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Rhipidistia

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63-476: Rhipidistia , also known as Dipnotetrapodomorpha , is a clade of lobe-finned fishes which includes the tetrapods and lungfishes . Rhipidistia formerly referred to a subgroup of Sarcopterygii consisting of the Porolepiformes and Osteolepiformes , a definition that is now obsolete. However, as cladistic understanding of the vertebrates has improved over the last few decades, a monophyletic Rhipidistia

126-444: A prehistoric lobe-finned fish is a stub . You can help Misplaced Pages by expanding it . Lobe-finned fish Sarcopterygii ( / ˌ s ɑːr k ɒ p t ə ˈ r ɪ dʒ i . aɪ / ; from Ancient Greek σάρξ (sárx)  'flesh' and πτέρυξ (ptérux)  'wing, fin') — sometimes considered synonymous with Crossopterygii (from Ancient Greek κροσσός (krossós)  'fringe') —

189-592: A common impurity. Tin ores are rare, as reflected in the fact there were no tin bronzes in Western Asia before 3000 BCE. The Bronze Age forms part of the three-age system for prehistoric societies. In this system, it follows the Neolithic in some areas of the world. While copper is a common ore, deposits of tin are rare in the Old World , and often had to be traded or carried considerable distances from

252-469: A hinge line, but this is lost in tetrapods and lungfish. Early sarcopterygians commonly exhibit a symmetrical tail, while all sarcopterygians possess teeth that are coated with genuine enamel . Most species of lobe-finned fishes are extinct. The largest known lobe-finned fish was Rhizodus hibberti from the Carboniferous period of Scotland which may have exceeded 7 meters in length. Among

315-674: A much more evident Mesolithic era, lasting millennia. In Northern Europe , societies were able to live well on rich food supplies from the marshlands fostered by the warmer climate. Such conditions produced distinctive human behaviours that are preserved in the material record, such as the Maglemosian and Azilian cultures. These conditions also delayed the coming of the Neolithic until as late as 4000 BCE (6,000  BP ) in northern Europe. Remains from this period are few and far between, often limited to middens . In forested areas,

378-472: A result, the phrase "lobe-finned fish" normally refers to not the entire clade but only aquatic members that are not tetrapods, i.e. a paraphyletic group. Non-tetrapod sarcopterygians were once the dominant predators of freshwater ecosystems during the Carboniferous and Permian periods, but suffered significant decline after the Great Dying . The only known extant non-tetrapod sarcopterygians are

441-654: A single room. Settlements might have a surrounding stone wall to keep domesticated animals in and hostile tribes out. Later settlements have rectangular mud-brick houses where the family lived in single or multiple rooms. Burial findings suggest an ancestor cult with preserved skulls of the dead. The Vinča culture may have created the earliest system of writing. The megalithic temple complexes of Ġgantija are notable for their gigantic structures. Although some late Eurasian Neolithic societies formed complex stratified chiefdoms or even states, states evolved in Eurasia only with

504-749: A single source. The emergence of metallurgy may have occurred first in the Fertile Crescent , where it gave rise to the Bronze Age in the 4th millennium BCE (the traditional view), although finds from the Vinča culture in Europe have now been securely dated to slightly earlier than those of the Fertile Crescent. Timna Valley contains evidence of copper mining 7,000 years ago. The process of transition from Neolithic to Chalcolithic in

567-570: A synthesis of rank-based Linnaean taxonomy and also reflects evolutionary relationships. Benton included the Superclass Tetrapoda in the Subclass Sarcopterygii in order to reflect the direct descent of tetrapods from lobe-finned fish, despite the former being assigned a higher taxonomic rank. Lobe-finned fishes and their sister group, the ray-finned fishes , make up the superclass Osteichthyes , characterized by

630-514: Is a clade (traditionally a class or subclass ) of vertebrate animals which includes a group of bony fish commonly referred to as lobe-finned fish . These vertebrates are characterised by prominent muscular limb buds (lobes) within their fins , which are supported by articulated appendicular skeletons . This is in contrast to the other clade of bony fish, the Actinopterygii , which have only skin -covered bony spines supporting

693-800: Is also a transition period between Stone Age and Bronze Age, the Chalcolithic or Copper Age. For the prehistory of the Americas see Pre-Columbian era . The notion of "prehistory" emerged during the Enlightenment in the work of antiquarians who used the word "primitive" to describe societies that existed before written records. The word "prehistory" first appeared in English in 1836 in the Foreign Quarterly Review . The geologic time scale for pre-human time periods, and

