most are trees, the majority having a monopodial growth form (a single, straight trunk with side branches) with strong apical dominance . Many conifers have distinctly scented resin , secreted to protect

4608-728: The Redcar Mudstone Formation , and was ratified in 2005. The beginning of the Pliensbachian is defined by the first appearance of the ammonite Bifericeras donovani . The village Thouars (Latin: Toarcium ), just south of Saumur in the Loire Valley of France , lends its name to the Toarcian Stage. The Toarcian was named by Alcide d'Orbigny in 1842, with the original locality being Vrines quarry around 2 km northwest of Thouars. The GSSP for

4736-460: The influence of environmental conditions, their anatomical characteristics record growth rate changes produced by these changing conditions. The microscopic structure of conifer wood consists of two types of cells : parenchyma , which have an oval or polyhedral shape with approximately identical dimensions in three directions, and strongly elongated tracheids. Tracheids make up more than 90% of timber volume. The tracheids of earlywood formed at

4864-402: The wind . In some (e.g. firs and cedars ), the cones disintegrate to release the seeds, and in others (e.g. the pines that produce pine nuts ) the nut-like seeds are dispersed by birds (mainly nutcrackers , and jays ), which break up the specially adapted softer cones. Ripe cones may remain on the plant for a varied amount of time before falling to the ground; in some fire-adapted pines,

4992-409: The 16 nutrient elements known to be essential to plants, 13 of which are obtained from the soil, including nitrogen , phosphorus , potassium , calcium , magnesium , and sulfur , all used in relatively large amounts. Nutrient concentrations in conifers also vary with season, age, and kind of tissue sampled, and analytical technique. The ranges of concentrations occurring in well-grown plants provide

5120-636: The Aalenian, precessionally forced climatic changes dictated peatland wildfire magnitude and frequency. The European climate appears to have become noticeably more humid at the Aalenian-Bajocian boundary but then became more arid during the middle Bajocian. A transient ice age possibly occurred in the late Bajocian. The Callovian-Oxfordian boundary at the end of the MJCI witnessed particularly notable global cooling, potentially even an ice age. This

5248-748: The Bajocian Stage after the town of Bayeux (Latin: Bajoce ) in Normandy, France. The GSSP for the base of the Bajocian is located in the Murtinheira section at Cabo Mondego , Portugal; it was ratified in 1997. The base of the Bajocian is defined by the first appearance of the ammonite Hyperlioceras mundum . The Bathonian is named after the city of Bath , England, introduced by Belgian geologist d'Omalius d'Halloy in 1843, after an incomplete section of oolitic limestones in several quarries in

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5376-593: The Black Jurassic and the overlying clayey sandstone and ferruginous oolite of the Brown Jurassic sequences of southwestern Germany. The GSSP for the base of the Aalenian is located at Fuentelsaz in the Iberian range near Guadalajara, Spain , and was ratified in 2000. The base of the Aalenian is defined by the first appearance of the ammonite Leioceras opalinum . Alcide d'Orbigny in 1842 named

5504-560: The Black Jurassic in England by William Conybeare and William Phillips in 1822. William Phillips, the geologist, worked with William Conybeare to find out more about the Black Jurassic in England. The French palaeontologist Alcide d'Orbigny in papers between 1842 and 1852 divided the Jurassic into ten stages based on ammonite and other fossil assemblages in England and France, of which seven are still used, but none has retained its original definition. The German geologist and palaeontologist Friedrich August von Quenstedt in 1858 divided

5632-655: The Callovian–Oxfordian boundary, peaking possibly as high as 140 metres above present sea level at the Kimmeridgian–Tithonian boundary. The sea levels falls in the late Tithonian, perhaps to around 100 metres, before rebounding to around 110 metres at the Tithonian–Berriasian boundary. The sea level within the long-term trends across the Jurassic was cyclical, with 64 fluctuations, 15 of which were over 75 metres. The most noted cyclicity in Jurassic rocks

5760-464: The Early Jurassic, but also including part of the preceding Rhaetian . The Hettangian Stage was named by Swiss palaeontologist Eugène Renevier in 1864 after Hettange-Grande in north-eastern France. The GSSP for the base of the Hettangian is located at the Kuhjoch Pass, Karwendel Mountains , Northern Calcareous Alps , Austria; it was ratified in 2010. The beginning of the Hettangian, and thus

5888-519: The Greek goddess of dawn . His name was chosen by Albert Oppel for this stratigraphical stage because the Tithonian finds itself hand in hand with the dawn of the Cretaceous. The base of the Tithonian currently lacks a GSSP. The working definition for the base of the Tithonian is the first appearance of the ammonite genus Gravesia . The upper boundary of the Jurassic is currently undefined, and

