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52-447: Rice is a cereal grain. Rice , Rice's , or Rices may also refer to: Rice Rice is a cereal grain and in its domesticated form is the staple food of over half of the world's population , particularly in Asia and Africa . Rice is the seed of the grass species Oryza sativa (Asian rice)—or, much less commonly, Oryza glaberrima (African rice). Asian rice

104-641: A pollen grain . The pollen is eventually released when the anther forms openings ( dehisces ). These may consist of longitudinal slits, pores, as in the heath family ( Ericaceae ), or by valves, as in the barberry family ( Berberidaceae ). In some plants, notably members of Orchidaceae and Asclepiadoideae , the pollen remains in masses called pollinia , which are adapted to attach to particular pollinating agents such as birds or insects. More commonly, mature pollen grains separate and are dispensed by wind or water, pollinating insects, birds or other pollination vectors. Pollen of angiosperms must be transported to

156-403: A branched inflorescence which arises from the last internode on the stem. There can be up to 350 spikelets in a panicle, each containing male and female flower parts ( anthers and ovule ). A fertilised ovule develops into the edible grain or caryopsis . Rice is a cereal belonging to the family Poaceae . As a tropical crop, it can be grown during the two distinct seasons (dry and wet) of

208-402: A great variety of patterns, some of them highly complex. It generally surrounds the gynoecium and is surrounded by the perianth . A few members of the family Triuridaceae , particularly Lacandonia schismatica and Lacandonia brasiliana , along with a few species of Trithuria (family Hydatellaceae ) are exceptional in that their gynoecia surround their androecia. Depending on

260-584: A millimeter in Wolfia spp up to five inches (13 centimeters) in Canna iridiflora and Strelitzia nicolai . The stamens in a flower are collectively called the androecium . The androecium can consist of as few as one-half stamen (i.e. a single locule ) as in Canna species or as many as 3,482 stamens which have been counted in the saguaro ( Carnegiea gigantea ). The androecium in various species of plants forms

312-411: A role in various religions and traditions, such as in weddings . The rice plant can grow to over 1 m (3 ft) tall; if in deep water, it can reach a length of 5 m (16 ft). A single plant may have several leafy stems or tillers . The upright stem is jointed with nodes along its length; a long slender leaf arises from each node. The self-fertile flowers are produced in a panicle ,

364-514: A single domestication event for O. sativa . Both indica and japonica forms of Asian rice sprang from a single domestication event in China from the wild rice Oryza rufipogon . Despite this evidence, it appears that indica rice arose when japonica arrived in India about 4,500 years ago and hybridised with another rice, whether an undomesticated proto- indica or wild O. nivara . Rice

416-440: A wide variety of pests including insects, nematodes, rodents such as rats, snails, and birds. Major rice insect pests include armyworms, rice bugs , black bugs , cutworms, field crickets, grasshoppers, leafhoppers, mealybugs, and planthoppers. High rates of nitrogen fertiliser application may worsen aphid outbreaks. Weather conditions can contribute to pest outbreaks: rice gall midge outbreaks are worsened by high rainfall in

468-436: Is rainfed like wheat or maize. Across Asia, unmilled rice or "paddy" (Indonesian and Malay padi ), was traditionally the product of smallholder agriculture, with manual harvesting . Larger farms make use of machines such as combine harvesters to reduce the input of labour. The grain is ready to harvest when the moisture content is 20–25%. Harvesting involves reaping , stacking the cut stalks, threshing to separate

520-678: Is 69% water, 29% carbohydrates , 2% protein , and contains negligible fat (table). In a reference serving of 100 grams (3.5 oz), cooked white rice provides 130 calories of food energy , and contains moderate levels of manganese (18% DV), with no other micronutrients in significant content (all less than 10% of the Daily Value ). In 2018, the World Health Organization strongly recommended fortifying rice with iron , and conditionally recommended fortifying it with vitamin A and with folic acid . Golden rice

572-627: Is a major food staple in Asia, Latin America, and some parts of Africa, feeding over half the world's population. However, a substantial part of the crop can be lost post-harvest through inefficient transportation, storage, and milling. A quarter of the crop in Nigeria is lost after harvest. Storage losses include damage by mould fungi if the rice is not dried sufficiently. In China, losses in modern metal silos were just 0.2%, compared to 7–13% when rice