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756-516: Is an example. In archaeology, the Iron Age refers to the advent of ferrous metallurgy . The adoption of iron coincided with other changes, often including more sophisticated agricultural practices, religious beliefs and artistic styles, which makes the archaeological Iron Age coincide with the " Axial Age " in the history of philosophy. Although iron ore is common, the metalworking techniques necessary to use iron are different from those needed for

819-488: Is anonymous. Because of this, reference terms that prehistorians use, such as " Neanderthal " or " Iron Age ", are modern labels with definitions sometimes subject to debate. The concept of a "Stone Age" is found useful in the archaeology of most of the world, although in the archaeology of the Americas it is called by different names and begins with a Lithic stage , or sometimes Paleo-Indian . The sub-divisions described below are used for Eurasia, and not consistently across

882-782: Is called the Lower Paleolithic (as in excavations it appears underneath the Upper Paleolithic), beginning with the earliest stone tools dated to around 3.3 million years ago at the Lomekwi site in Kenya. These tools predate the genus Homo and were probably used by Kenyanthropus . Evidence of control of fire by early hominins during the Lower Palaeolithic Era is uncertain and has at best limited scholarly support. The most widely accepted claim

945-412: Is from Greek : δίπνοος , romanized :  dipnoos , lit.   'with two breathing structures' (from δι- meaning "twice", and πνοή meaning "breathing, breath"); from Ancient Greek: τετρα- , romanized:  tetra- , lit.   'four', the combining form of the numeral τέτταρες ( tettares ); from Ancient Greek: -ποδ- , romanized:  -pod- ,

1008-479: Is now understood to include the whole of Tetrapoda and the lungfishes . Rhipidistia includes Porolepiformes and Dipnoi. Extensive fossilization of lungfishes has contributed to many evolutionary studies of this group. Evolution of autostylic jaw suspension, in which the palatoquadrate bone fuses to the cranium, and the lymph pumping " lymph heart " (later lost in mammals and flying birds ), are unique to this group. Another feature shared by lungfish and tetrapods

1071-558: Is provided by a wide variety of natural and social sciences, such as anthropology , archaeology , archaeoastronomy , comparative linguistics , biology , geology , molecular genetics , paleontology , palynology , physical anthropology , and many others. Human prehistory differs from history not only in terms of its chronology , but in the way it deals with the activities of archaeological cultures rather than named nations or individuals . Restricted to material processes, remains, and artefacts rather than written records, prehistory

1134-551: Is seen as a transition period between the Stone Age and Bronze Age. An archaeological site in Serbia contains the oldest securely dated evidence of copper making at high temperature, from 7,500 years ago. The find in 2010 extends the known record of copper smelting by about 800 years, and suggests that copper smelting may have been invented independently in separate parts of Asia and Europe at that time, rather than spreading from

1197-500: Is that H. erectus or H. ergaster made fires between 790,000 and 690,000 BP in a site at Bnot Ya'akov Bridge , Israel . The use of fire enabled early humans to cook food, provide warmth, have a light source, deter animals at night and meditate. Early Homo sapiens originated some 300,000 years ago, ushering in the Middle Palaeolithic . Anatomic changes indicating modern language capacity also arise during

1260-446: Is the divided atrium . The precise time at which the choana of tetrapods evolved is debated, with some considering early rhipidistians as the first choanates. The feature is also present in modern lungfish but is probably a case of convergent evolution . The basal stem-lungfish Diabolepis did not possess it. Instead, it had four nostrils (two anterior and two posterior) like most fish. However, its posterior nares are very close to

1323-489: Is the period of human history between the first known use of stone tools by hominins c.  3.3   million years ago and the beginning of recorded history with the invention of writing systems . The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared c.  5,200 years ago. It took thousands of years for writing systems to be widely adopted, with writing having spread to almost all cultures by

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1386-516: Is when the first signs of human presence have been found; however, Africa and Asia contain sites dated as early as c.  2.5 and 1.8 million years ago, respectively. Depending on the date when relevant records become a useful academic resource, its end date also varies. For example, in Egypt it is generally accepted that prehistory ended around 3100 BCE, whereas in New Guinea