6016-792: The Jura Mountains as geologically distinct from the Triassic aged Muschelkalk of southern Germany , but he erroneously concluded that they were older. He then named them Jura-Kalkstein ('Jura limestone') in 1799. In 1829, the French naturalist Alexandre Brongniart published a book entitled Description of the Terrains that Constitute the Crust of the Earth or Essay on the Structure of

6144-483: The Jurassic as a whole, is marked by the first appearance of the ammonite Psiloceras spelae tirolicum in the Kendlbach Formation exposed at Kuhjoch. The base of the Jurassic was previously defined as the first appearance of Psiloceras planorbis by Albert Oppel in 1856–58, but this was changed as the appearance was seen as too localised an event for an international boundary. The Sinemurian Stage

6272-608: The Jurassic of Asia has strap-shaped ginkgo-like leaves with highly distinct reproductive structures with similarities to those of peltasperm and corystosperm seed ferns, has been suggested to be a member of Ginkgoales sensu lato. Conifer Conifers are a group of cone-bearing seed plants , a subset of gymnosperms . Scientifically, they make up the division Pinophyta ( / p ɪ ˈ n ɒ f ɪ t ə , ˈ p aɪ n oʊ f aɪ t ə / ), also known as Coniferophyta ( / ˌ k ɒ n ɪ f ə ˈ r ɒ f ɪ t ə , - oʊ f aɪ t ə / ) or Coniferae . The division contains

6400-655: The Jurassic was marked by the major Triassic–Jurassic extinction event , associated with the eruption of the Central Atlantic Magmatic Province (CAMP). The beginning of the Toarcian Age started around 183 million years ago and is marked by the Toarcian Oceanic Anoxic Event , a global episode of oceanic anoxia , ocean acidification , and elevated global temperatures associated with extinctions, likely caused by

6528-513: The Jurassic was warmer than the present, and there were no ice caps . Forests grew close to the poles, with large arid expanses in the lower latitudes. On land, the fauna transitioned from the Triassic fauna, dominated jointly by dinosauromorph and pseudosuchian archosaurs , to one dominated by dinosaurs alone. The first stem-group birds appeared during the Jurassic, evolving from a branch of theropod dinosaurs. Other major events include

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6656-797: The Jurassic, both the North and South Pole were covered by oceans. Beginning in the Early Jurassic, the Boreal Ocean was connected to the proto-Atlantic by the "Viking corridor" or Transcontinental Laurasian Seaway, a passage between the Baltic Shield and Greenland several hundred kilometers wide. During the Callovian, the Turgai Epicontinental Sea formed, creating a marine barrier between Europe and Asia. Madagascar and Antarctica began to rift away from Africa during

6784-686: The Jurassic, the North Atlantic Ocean remained relatively narrow, while the South Atlantic did not open until the Cretaceous. The continents were surrounded by Panthalassa , with the Tethys Ocean between Gondwana and Asia. At the end of the Triassic, there was a marine transgression in Europe, flooding most parts of central and western Europe transforming it into an archipelago of islands surrounded by shallow seas. During

6912-572: The Jurassic–Cretaceous boundary is currently the only system boundary to lack a defined GSSP. Placing a GSSP for this boundary has been difficult because of the strong regionality of most biostratigraphic markers, and lack of any chemostratigraphic events, such as isotope excursions (large sudden changes in ratios of isotopes ), that could be used to define or correlate a boundary. Calpionellids , an enigmatic group of planktonic protists with urn-shaped calcitic tests briefly abundant during

7040-713: The Known Lands of the Earth. In this book, Brongniart used the phrase terrains jurassiques when correlating the "Jura-Kalkstein" of Humboldt with similarly aged oolitic limestones in Britain, thus coining and publishing the term "Jurassic". The German geologist Leopold von Buch in 1839 established the three-fold division of the Jurassic, originally named from oldest to the youngest: the Black Jurassic , Brown Jurassic , and White Jurassic . The term " Lias " had previously been used for strata of equivalent age to

7168-526: The Late Jurassic (Kimmeridgian) of Scotland, which remains the only known unequivocal fossil of the group before the Cretaceous. Despite being the earliest known member of the Pinaceae, Eathiestrobus appears to be a member of the pinoid clade of the family, suggesting that the initial diversification of Pinaceae occurred earlier than has been found in the fossil record. The earliest record of

7296-416: The Middle Jurassic. Also abundant during the Jurassic is the extinct family Cheirolepidiaceae , often recognised through their highly distinctive Classopolis pollen. Jurassic representatives include the pollen cone Classostrobus and the seed cone Pararaucaria . Araucarian and Cheirolepidiaceae conifers often occur in association. The oldest definitive record of the cypress family ( Cupressaceae )

