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60-563: Over 180, see text Erysimum , or wallflower , is a genus of flowering plants in the cabbage family, Brassicaceae . It includes more than 150 species , both popular garden plants and many wild forms. Erysimum is characterised by star-shaped and/or two-sided) trichomes growing from the stem, with yellow, red, pink or orange flowers and multiseeded seed pods. Wallflowers are annuals , herbaceous perennials or sub-shrubs . The perennial species are short-lived and in cultivation treated as biennials . Most species have stems erect, with

120-557: A species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in the nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

180-480: A broad neocortex region of the brain, fur or hair , and three middle ear bones . These characteristics distinguish them from reptiles and birds , from which their ancestors diverged in the Carboniferous Period over 300 million years ago. Around 6,400 extant species of mammals have been described and divided into 27 orders . The largest orders of mammals, by number of species , are

240-556: A covering of bifid hairs, usually 25 ± 53cm × 2–3mm in size. The leaves are narrow and fixed. The lower leaves are broad and round with backwardly directed lobes, 50–80mm × 0.5–3mm. Stem leaves are linear, entire, growing whitish with 2-fid hairs; 21–43mm × 1.5–2mm. Flower clusters grow at intervals on short equal stalks along the stem, with bright yellow to red or pink bilateral flowers . Flowering occurs during spring and summer. One species, Erysimum semperflorens , native to Morocco and Algeria, has white flowers. The flowering part of

300-517: A disease of Brassicaceae . Growth is best in dry soils with very good drainage, and they are often grown successfully in loose wall mortar, hence the vernacular name. There is a wide range of flower color in the warm spectrum, including white, yellow, orange, red, pink, maroon, purple and russet. The flowers, appearing in spring, usually have a strong, pleasant fragrance. Wallflowers are often associated in spring bedding schemes with tulips and forget-me-nots . The cultivar 'Bowles's Mauve' has gained

360-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

420-583: A long history of use in traditional medicine. In Naturalis Historia by Pliny the Elder ( c.  77 ), Erysimum is classified as a medicinal rather than a food plant. Erysimum cheiri is described as a medicinal herb in De Materia Medica by Pedanius Dioscorides ( c.  70 ), which was the predominant European medical pharmacopeia for more than 1,500 years. Other medieval descriptions of medicinal herbs and their uses, including

480-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

540-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

600-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

660-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

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720-630: Is deterred from feeding and oviposition by cardenolides in Erysimum cheiranthoides . Similarly, Anthocharis cardamines (orange tip butterfly), which oviposits on almost all crucifer species, avoids E. cheiranthoides . Erysimum asperum (western wallflower) is resistant to feeding and oviposition of Pieris napi macdunnoughii (synonym Pieris marginalis , margined white butterfly). Two crucifer-feeding beetles, Phaedon sp. and Phyllotreta sp., were deterred from feeding by cardenolides that were applied to their preferred food plants. Consistent with

780-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

840-699: Is pollinated almost exclusively by Anthophora alluadii . Like most Brassicaceae , species in the genus Erysimum produce glucosinolates as defensive compounds. However, unlike almost all other genera in the Brassicaceae, Erysimum also accumulates cardiac glycosides , another class of phytochemicals with an ecological importance in insect defense. Cardiac glycosides specifically function to prevent insect herbivory and/or oviposition by blocking ion channel function in muscle cells. These chemicals are toxic enough to deter generalist, and even some specialist insect herbivores. Cardiac glycoside production

900-572: Is primarily driven by human poaching and habitat destruction , primarily deforestation . Over 70% of mammal species are in the orders Rodentia , Chiroptera , and Eulipotyphla . Mammal classification has been through several revisions since Carl Linnaeus initially defined the class, and at present , no classification system is universally accepted. McKenna & Bell (1997) and Wilson & Reeder (2005) provide useful recent compendiums. Simpson (1945) provides systematics of mammal origins and relationships that had been taught universally until

960-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

1020-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

1080-426: Is the viviparous placental mammals , so named for the temporary organ ( placenta ) used by offspring to draw nutrition from the mother during gestation . Most mammals are intelligent , with some possessing large brains, self-awareness , and tool use . Mammals can communicate and vocalize in several ways, including the production of ultrasound , scent marking , alarm signals , singing , echolocation ; and, in