1449-708: The Copper Age or Bronze Age ; or, in some geographical regions, in the Iron Age ). The term Neolithic is commonly used in the Old World ; its application to cultures in the Americas and Oceania is complicated by the fact standard progression from stone to metal tools, as seen in the Old World, does not neatly apply. Early Neolithic farming was limited to a narrow range of plants, both wild and domesticated, which included einkorn wheat , millet and spelt , and

1512-531: The Paleolithic , by the Neolithic only Homo sapiens sapiens remained. This was a period of technological and social developments which established most of the basic elements of historical cultures, such as the domestication of crops and animals , and the establishment of permanent settlements and early chiefdoms. The era commenced with the beginning of farming , which produced the " Neolithic Revolution ". It ended when metal tools became widespread (in

1575-513: The Permian periods. There are three major hypotheses as to how lungfish evolved their stubby fins (proto-limbs). The first tetrapodomorphs, which included the gigantic rhizodonts , had the same general anatomy as the lungfish, who were their closest kin, but they appear not to have left their water habitat until the late Devonian epoch (385–359 Ma), with the appearance of tetrapods (four-legged vertebrates). Tetrapods and megalichthyids are

1638-457: The lungfish and the tetrapodomorphs , and both of them evolved their swim bladders into air-breathing lungs. Lungfish radiated into their greatest diversity during the Triassic period; today fewer than a dozen genera remain, having evolved the first proto-lungs and proto-limbs, adapting to living outside a submerged water environment by the middle Devonian (397–385 Ma). The tetrapodomorphs, on

1701-606: The three-age system for human prehistory, were systematised during the nineteenth century in the work of British, French, German, and Scandinavian anthropologists , archaeologists , and antiquarians . The main source of information for prehistory is archaeology (a branch of anthropology), but some scholars are beginning to make more use of evidence from the natural and social sciences. The primary researchers into human prehistory are archaeologists and physical anthropologists who use excavation, geologic and geographic surveys, and other scientific analysis to reveal and interpret

1764-475: The "Chalcolithic", "Eneolithic", or "Copper Age" refers to a transitional period where early copper metallurgy appeared alongside the widespread use of stone tools. During this period, some weapons and tools were made of copper. This period was still largely Neolithic in character. It is a phase of the Bronze Age before it was discovered that adding tin to copper formed the harder bronze . The Copper Age

1827-623: The 19th century. The end of prehistory therefore came at different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age , Sumer in Mesopotamia , the Indus Valley Civilisation , and ancient Egypt were the first civilizations to develop their own scripts and keep historical records, with their neighbours following. Most other civilizations reached their end of prehistory during

1890-623: The Americas, these areas did not develop complex writing systems before the arrival of Eurasians, so their prehistory reaches into relatively recent periods; for example, 1788 is usually taken as the end of the prehistory of Australia . The period when a culture is written about by others, but has not developed its own writing system, is often known as the protohistory of the culture. By definition, there are no written records from human prehistory, which can only be known from material archaeological and anthropological evidence: prehistoric materials and human remains. These were at first understood by

1953-705: The Azilian cultures, before spreading to Europe through the Iberomaurusian culture of Northern Africa and the Kebaran culture of the Levant . However, independent discovery is not ruled out. "Neolithic" means "New Stone Age", from about 10,200 BCE in some parts of the Middle East, but later in other parts of the world, and ended between 4,500 and 2,000 BCE. Although there were several species of humans during

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2016-817: The Early Devonian (416–397 Ma), the sarcopterygians, or lobe-finned fishes, split into two main lineages: the coelacanths and the rhipidistians . Coelacanths never left the oceans and their heyday was the late Devonian and Carboniferous , from 385 to 299 Ma, as they were more common during those periods than in any other period in the Phanerozoic . Actinistians, a group within the lobe-finned fish, have been around for almost 380 million years. Over time, researchers have identified 121 species spread across 47 genera. Some species are well-documented in their evolutionary placement, while others are harder to track.The greatest boom in actinistian diversity happened during

2079-559: The Early Triassic, just after the Great Dying. Coelacanths of the genus Latimeria still live today in the open oceans and retained many primordial features of ancient sarcopterygians, earning them a reputation as living fossils. The Rhipidistians, whose ancestors probably lived in the oceans near river mouths and estuaries , left the marine world and migrated into freshwater habitats. They then split into two major groups:

2142-460: The Iron Age, often through conquest by empires, which continued to expand during this period. For example, in most of Europe conquest by the Roman Empire means the term Iron Age is replaced by "Roman", " Gallo-Roman ", and similar terms after the conquest. Even before conquest, many areas began to have a protohistory, as they were written about by literate cultures; the protohistory of Ireland

2205-694: The Middle East is characterized in archaeological stone tool assemblages by a decline in high quality raw material procurement and use. North Africa and the Nile Valley imported its iron technology from the Near East and followed the Near Eastern course of Bronze Age and Iron Age development. The Bronze Age is the earliest period in which some civilizations reached the end of prehistory, by introducing written records. The Bronze Age, or parts thereof, are thus considered to be part of prehistory only for

2268-739: The Middle Palaeolithic. During the Middle Palaeolithic Era, there is the first definitive evidence of human use of fire. Sites in Zambia have charred logs, charcoal and carbonized plants, that have been dated to 180,000 BP. The systematic burial of the dead , music , prehistoric art , and the use of increasingly sophisticated multi-part tools are highlights of the Middle Paleolithic. The Upper Paleolithic extends from 50,000 and 12,000 years ago, with

2331-569: The case of Indigenous Australian "highways" known as songlines . The Mesolithic, or Middle Stone Age (from the Greek mesos , 'middle', and lithos , 'stone'), was a period in the development of human technology between the Palaeolithic and Neolithic . The Mesolithic period began with the retreat of glaciers at the end of the Pleistocene epoch, some 10,000 BP, and ended with

2394-735: The coelacanths display a strong resemblance to the presumed ancestral form of tetrapod limbs. Lobe-finned fishes seemingly underwent two distinct evolutionary paths, leading to their classification into two subclasses: the Rhipidistia (comprising the Dipnoi, or lungfish , and the Tetrapodomorpha , which includes the Tetrapoda) and the Actinistia (represented by coelacanths). The classification below follows Benton (2004), and uses

2457-463: The collection of folklore and by analogy with pre-literate societies observed in modern times. The key step to understanding prehistoric evidence is dating, and reliable dating techniques have developed steadily since the nineteenth century. The most common of these dating techniques is radiocarbon dating . Further evidence has come from the reconstruction of ancient spoken languages . More recent techniques include forensic chemical analysis to reveal

2520-946: The combining form of πούς ( pous ) meaning "foot"; and from Ancient Greek: -μορφος , romanized:  -morphos , the combining form of μορφή ( morph ) meaning "physical shape". The cladogram presented below is based on studies compiled by Philippe Janvier and others for the Tree of Life Web Project , and Swartz 2012. † Onychodontidae Actinistia (coelacanths) † Styloichthys † Porolepiformes Dipnoi (lungfishes) ?† Tungsenia † Kenichthys † Rhizodontiformes ?† Thysanolepidae † Canowindridae † Osteolepiformes † Tristichopteridae † Tinirau † Platycephalichthys † Panderichthys † Elpistostegidae † Elginerpetontidae † Metaxygnathus † Ventastega Tetrapoda [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] This article about

2583-506: The end of the Bronze Age large states, whose armies imposed themselves on people with a different culture, and are often called empires, had arisen in Egypt, China, Anatolia (the Hittites ), and Mesopotamia , all of them literate. The Iron Age is not part of prehistory for all civilizations who had introduced written records during the Bronze Age. Most remaining civilizations did so during

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2646-639: The end of the prehistoric era is set much more recently, in the 1870s, when the Russian anthropologist Nicholai Miklukho-Maklai spent several years living among native peoples, and described their way of life in a comprehensive treatise. In Europe the relatively well-documented classical cultures of Ancient Greece and Ancient Rome had neighbouring cultures, including the Celts and the Etruscans , with little writing. Historians debate how much weight to give to

2709-419: The evolutionary shift from aquatic to terrestrial existence. Pectoral and pelvic fins have articulations resembling those of tetrapod limbs. The first tetrapod land vertebrates, basal amphibian organisms, possessed legs derived from these fins. Sarcopterygians also possess two dorsal fins with separate bases, as opposed to the single dorsal fin in ray-finned fish. The braincase of sarcopterygians primitively has