7424-501: The Middle to Late Jurassic, corresponding to the increasing aridity of the region. Ginkgoales , of which the sole living species is Ginkgo biloba , were more diverse during the Jurassic: they were among the most important components of Eurasian Jurassic floras and were adapted to a wide variety of climatic conditions. The earliest representatives of the genus Ginkgo , represented by ovulate and pollen organs similar to those of

7552-743: The North American Forest Tree Nursery Soils Workshop at Syracuse in 1980 provided strong contrary evidence: Bob Eastman, President of the Western Maine Forest Nursery Co. stated that for 15 years he has been successful in avoiding winter “burn” to Norway spruce and white spruce in his nursery operation by fertilizing with 50–80 lb/ac (56–90 kg/ha) nitrogen in September, whereas previously winter burn had been experienced annually, often severely. Eastman also stated that

7680-653: The North Atlantic Ocean with eastern Panthalassa. Palaeontological data suggest that the seaway had been open since the Early Jurassic. As part of the Nevadan orogeny , which began during the Triassic, the Cache Creek Ocean closed, and various terranes including the large Wrangellia Terrane accreted onto the western margin of North America. By the Middle Jurassic the Siberian plate and

7808-462: The North China-Amuria block had collided, resulting in the closure of the Mongol-Okhotsk Ocean . During the Early Jurassic, around 190 million years ago, the Pacific Plate originated at the triple junction of the Farallon , Phoenix , and Izanagi tectonic plates , the three main oceanic plates of Panthalassa. The previously stable triple junction had converted to an unstable arrangement surrounded on all sides by transform faults because of

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7936-409: The Pinales without Taxales as paraphyletic , and the latter order is no longer considered distinct. A more accurate subdivision would be to split the class into three orders, Pinales containing only Pinaceae, Araucariales containing Araucariaceae and Podocarpaceae, and Cupressales containing the remaining families (including Taxaceae), but there has not been any significant support for such a split, with

8064-502: The TOAE represented one of the most severe extinctions in their evolutionary history. While the event had significant impact on marine invertebrates, it had little effect on marine reptiles. During the TOAE, the Sichuan Basin was transformed into a giant lake , probably three times the size of modern-day Lake Superior , represented by the Da'anzhai Member of the Ziliujing Formation . The lake likely sequestered ~460 gigatons (Gt) of organic carbon and ~1,200 Gt of inorganic carbon during

8192-414: The Tithonian, known as the Early Tithonian Cooling Event (ETCE). The end of the Jurassic was marked by the Tithonian–early Barremian Cool Interval (TBCI), beginning 150 million years ago and continuing into the Early Cretaceous . The Toarcian Oceanic Anoxic Event (TOAE), also known as the Jenkyns Event, was an episode of widespread oceanic anoxia during the early part of the Toarcian Age, c. 183 Mya. It

8320-529: The Toarcian around 174 million years ago. During the Toarcian Warm Interval, ocean surface temperatures likely exceeded 30 °C (86 °F), and equatorial and subtropical (30°N–30°S) regions are likely to have been extremely arid, with temperatures in the interior of Pangea likely in excess of 40 °C (104 °F).The Toarcian Warm Interval is followed by the Middle Jurassic Cool Interval (MJCI) between 174 and 164 million years ago, which may have been punctuated by brief, ephemeral icehouse intervals. During

8448-489: The appearance of the earliest crabs and modern frogs , salamanders and lizards . Mammaliaformes , one of the few cynodont lineages to survive the end of the Triassic, continued to diversify throughout the period, with the Jurassic seeing the emergence of the first crown group mammals . Crocodylomorphs made the transition from a terrestrial to an aquatic life. The oceans were inhabited by marine reptiles such as ichthyosaurs and plesiosaurs , while pterosaurs were

8576-428: The atmosphere, as well as the possible associated release of methane clathrates . This likely accelerated the hydrological cycle and increased silicate weathering , as evidenced by an increased amount of organic matter of terrestrial origin found in marine deposits during the TOAE. Groups affected include ammonites, ostracods , foraminifera , bivalves , cnidarians , and especially brachiopods , for which

8704-407: The base of the Toarcian is located at Peniche, Portugal , and was ratified in 2014. The boundary is defined by the first appearance of ammonites belonging to the subgenus Dactylioceras ( Eodactylites ) . The Aalenian is named after the city of Aalen in Germany. The Aalenian was defined by Swiss geologist Karl Mayer-Eymar in 1864. The lower boundary was originally between the dark clays of