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624-504: Is a variety produced through genetic engineering to synthesize beta-carotene , a precursor of vitamin A, in the endosperm of the rice grain. It is intended to be grown and eaten in parts of the world where Vitamin A deficiency is prevalent. Golden rice has been opposed by activists, such as in the Philippines . In 2016 more than 100 Nobel laureates encouraged the use of genetically modified organisms , such as golden rice, for

676-581: Is based on four principles: biodiversity, host plant resistance, landscape ecology, and hierarchies in a landscape—from biological to social. Farmers' pesticide applications are often unnecessary. Pesticides may actually induce resurgence of populations of rice pests such as the brown planthopper , both by destroying beneficial insects and by enhancing the pest's reproduction. The International Rice Research Institute (IRRI) demonstrated in 1993 that an 87.5% reduction in pesticide use can lead to an overall drop in pest numbers. Farmers in China, Indonesia and

728-458: Is called appendiculate , e.g. Nerium odorum and some other species of Apocynaceae . In Nerium , the appendages are united as a staminal corona. A column formed from the fusion of multiple filaments is known as an androphore . Stamens can be connate (fused or joined in the same whorl) as follows: Anther shapes are variously described by terms such as linear , rounded , sagittate , sinuous , or reniform . The anther can be attached to

780-464: Is called a staminate flower , or (inaccurately) a male flower. A flower with a functional pistil but no functional stamens is called a pistillate flower , or (inaccurately) a female flower. An abortive or rudimentary stamen is called a staminodium or staminode , such as in Scrophularia nodosa . The carpels and stamens of orchids are fused into a column . The top part of the column

832-410: Is formed by the anther, which is covered by an anther cap . Stamens can also be adnate (fused or joined from more than one whorl): They can have different lengths from each other: or respective to the rest of the flower ( perianth ): They may be arranged in one of two different patterns: They may be arranged, with respect to the petals : Where the connective is very small, or imperceptible,

884-508: Is from Asia. The average world yield for rice was 4.7 metric tons per hectare (2.1 short tons per acre), in 2022. Yuan Longping of China's National Hybrid Rice Research and Development Center set a world record for rice yield in 1999 at 17.1 metric tons per hectare (7.6 short tons per acre) on a demonstration plot. This employed specially developed hybrid rice and the System of Rice Intensification (SRI), an innovation in rice farming. Rice

936-812: Is sister to the bamboos, Bambusoideae , and the cereal subfamily Pooideae . The rice genus Oryza is one of eleven in the Oryzeae; it is sister to the Phyllorachideae . The edible rice species O. sativa and O. glaberrima are among some 300 species or subspecies in the genus. other grasses (inc. the C4 grasses , maize , sorghum ) Streptogyneae Ehrharteae Phyllorachideae Wild rices inc. Zizania other rice species and subspecies O. sativa (Asian rice) O. glaberrima (African rice) Bambusoideae (bamboos) Pooideae (grasses and cereals inc. wheat , barley ) Oryza sativa rice

988-694: Is stickier, and is used for sweet dishes, and in Italy for risotto ; and sticky short-grain rice is used in Japanese sushi as it keeps its shape when cooked. White rice when cooked contains 29% carbohydrate and 2% protein, with some manganese . Golden rice is a variety produced by genetic engineering to contain vitamin A . Production of rice is estimated to have caused over 1% of global greenhouse gas emissions in 2022. Predictions of how rice yields will be affected by climate change vary across geographies and socioeconomic contexts. In human culture, rice plays

1040-438: Is subjected to a steaming process before it is milled. This makes the grain harder, and moves some of the grain's vitamins and minerals into the white part of the rice so these are retained after milling. Rice does not contain gluten , so is suitable for people on a gluten-free diet . Rice is a good source of protein and a staple food in many parts of the world, but it is not a complete protein as it does not contain all of

1092-406: Is to flood the field to a depth of 5 cm (2 in), then to let the water level drop to 15 cm (6 in) below surface level, as measured by looking into a perforated field water tube sunk into the soil, and then repeating the cycle. Deepwater rice varieties tolerate flooding to a depth of over 50 centimetres for at least a month. Upland rice is grown without flooding, in hilly or mountainous regions; it