2772-446: The few mines, stimulating the creation of extensive trading routes. In many areas as far apart as China and England, the valuable new material was used for weapons, but for a long time apparently not available for agricultural tools. Much of it seems to have been hoarded by social elites, and sometimes deposited in extravagant quantities, from Chinese ritual bronzes and Indian copper hoards , to European hoards of unused axe-heads. By

2835-517: The fins. The tetrapods , a mostly terrestrial superclass of vertebrates, are now recognized as having evolved from sarcopterygian ancestors and are most closely related to lungfishes . Their paired pectoral and pelvic fins evolved into limbs , and their foregut diverticulum eventually evolved into air-breathing lungs . Cladistically , this would make the tetrapods a subgroup within Sarcopterygii and thus sarcopterygians themselves. As

2898-493: The first organized settlements and blossoming of artistic work. Throughout the Palaeolithic, humans generally lived as nomadic hunter-gatherers . Hunter-gatherer societies tended to be very small and egalitarian, although hunter-gatherer societies with abundant resources or advanced food-storage techniques sometimes developed sedentary lifestyles with complex social structures such as chiefdoms, and social stratification . Long-distance contacts may have been established, as in

2961-497: The first signs of deforestation have been found, although this would only begin in earnest during the Neolithic, when more space was needed for agriculture . The Mesolithic is characterized in most areas by small composite flint tools: microliths and microburins . Fishing tackle , stone adzes , and wooden objects such as canoes and bows have been found at some sites. These technologies first occur in Africa, associated with

3024-461: The following Iron Age . The three-age division of prehistory into Stone Age , Bronze Age , and Iron Age remains in use for much of Eurasia and North Africa , but is not generally used in those parts of the world where the working of hard metals arrived abruptly from contact with Eurasian cultures, such as Oceania , Australasia , much of Sub-Saharan Africa , and parts of the Americas . With some exceptions in pre-Columbian civilizations in

3087-473: The introduction of agriculture , the date of which varied by geographic region. In some areas, such as the Near East , agriculture was already underway by the end of the Pleistocene , and there the Mesolithic is short and poorly defined. In areas with limited glacial impact, the term " Epipalaeolithic " is preferred. Regions that experienced greater environmental effects as the last ice age ended have

3150-474: The keeping of dogs , sheep , and goats . By about 6,900–6,400 BCE, it included domesticated cattle and pigs, the establishment of permanently or seasonally inhabited settlements, and the use of pottery . The Neolithic period saw the development of early villages , agriculture , animal domestication , tools , and the onset of the earliest recorded incidents of warfare. Settlements became more permanent, some with circular houses made of mudbrick with

3213-401: The lip, meaning a ventral 'displacement' of the posterior nostril can be considered a synapomorphy of the lungfish-tetrapod clade. The complete choana then seems to have developed independently in the two surviving clades. The word "Rhipidistia" is from Ancient Greek : ῥιπίδιον , romanized :  rhipídion , lit.   'small bellows '. The word "Dipnotetrapodomorpha"

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3276-401: The nature and behavior of pre-literate and non-literate peoples. Human population geneticists and historical linguists are also providing valuable insight. Cultural anthropologists help provide context for societal interactions, by which objects of human origin pass among people, allowing an analysis of any article that arises in a human prehistoric context. Therefore, data about prehistory

3339-1470: The only tetrapodomorphs which survived after the Devonian, with the latter group disappearing during the Permian. Non-tetrapod sarcopterygians continued until towards the end of Paleozoic era, suffering heavy losses during the Permian–Triassic extinction event (251 Ma). The cladogram presented below is based on studies compiled by Janvier et al . (1997) for the Tree of Life Web Project , Mikko's Phylogeny Archive and Swartz (2012). † Onychodontidae Actinistia (coelacanths) † Styloichthys changae Zhu & Yu, 2002 † Porolepiformes Dipnoi (lungfishes) ?† Tungsenia paradoxa Lu et al. , 2012 † Kenichthys campbelli Chang & Zhu, 1993 † Rhizodontiformes ?† Thysanolepidae † Canowindridae † Osteolepiformes † Tristichopteridae † Tinirau clackae Swartz, 2012 † Platycephalichthys Vorobyeva, 1959 † Panderichthys rhombolepis Gross, 1941 † Elpistostegidae † Elginerpeton † Metaxygnathus denticulus Campbell & Bell, 1977 † Ventastega curonica Tetrapoda s.s. ==References== [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] Prehistoric Prehistory , also called pre-literary history ,