8832-558: The beginning of a growing season have large radial sizes and smaller, thinner cell walls . Then, the first tracheids of the transition zone are formed, where the radial size of cells and the thickness of their cell walls changes considerably. Finally, latewood tracheids are formed, with small radial sizes and greater cell wall thickness. This is the basic pattern of the internal cell structure of conifer tree rings. Most conifers are monoecious , but some are subdioecious or dioecious ; all are wind-pollinated . Conifer seeds develop inside

8960-587: The cone consists of several fused scales, while in others (e.g. Taxaceae), the cone is reduced to just one seed scale or (e.g. Cephalotaxaceae) the several scales of a cone develop into individual arils, giving the appearance of a cluster of berries. The male cones have structures called microsporangia that produce yellowish pollen through meiosis. Pollen is released and carried by the wind to female cones. Pollen grains from living pinophyte species produce pollen tubes, much like those of angiosperms. The gymnosperm male gametophytes (pollen grains) are carried by wind to

9088-402: The dispersal of conifer seeds. Wind-born seed dispersal involves two processes, namely; local neighborhood dispersal and long-distance dispersal. Long-distance dispersal distances range from 11.9–33.7 kilometres (7.4–20.9 mi) from the source. Birds of the crow family, Corvidae , are the primary distributor of the conifer seeds. These birds are known to cache 32,000 pine seeds and transport

9216-447: The dominant flying vertebrates . Modern sharks and rays first appeared and diversified during the period, while the first known crown-group teleost fish appeared near the end of the period. The flora was dominated by ferns and gymnosperms , including conifers , of which many modern groups made their first appearance during the period, as well as other groups like the extinct Bennettitales . The chronostratigraphic term "Jurassic"

9344-688: The end of the Jurassic. The oldest unambiguous members of Podocarpaceae are known from the Jurassic, found across both hemispheres, including Scarburgia and Harrisiocarpus from the Middle Jurassic of England, as well as unnamed species from the Middle-Late Jurassic of Patagonia. During the Early Jurassic, the flora of the mid-latitudes of Eastern Asia were dominated by the extinct deciduous broad leafed conifer Podozamites , which appears to not be closely related to any living family of conifer. Its range extended northwards into polar latitudes of Siberia and then contracted northward in

9472-653: The end of the Triassic, the Peltaspermaceae became extinct in most parts of the world, with Lepidopteris persisting into the Early Jurassic in Patagonia. Dicroidium , a corystosperm seed fern that was a dominant part of Gondwanan floral communities during the Triassic, also declined at the Triassic–Jurassic boundary, surviving as a relict in Antarctica into the Early Jurassic. Conifers formed

9600-465: The eruption of the Karoo-Ferrar large igneous provinces . The end of the Jurassic, however, has no clear, definitive boundary with the Cretaceous and is the only boundary between geological periods to remain formally undefined. By the beginning of the Jurassic, the supercontinent Pangaea had begun rifting into two landmasses: Laurasia to the north and Gondwana to the south. The climate of

9728-468: The event. Seawater pH , which had already substantially decreased prior to the event, increased slightly during the early stages of the TOAE, before dropping to its lowest point around the middle of the event. This ocean acidification is the probable cause of the collapse of carbonate production. Additionally, anoxic conditions were exacerbated by enhanced recycling of phosphorus back into ocean water as

9856-506: The evidence for this is primarily European, probably controlled by changes in eustatic sea level. There is no evidence of a mass extinction of plants at the Triassic–Jurassic boundary. At the Triassic–Jurassic boundary in Greenland, the sporomorph (pollen and spores) record suggests a complete floral turnover. An analysis of macrofossil floral communities in Europe suggests that changes were mainly due to local ecological succession . At

9984-432: The families had angular venter, or keel for at least part of their ontogeny. Most of the members of this superfamily had only simple ribs, but few of them have evolved also secondary ribbing. While some members are involute and some Oxynoticeratidae were oxycone, most of the species were evolute. This Ammonitina -related article is a stub . You can help Misplaced Pages by expanding it . Jurassic The start of

10112-567: The female cone is termed fruit , which undergoes ripening (maturation). It was found recently that the pollen of conifers transfers the mitochondrial organelles to the embryo , a sort of meiotic drive that perhaps explains why Pinus and other conifers are so productive, and perhaps also has bearing on observed sex-ratio bias. Conifers are heterosporous , generating two different types of spores: male microspores and female megaspores . These spores develop on separate male and female sporophylls on separate male and female cones. In

10240-686: The first appearance of Cardioceras redcliffense as the lower boundary. The village of Kimmeridge on the coast of Dorset , England, is the origin of the name of the Kimmeridgian. The stage was named by Alcide d'Orbigny in 1842 in reference to the Kimmeridge Clay . The GSSP for the base of the Kimmeridgian is the Flodigarry section at Staffin Bay on the Isle of Skye , Scotland , which