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1144-471: Is used as a model organism in biology. Dry rice grain is milled to remove the outer layers; depending on how much is removed, products range from brown rice to rice with germ and white rice. Some is parboiled to make it easy to cook. Rice contains no gluten ; it provides protein but not all the essential amino acids needed for good health. Rice of different types is eaten around the world. Long-grain rice tends to stay intact on cooking; medium-grain rice

1196-440: The essential amino acids in sufficient amounts for good health. World trade figures are much smaller than those for production, as less than 8% of rice produced is traded internationally. China, an exporter of rice in the early 2000s, had become the world's largest importer of rice by 2013. Developing countries are the main players in the world rice trade; by 2012, India was the largest exporter of rice, with Thailand and Vietnam

1248-443: The stigma , the receptive surface of the carpel , of a compatible flower, for successful pollination to occur. After arriving, the pollen grain (an immature microgametophyte) typically completes its development. It may grow a pollen tube and undergo mitosis to produce two sperm nuclei. In the typical flower (that is, in the majority of flowering plant species) each flower has both carpels and stamens . In some species, however,

1300-478: The 21st century as people in Asia and elsewhere ate less grain and more meat. An exception is Sub-Saharan Africa, where both per capita consumption of rice and population are increasing. Rice is a commonly-eaten food around the world. The varieties of rice are typically classified as short-, medium-, and long-grained. Oryza sativa indica varieties are usually long-grained; Oryza sativa japonica varieties are usually short- or medium-grained. Short-grain rice, with

1352-704: The Americas by the Spanish. In British North America by the time of the start of the American War of Independence , rice had become the fourth most valuable export commodity behind only tobacco, wheat, and fish. In 2021, world production of rice was 787 million tonnes , led by China and India with a combined 52% of the total. This placed rice fourth in the list of crops by production, after sugarcane , maize , and wheat . Other major producers were Bangladesh , Indonesia and Vietnam . 90% of world production

1404-466: The Philippines have traditionally managed weeds and pests by the polycultural practice of raising ducks and sometimes fish in their rice paddies. These produce valuable additional crops, eat small pest animals, manure the rice, and in the case of ducks also control weeds. Rice plants produce their own chemical defences to protect themselves from pest attacks. Some synthetic chemicals, such as

1456-474: The anther (anther sacs or pollen sacs). The two separate locules on each side of an anther may fuse into a single locule. Each microsporangium is lined with a nutritive tissue layer called the tapetum and initially contains diploid pollen mother cells. These undergo meiosis to form haploid spores. The spores may remain attached to each other in a tetrad or separate after meiosis. Each microspore then divides mitotically to form an immature microgametophyte called

1508-425: The anther lobes are close together, and the connective is referred to as discrete , e.g. Euphorbia pp., Adhatoda zeylanica . Where the connective separates the anther lobes, it is called divaricate , e.g. Tilia , Justicia gendarussa . The connective may also be a long and stalk-like, crosswise on the filament, this is a distractile connective, e.g. Salvia . The connective may also bear appendages, and

1560-404: The base or in the middle area of the anther. The sterile tissue between the lobes is called the connective , an extension of the filament containing conducting strands. It can be seen as an extension on the dorsal side of the anther. A pollen grain develops from a microspore in the microsporangium and contains the male gametophyte . The size of anthers differs greatly, from a tiny fraction of

1612-465: The benefits these could bring. In 2022, greenhouse gas emissions from rice cultivation were estimated at 5.7 billion tonnes CO2eq, representing 1.2% of total emissions. Within the agriculture sector, rice produces almost half the greenhouse gas emissions from croplands , some 30% of agricultural methane emissions , and 11% of agricultural nitrous oxide emissions. Methane is released from rice fields subject to long-term flooding, as this inhibits

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1664-511: The crop would be lost under these conditions. In the Po Valley in Italy, the arborio and carnaroli risotto rice varieties have suffered poor harvests through drought in the 21st century. The Ente Nazionale Risi  [ it ] is developing drought-resistant varieties; its nuovo prometeo variety has deep roots that enable it to tolerate drought, but is not suitable for risotto. Rice yield can be reduced by weed growth, and