3402-613: The other hand, evolved into the fully-limbed stegocephalians and later the fully terrestrial tetrapods during the Late Devonian , when the Late Devonian Extinction bottlenecked and selected against the more aquatically adapted groups among stem-tetrapods . The surviving tetrapods then underwent adaptive radiation on dry land and become the dominant terrestrial animals during the Carboniferous and

3465-465: The presence of swim bladders (which share ancestry with lungs) as well as the evolution of ossified endoskeleton instead of cartilages like the skeletons of acanthodians , chondrichthyians and most placoderms . There are otherwise vast differences in fin, respiratory and circulatory structures between the Sarcopterygii and the Actinopterygii, such as the presence of cosmoid layers in

3528-658: The regions and civilizations who developed a system of keeping written records during later periods. The invention of writing coincides in some areas with the beginnings of the Bronze Age. After the appearance of writing, people started creating texts including written records of administrative matters. The Bronze Age refers to a period in human cultural development when the most advanced metalworking (at least in systematic and widespread use) included techniques for smelting copper and tin from naturally occurring outcroppings of ores, and then combining them to cast bronze . These naturally occurring ores typically included arsenic as

3591-502: The rise of metallurgy, and most Neolithic societies on the whole were relatively simple and egalitarian. Most clothing appears to have been made of animal skins, as indicated by finds of large numbers of bone and antler pins which are ideal for fastening leather. Wool cloth and linen might have become available during the later Neolithic, as suggested by finds of perforated stones that (depending on size) may have served as spindle whorls or loom weights. In Old World archaeology,

3654-459: The scales of sarcopterygians. The earliest sarcopterygian fossils were found in the uppermost Silurian , about 418  Ma . They closely resembled the acanthodians (the "spiny fish", a taxon that became extinct at the end of the Paleozoic). In the early–middle Devonian (416–385 Ma), while the predatory placoderms dominated the seas, some sarcopterygians came into freshwater habitats. In

3717-635: The sometimes biased accounts in Greek and Roman literature, of these protohistoric cultures. In dividing up human prehistory in Eurasia, historians typically use the three-age system, whereas scholars of pre-human time periods typically use the well-defined geologic record and its internationally defined stratum base within the geologic time scale . The three-age system is the periodization of human prehistory into three consecutive time periods , named for their predominant tool-making technologies: Stone Age , Bronze Age and Iron Age . In some areas, there

3780-634: The two groups of living species, the coelacanths and the lungfishes , the largest species is the West Indian Ocean coelacanth , reaching 2 m (6 ft 7 in) in length and weighing up 110 kg (240 lb). The largest lungfish is the marbled lungfish which can reach 2 m (6.6 ft) in length and weigh up to 50 kg (110 lb). Taxonomists who adhere to the cladistic approach include Tetrapoda within this classification, encompassing all species of vertebrates with four limbs. The fin-limbs found in lobe-finned fishes like

3843-654: The two species of coelacanths and six species of lungfishes . Early lobe-finned fishes are bony fish with fleshy, lobed, paired fins, which are joined to the body by a single bone. The fins of lobe-finned fishes differ from those of all other fish in that each is borne on a fleshy, lobelike, scaly stalk extending from the body that resembles a limb bud . The scales of sarcopterygians are true scaloids, consisting of lamellar bone surrounded by layers of vascular bone, cosmine (similar to dentin ), and external keratin . The physical structure of tetrapodomorphs, fish bearing resemblance to tetrapods, provides valuable insights into

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3906-587: The use and provenance of materials, and genetic analysis of bones to determine kinship and physical characteristics of prehistoric peoples. The beginning of prehistory is normally taken to be marked by human-like beings appearing on Earth. The date marking its end is typically defined as the advent of the contemporary written historical record. Both dates consequently vary widely from region to region. For example, in European regions, prehistory cannot begin before c.  1.3  million years ago, which

3969-460: The whole area. "Palaeolithic" means "Old Stone Age", and begins with the first use of stone tools . The Paleolithic is the earliest period of the Stone Age . It extends from the earliest known use of stone tools by hominins c.  3.3  million years ago, to the end of the Pleistocene c.  11,650   BP (before the present period). The early part of the Palaeolithic

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