10368-465: The first appearance of the ammonite Strambergella jacobi , formerly placed in the genus Berriasella , but its use as a stratigraphic indicator has been questioned, as its first appearance does not correlate with that of C. alpina . The Kimmeridge Clay and equivalents are the major source rock for the North Sea oil . The Arabian Intrashelf Basin, deposited during the Middle and Late Jurassic,

10496-449: The first year spring and become conelets. The conelet goes through another winter rest and, in the spring of the second year archegonia form in the conelet. Fertilization of the archegonia occurs by early summer of the second year, so the pollination-fertilization interval exceeds a year. After fertilization, the conelet is considered an immature cone. Maturation occurs by autumn of the second year, at which time seeds are shed. In summary,

10624-466: The following spring. Female strobili emerge then pollination occurs in spring of the second year then the pollinated strobili become conelets in the same year (i.e. the second year). The female gametophytes in the conelet develop so slowly that the megaspore does not go through free-nuclear divisions until autumn of the third year. The conelet then overwinters again in the free-nuclear female gametophyte stage. Fertilization takes place by early summer of

10752-492: The following summer when larvae occasionally return to the surface of the wood and extend the feeding channels generally in a U-shaped configuration. During this time, small piles of frass extruded by the larvae accumulate under logs. Early in the spring of the second year following egg-laying, the larvae, about 30 mm long, pupate in the tunnel enlargement just below the wood surface. The resulting adults chew their way out in early summer, leaving round exit holes, so completing

10880-399: The fourth year and seeds mature in the cones by autumn of the fourth year. The growth and form of a forest tree are the result of activity in the primary and secondary meristems , influenced by the distribution of photosynthate from its needles and the hormonal gradients controlled by the apical meristems. External factors also influence growth and form. Fraser recorded the development of

11008-402: The genus Elatides . The Jurassic also saw the first appearances of some modern genera of cypresses, such as Sequoia . Members of the extinct genus Schizolepidopsis which likely represent a stem-group to the pine family ( Pinaceae ), were widely distributed across Eurasia during the Jurassic. The oldest unambiguous record of Pinaceae is the pine cone Eathiestrobus , known from

11136-419: The growing embryo) and its surrounding integument, becomes a seed . Eventually, the seed may fall to the ground and, if conditions permit, grow into a new plant. In forestry , the terminology of flowering plants has commonly though inaccurately been applied to cone-bearing trees as well. The male cone and unfertilized female cone are called male flower and female flower , respectively. After fertilization,

11264-530: The immense conifer forests of the world represent the largest terrestrial carbon sink . Conifers are of great economic value for softwood lumber and paper production. Conifer is a Latin word, a compound of conus (cone) and ferre (to bear), meaning "the one that bears (a) cone(s)". The division name Pinophyta conforms to the rules of the International Code of Nomenclature for algae, fungi, and plants (ICN), which state (Article 16.1) that

11392-535: The largest and economically most important component group of gymnosperms, but nevertheless they comprise only one of the four groups. The division Pinophyta consists of just one class, Pinopsida, which includes both living and fossil taxa. Subdivision of the living conifers into two or more orders has been proposed from time to time. The most commonly seen in the past was a split into two orders, Taxales (Taxaceae only) and Pinales (the rest), but recent research into DNA sequences suggests that this interpretation leaves

11520-545: The late Early Jurassic in association with the eruption of the Karoo-Ferrar large igneous provinces , opening the western Indian Ocean and beginning the fragmentation of Gondwana. At the beginning of the Jurassic, North and South America remained connected, but by the beginning of the Late Jurassic they had rifted apart to form the Caribbean Seaway, also known as the Hispanic Corridor, which connected

11648-472: The late Sinemurian–Pliensbachian before regressing to near present levels by the late Pliensbachian. There seems to have been a gradual rise to a peak of ~75 m above present sea level during the Toarcian. During the latest part of the Toarcian, the sea level again dropped by several tens of metres. It progressively rose from the Aalenian onwards, aside from dips of a few tens of metres in the Bajocian and around

11776-403: The latest Jurassic to earliest Cretaceous, have been suggested to represent the most promising candidates for fixing the Jurassic–Cretaceous boundary In particular, the first appearance Calpionella alpina , co-inciding with the base of the eponymous Alpina subzone, has been proposed as the definition of the base of the Cretaceous. The working definition for the boundary has often been placed as

11904-417: The leaf bases are twisted to present the leaves in a very flat plane for maximum light capture. Leaf size varies from 2 mm in many scale-leaved species, up to 400 mm long in the needles of some pines (e.g. Apache pine, Pinus engelmannii ). The stomata are in lines or patches on the leaves and can be closed when it is very dry or cold. The leaves are often dark green in colour, which may help absorb