1716-425: The exception of Spanish Bomba, is usually sticky when cooked, and is suitable for puddings. Thai Jasmine rice is aromatic, and unusually for a long-grain rice has some stickiness, with a soft texture. Indian Basmati rice is very long-grained and aromatic. Italian Arborio rice , used for risotto , is of medium length, oval, and quite sticky. Japanese sushi rice is a sticky short-grain variety. Cooked white rice

1768-425: The field where they will grow, or seedlings can be grown in a seedbed and transplanted into the field. Direct seeding needs some 60 to 80 kg of grain per hectare, while transplanting needs less, around 40 kg per hectare, but requires far more labour. Most rice in Asia is transplanted by hand. Mechanical transplanting takes less time but requires a carefully-prepared field and seedlings raised on mats or in trays to fit

1820-400: The flowers are unisexual with only carpels or stamens. ( monoecious = both types of flowers found on the same plant; dioecious = the two types of flower found only on different plants). A flower with only stamens is called androecious . A flower with only carpels is called gynoecious . A pistil consists of one or more carpels. A flower with functional stamens but no functional pistil

1872-511: The grain, and cleaning by winnowing or screening . The rice grain is dried as soon as possible to bring the moisture content down to a level that is safe from mould fungi. Traditional drying relies on the heat of the sun, with the grain spread out on mats or on pavements. The edible rice species are members of the BOP clade within the grass family, the Poaceae . The rice subfamily, Oryzoideae ,

1924-419: The great diversity and continuous evolution of pests. Resistance genes are being sought from wild species of rice, and genetic engineering techniques are being applied. Anther A stamen typically consists of a stalk called the filament and an anther which contains microsporangia . Most commonly anthers are two-lobed (each lobe is termed a locule ) and are attached to the filament either at

1976-575: The herbicide 2,4-D , cause the plant to increase the production of certain defensive chemicals and thereby increase the plant's resistance to some types of pests. Conversely, other chemicals, such as the insecticide imidacloprid , appear to induce changes in the gene expression of the rice that make the plant more susceptible to certain pests. Plant breeders have created rice cultivars incorporating resistance to various insect pests . Conventional plant breeding of resistant varieties has been limited by challenges such as rearing insect pests for testing, and

2028-410: The later years of the 20th century decreased rice yield by between 10% and 20% across 200 farms in seven Asian countries. This may have been caused by increased night-time respiration. IRRI has predicted that Asian rice yields will fall by some 20% per 1°C rise in global mean temperature. Further, rice is unable to yield grain if the flowers experience a temperature of 35 °C or more for over one hour, so

2080-417: The machine. Rice does not thrive if continuously submerged. Rice can be grown in different environments, depending upon water availability. The usual arrangement is for lowland fields to be surrounded by bunds and flooded to a depth of a few centimetres until around a week before harvest time; this requires a large amount of water. The "alternate wetting and drying" technique uses less water. One form of this

2132-428: The other largest exporters. As of 2016, the countries that consumed the most rice were China (29% of total), India, and Indonesia. By 2020, Bangladesh had taken third place from Indonesia. On an annual average from 2020-23, China consumed 154 million tonnes of rice, India consumed 109 million tonnes, and Bangladesh and Indonesia consumed about 36 million tonnes each. Across the world, rice consumption per capita fell in

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2184-812: The rice produced in developing nations is lost after harvest through factors such as poor transport and storage. Rice yields can be reduced by pests including insects , rodents , and birds , as well as by weeds , and by diseases such as rice blast . Traditional rice polycultures such as rice-duck farming , and modern integrated pest management seek to control damage from pests in a sustainable way. Many varieties of rice have been bred to improve crop quality and productivity. Biotechnology has created Green Revolution rice able to produce high yields when supplied with nitrogen fertiliser and managed intensively. Other products are rice able to express human proteins for medicinal use; flood-tolerant or deepwater rice ; and drought-tolerant and salt-tolerant varieties. Rice