12032-402: The majority of opinion preferring retention of all the families within a single order Pinales, despite their antiquity and diverse morphology . There were seven families of conifers c.  2011 , with 65–70 genera and over 600 living species ( c.  2002 ). The seven most distinct families are linked in the box above right and phylogenetic diagram left. In other interpretations,

12160-403: The male cones, microspores are produced from microsporocytes by meiosis . The microspores develop into pollen grains, which contain the male gametophytes. Large amounts of pollen are released and carried by the wind. Some pollen grains will land on a female cone for pollination. The generative cell in the pollen grain divides into two haploid sperm cells by mitosis leading to the development of

12288-428: The modern species, are known from the Middle Jurassic in the Northern Hemisphere. Several other lineages of ginkgoaleans are known from Jurassic rocks, including Yimaia , Grenana , Nagrenia and Karkenia . These lineages are associated with Ginkgo- like leaves, but are distinguished from living and fossil representatives of Ginkgo by having differently arranged reproductive structures. Umaltolepis from

12416-466: The names of higher taxa in plants (above the rank of family) are either formed from the name of an included family (usually the most common and/or representative), in this case Pinaceae (the pine family), or are descriptive. A descriptive name in widespread use for the conifers (at whatever rank is chosen) is Coniferae (Art 16 Ex 2). According to the ICN, it is possible to use a name formed by replacing

12544-422: The ocean floor acting as a buffer against large CO 2 emissions. The climate of the Jurassic was generally warmer than that of present, by around 5–10 °C (9–18 °F), with atmospheric carbon dioxide likely about four times higher. Intermittent "cold snap" intervals are known to have occurred during this time period, however, interrupting the otherwise warm greenhouse climate. Forests likely grew near

12672-460: The oceans, resulting in large areas of desert and scrubland in the lower latitudes between 40° N and S of the equator. Tropical rainforest and tundra biomes are likely to have been rare or absent. The Jurassic also witnessed the decline of the Pangaean megamonsoon that had characterised the preceding Permian and Triassic periods. Variation in the frequency of wildfire activity in the Jurassic

12800-486: The one 11 m tall white spruce, Fraser et al. (1964) speculated that if the photosynthate used in making apical growth in 1961 was manufactured the previous year, then the 4 million needles that were produced up to 1960 manufactured food for about 600,000 mm of apical growth or 730 g dry weight, over 12 million mm of wood for the 1961 annual ring, plus 1 million new needles, in addition to new tissue in branches, bark, and roots in 1960. Added to this would be

12928-424: The one-year and the two-year cycles differ mainly in the duration of the pollination-fertilization interval. Three-year reproductive cycle : Three of the conifer species are pine species ( Pinus pinea , Pinus leiophylla , Pinus torreyana ) which have pollination and fertilization events separated by a two-year interval. Female strobili initiated during late summer or autumn of a year, then overwinter until

13056-425: The overwintering storage capacity of stock thus treated was much improved (Eastman 1980). The concentrations of nutrients in plant tissues depend on many factors, including growing conditions. Interpretation of concentrations determined by analysis is easy only when a nutrient occurs in excessively low or occasionally excessively high concentration. Values are influenced by environmental factors and interactions among

13184-569: The paraphyletic assemblage of " walchian conifers ", which were small trees, and probably originated in dry upland habitats. The range of conifers expanded during the Early Permian ( Cisuralian ) to lowlands due to increasing aridity. Walchian conifers were gradually replaced by more advanced voltzialean or "transition" conifers. Conifers were largely unaffected by the Permian–Triassic extinction event , and were dominant land plants of

13312-429: The photosynthate to produce energy to sustain respiration over this period, an amount estimated to be about 10% of the total annual photosynthate production of a young healthy tree. On this basis, one needle produced food for about 0.19 mg dry weight of apical growth, 3 mm wood, one-quarter of a new needle, plus an unknown amount of branch wood, bark and roots. The order of priority of photosynthate distribution

13440-505: The plant for several (2–40) years before falling, but five genera ( Larix , Pseudolarix , Glyptostrobus , Metasequoia and Taxodium ) are deciduous , shedding their leaves in autumn. The seedlings of many conifers, including most of the Cupressaceae, and Pinus in Pinaceae, have a distinct juvenile foliage period where the leaves are different, often markedly so, from the typical adult leaves. Tree rings are records of

13568-611: The poles, where they experienced warm summers and cold, sometimes snowy winters; there were unlikely to have been ice sheets given the high summer temperatures that prevented the accumulation of snow, though there may have been mountain glaciers. Dropstones and glendonites in northeastern Siberia during the Early to Middle Jurassic indicate cold winters. The ocean depths were likely 8 °C (14 °F) warmer than present, and coral reefs grew 10° of latitude further north and south. The Intertropical Convergence Zone likely existed over