2236-1011: The soil from absorbing atmospheric oxygen, resulting in anaerobic fermentation of organic matter in the soil. Emissions can be limited by planting new varieties, not flooding continuously, and removing straw. It is possible to cut methane emissions in rice cultivation by improved water management, combining dry seeding and one drawdown, or executing a sequence of wetting and drying . This results in emission reductions of up to 90% compared to full flooding and even increased yields. Predictions of climate change's effects on rice cultivation vary. Global rice yield has been projected to decrease by around 3.2% with each 1°C increase in global average temperature while another study predicts global rice cultivation will increase initially, plateauing at about 3°C warming (2091–2100 relative to 1850–1900). The impacts of climate change on rice cultivation vary across geographic location and socioeconomic context. For example, rising temperatures and decreasing solar radiation during

2288-432: The species of plant, some or all of the stamens in a flower may be attached to the petals or to the floral axis . They also may be free-standing or fused to one another in many different ways, including fusion of some but not all stamens. The filaments may be fused and the anthers free, or the filaments free and the anthers fused. Rather than there being two locules, one locule of a stamen may fail to develop, or alternatively

2340-496: The two locules may merge late in development to give a single locule. Extreme cases of stamen fusion occur in some species of Cyclanthera in the family Cucurbitaceae and in section Cyclanthera of genus Phyllanthus (family Euphorbiaceae ) where the stamens form a ring around the gynoecium, with a single locule. Plants having a single stamen are referred to as "monandrous." A typical anther contains four microsporangia. The microsporangia form sacs or pockets ( locules ) in

2392-701: The voyage to the rest of the Pacific. It reached Austroasiatic and Kra-Dai -speakers in Mainland Southeast Asia and southern China by 5000 years ago. Rice spread around the rest of the world through cultivation, migration and trade, eventually to the Americas as part of the Columbian exchange after 1492. The now less common Oryza glaberrima (African rice) was independently domesticated in Africa around 3,000 years ago, and introduced to

2444-785: The wet season, while thrips outbreaks are associated with drought. Rice blast , caused by the fungus Magnaporthe grisea , is the most serious disease of growing rice. It and bacterial leaf streak (caused by Xanthomonas oryzae pv. oryzae ) are perennially the two worst rice diseases worldwide; they are both among the ten most important diseases of all crop plants. Other major rice diseases include sheath blight (caused by Rhizoctonia solani ), false smut ( Ustilaginoidea virens ), and bacterial panicle blight ( Burkholderia glumae ). Viral diseases include rice bunchy stunt, rice dwarf, rice tungro , and rice yellow mottle. Crop protection scientists are developing sustainable techniques for managing rice pests. Sustainable pest management

2496-539: The year provided that sufficient water is made available. It is normally an annual, but in the tropics it can survive as a perennial , producing a ratoon crop. Like all crops, rice depends for its growth on both biotic and abiotic environmental factors. The principal biotic factors are crop variety, pests , and plant diseases . Abiotic factors include the soil type, whether lowland or upland, amount of rain or irrigation water, temperature, day length , and intensity of sunlight. Rice grains can be planted directly into

2548-464: Was domesticated in China some 13,500 to 8,200 years ago; African rice was domesticated in Africa about 3,000 years ago. Rice has become commonplace in many cultures worldwide; in 2021, 787 million tons were produced, placing it fourth after sugarcane , maize , and wheat . Only some 8% of rice is traded internationally. China, India, and Indonesia are the largest consumers of rice. A substantial amount of

2600-469: Was first domesticated in China 9,000 years ago, by people of Neolithic cultures in the Upper and Lower Yangtze , associated with Hmong-Mien -speakers and pre-Austronesians , respectively. The functional allele for nonshattering , the critical indicator of domestication in grains, as well as five other single-nucleotide polymorphisms , is identical in both indica and japonica . This implies

2652-574: Was introduced early into Sino-Tibetan cultures in northern China by around 6000 to 5600 years ago, and to the Korean peninsula and Japan by around 5500 to 3200 years ago. It was also carried into Taiwan by the Dapenkeng culture by 5500 to 4000 years ago, before spreading southwards via the Austronesian migrations to Island Southeast Asia , Madagascar , and Guam , but did not survive

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2704-439: Was stored by rural households. The dry grain is milled to remove the outer layers, namely the husk and bran . These can be removed in a single step, in two steps, or as in commercial milling in a multi-step process of cleaning, dehusking, separation, polishing, grading, and weighing. Brown rice only has the inedible husk removed. Further milling removes bran and the germ to create successively whiter products. Parboiled rice

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