13696-416: The pollen tube. At fertilization, one of the sperm cells unites its haploid nucleus with the haploid nucleus of an egg cell. The female cone develops two ovules, each of which contains haploid megaspores. A megasporocyte is divided by meiosis in each ovule. Each winged pollen grain is a four celled male gametophyte . Three of the four cells break down leaving only a single surviving cell which will develop into

13824-648: The region. The GSSP for the base of the Bathonian is Ravin du Bès, Bas-Auran area, Alpes de Haute Provence , France; it was ratified in 2009. The base of the Bathonian is defined by the first appearance of the ammonite Gonolkites convergens , at the base of the Zigzagiceras zigzag ammonite zone . The Callovian is derived from the Latinized name of the village of Kellaways in Wiltshire , England, and

13952-514: The removal of individual plants beyond plantations is encouraged. At least 20 species of roundheaded borers of the family Cerambycidae feed on the wood of spruce , fir , and hemlock (Rose and Lindquist 1985). Borers rarely bore tunnels in living trees, although when populations are high, adult beetles feed on tender twig bark, and may damage young living trees. One of the most common and widely distributed borer species in North America

14080-453: The same effect in 105-day-old white spruce. The general short-term effect of nitrogen fertilization on coniferous seedlings is to stimulate shoot growth more so than root growth (Armson and Carman 1961). Over a longer period, root growth is also stimulated. Many nursery managers were long reluctant to apply nitrogenous fertilizers late in the growing season, for fear of increased danger of frost damage to succulent tissues. A presentation at

14208-927: The seeds as far as 12–22 km (7.5–13.7 mi) from the source. The birds store the seeds in the soil at depths of 2–3 cm ( 3 ⁄ 4 – 1 + 1 ⁄ 4  in) under conditions which favor germination . Conifers are the dominant plants over large areas of land, most notably the taiga of the Northern Hemisphere , but also in similar cool climates in mountains further south. A number of conifers originally introduced for forestry have become invasive species in parts of New Zealand , including radiata pine ( Pinus radiata ), lodgepole pine ( P. contorta ), Douglas fir ( Pseudotsuga mensiezii ) and European larch ( Larix decidua ). In parts of South Africa , maritime pine ( Pinus pinaster ), patula pine ( P. patula ) and radiata pine have been declared invasive species. These wilding conifers are

14336-545: The seeds may be stored in closed cones for up to 60–80 years, being released only when a fire kills the parent tree. In the families Podocarpaceae , Cephalotaxaceae , Taxaceae , and one Cupressaceae genus ( Juniperus ), the scales are soft, fleshy, sweet, and brightly colored, and are eaten by fruit-eating birds, which then pass the seeds in their droppings. These fleshy scales are (except in Juniperus ) known as arils . In some of these conifers (e.g. most Podocarpaceae),

14464-429: The sole source of nitrogen guanidine compounds were less prominent. Durzan and Steward noted that their results, drawn from determinations made in late summer, did not rule out the occurrence of different interim responses at other times of the year. Ammonium nitrogen produced significantly heavier (dry weight) seedlings with a higher nitrogen content after 5 weeks than did the same amount of nitrate nitrogen. Swan found

14592-517: The termination -aceae in the name of an included family, in this case preferably Pinaceae , by the appropriate termination, in the case of this division -ophyta . Alternatively, " descriptive botanical names " may also be used at any rank above family. Both are allowed. This means that if conifers are considered a division, they may be called Pinophyta or Coniferae. As a class, they may be called Pinopsida or Coniferae. As an order they may be called Pinales or Coniferae or Coniferales . Conifers are

14720-573: The three series of von Buch in the Swabian Jura into six subdivisions defined by ammonites and other fossils. The German palaeontologist Albert Oppel in his studies between 1856 and 1858 altered d'Orbigny's original scheme and further subdivided the stages into biostratigraphic zones, based primarily on ammonites. Most of the modern stages of the Jurassic were formalized at the Colloque du Jurassique à Luxembourg in 1962. The Jurassic Period

14848-570: The total number of species is relatively small, conifers are ecologically important. They are the dominant plants over large areas of land, most notably the taiga of the Northern Hemisphere , but also in similar cool climates in mountains further south. Boreal conifers have many wintertime adaptations. The narrow conical shape of northern conifers, and their downward-drooping limbs, help them shed snow. Many of them seasonally alter their biochemistry to make them more resistant to freezing. While tropical rainforests have more biodiversity and turnover,

14976-475: The tree against insect infestation and fungal infection of wounds. Fossilized resin hardens into amber , which has been commercially exploited historically (for example, in New Zealand's 19th-century kauri gum industry). The size of mature conifers varies from less than one metre to over 100 metres in height. The world's tallest, thickest, largest, and oldest living trees are all conifers. The tallest

15104-416: The usual 2-year life cycle. Conifers – notably Abies (fir), Cedrus , Chamaecyparis lawsoniana (Lawson's cypress), Cupressus (cypress), juniper , Picea (spruce), Pinus (pine), Taxus (yew), Thuja (cedar) – have been the subject of selection for ornamental purposes. Plants with unusual growth habits, sizes, and colours are propagated and planted in parks and gardens throughout

15232-407: The world's major landmasses were coalesced into the supercontinent Pangaea , which during the Early Jurassic began to break up into northern supercontinent Laurasia and the southern supercontinent Gondwana . The rifting between North America and Africa was the first to initiate, beginning in the early Jurassic, associated with the emplacement of the Central Atlantic Magmatic Province . During

15360-468: The world. Conifers can absorb nitrogen in either the ammonium (NH 4 ) or nitrate (NO 3 ) form, but the forms are not physiologically equivalent. Form of nitrogen affected both the total amount and relative composition of the soluble nitrogen in white spruce tissues (Durzan and Steward). Ammonium nitrogen was shown to foster arginine and amides and lead to a large increase of free guanidine compounds, whereas in leaves nourished by nitrate as

15488-403: The years 1955 through 1961, respectively. The total number of needles of all ages present on the 36-year-old tree in 1961 was 5.25 million weighing 14.25 kg. In 1961, needles as old as 13 years remained on the tree. The ash weight of needles increased progressively with age from about 4% in first-year needles in 1961 to about 8% in needles 10 years old. In discussing the data obtained from

15616-585: The yew family ( Taxaceae ) is Palaeotaxus rediviva , from the Hettangian of Sweden, suggested to be closely related to the living Austrotaxus , while Marskea jurassica from the Middle Jurassic of Yorkshire, England and material from the Callovian–Oxfordian Daohugou Bed in China are thought to be closely related to Amentotaxus , with the latter material assigned to the modern genus, indicating that Taxaceae had substantially diversified by

15744-474: Was first defined and introduced into scientific literature by Alcide d'Orbigny in 1842. It takes its name from the French town of Semur-en-Auxois , near Dijon . The original definition of Sinemurian included what is now the Hettangian. The GSSP of the Sinemurian is located at a cliff face north of the hamlet of East Quantoxhead , 6 kilometres east of Watchet , Somerset , England , within the Blue Lias , and

15872-569: Was governed by the 405 kyr eccentricity cycle. Thanks to the breakup of Pangaea, the hydrological cycle during the Jurassic was significantly enhanced. The beginning of the Jurassic was likely marked by a thermal spike corresponding to the Triassic–Jurassic extinction and eruption of the Central Atlantic magmatic province. The first part of the Jurassic was marked by the Early Jurassic Cool Interval between 199 and 183 million years ago. It has been proposed that glaciation

16000-582: Was named by Alcide d'Orbigny in 1852, originally the base at the contact between the Forest Marble Formation and the Cornbrash Formation . However, this boundary was later found to be within the upper part of the Bathonian. The base of the Callovian does not yet have a certified GSSP. The working definition for the base of the Callovian is the first appearance of ammonites belonging to the genus Kepplerites . The Oxfordian

16128-490: Was present in the Northern Hemisphere during both the early Pliensbachian and the latest Pliensbachian. There was a spike in global temperatures of around 4–8 °C (7–14 °F) during the early part of the Toarcian corresponding to the Toarcian Oceanic Anoxic Event and the eruption of the Karoo-Ferrar large igneous provinces in southern Gondwana, with the warm interval extending to the end of

16256-687: Was ratified in 2000. The beginning of the Sinemurian is defined by the first appearance of the ammonite Vermiceras quantoxense . Albert Oppel in 1858 named the Pliensbachian Stage after the hamlet of Pliensbach in the community of Zell unter Aichelberg in the Swabian Alb , near Stuttgart , Germany. The GSSP for the base of the Pliensbachian is found at the Wine Haven locality in Robin Hood's Bay , Yorkshire , England, in

16384-430: Was ratified in 2021. The boundary is defined by the first appearance of ammonites marking the boreal Bauhini Zone and the subboreal Baylei Zone. The Tithonian was introduced in scientific literature by Albert Oppel in 1865. The name Tithonian is unusual in geological stage names because it is derived from Greek mythology rather than a place name. Tithonus was the son of Laomedon of Troy and fell in love with Eos ,

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