1140-539: Is widespread in Erysimum , with at least 48 species in the genus containing these compounds. Accumulation of cardiac glycosides in Erysimum crepidifolium , but not other tested species, is induced by treatment with jasmonic acid and methyl jasmonate , endogenous elicitors of chemical defenses in many plant species. Molecular phylogenetic analysis indicates that Erysimum diversification from other Brassicaceae species that do not produce cardiac glycosides began in

1200-473: The Cretaceous . The relationships between these three lineages is contentious, and all three possible hypotheses have been proposed with respect to which group is basal . These hypotheses are Atlantogenata (basal Boreoeutheria), Epitheria (basal Xenarthra) and Exafroplacentalia (basal Afrotheria). Boreoeutheria in turn contains two major lineages— Euarchontoglires and Laurasiatheria . Estimates for

1260-545: The Dispensatorium des Cordus by Valerius Cordus (1542), Bocks Kräuterbuch by Hieronymus Bock (1577), and Tabernaemontanus' Neuw Kreuterbuch by Jacobus Theodorus Tabernaemontanus (1588), also discuss applications of E. cheiri . In traditional Chinese medicine, Erysimum cheiranthoides has been used to treat heart disease and other ailments. Although medical uses of Erysimum became uncommon in Europe after

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1320-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

1380-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

1440-564: The International Union for Conservation of Nature (IUCN) completed a five-year Global Mammal Assessment for its IUCN Red List , which counted 5,488 species. According to research published in the Journal of Mammalogy in 2018, the number of recognized mammal species is 6,495, including 96 recently extinct. The word " mammal " is modern, from the scientific name Mammalia coined by Carl Linnaeus in 1758, derived from

1500-556: The Latin mamma ("teat, pap"). In an influential 1988 paper, Timothy Rowe defined Mammalia phylogenetically as the crown group of mammals, the clade consisting of the most recent common ancestor of living monotremes ( echidnas and platypuses ) and therian mammals ( marsupials and placentals ) and all descendants of that ancestor. Since this ancestor lived in the Jurassic period, Rowe's definition excludes all animals from

1560-496: The Middle Ages , Erysimum diffusum, as well as purified erysimin and erysimoside, have been applied more recently as Ukrainian ethnobotanical treatments. Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including

1620-485: The Pliocene (2.33–5.2 million years ago), suggesting relatively recent evolution of cardiac glycosides as a defensive trait in this genus. The evolution of novel chemical defenses in plants, such as cardenolides in the genus Erysimum , is predicted to allow escape from herbivory by specialist herbivores and expansion into new ecological niches. The crucifer-feeding specialist Pieries rapae (white cabbage butterfly)

1680-493: The Royal Horticultural Society 's Award of Garden Merit . It can become a bushy evergreen perennial in milder locations. It is strongly scented and attractive to bees. Erysimum is found in a range of habitats across the northern hemisphere, and has developed diverse morphology and growth habits (herbaceous annual or perennial, and woody perennial). Different Erysimum species are used as food plants by

1740-606: The biological classification scheme used, are the primates : apes , monkeys , and lemurs ; the Cetartiodactyla : whales and even-toed ungulates ; and the Carnivora which includes cats , dogs , weasels , bears , seals , and allies. According to Mammal Species of the World , 5,416 species were identified in 2006. These were grouped into 1,229  genera , 153  families and 29 orders. In 2008,

1800-406: The dominant terrestrial animal group from 66 million years ago to the present. The basic mammalian body type is quadrupedal , with most mammals using four limbs for terrestrial locomotion ; but in some, the limbs are adapted for life at sea , in the air , in trees or underground . The bipeds have adapted to move using only the two lower limbs, while the rear limbs of cetaceans and

1860-736: The larvae of some Lepidoptera ( butterflies and moths ) species including the garden carpet ( Xanthorhoe fluctuata ). In addition, some species of weevils, like Ceutorhynchus chlorophanus , live inside the fruits feeding on the developing seeds. Many species of beetles, bugs and grasshoppers eat the leaves and stalks. Some mammalian herbivores, for example mule deer ( Odocoileus hemionus ) in North America, argali ( Ovis ammon ) in Mongolia, red deer ( Cervus elaphus ) in Central Europe, or Spanish ibex ( Capra pyrenaica ) in

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1920-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

1980-428: The rodents , bats , and Eulipotyphla (including hedgehogs , moles and shrews ). The next three are the primates (including humans , monkeys and lemurs ), the even-toed ungulates (including pigs , camels , and whales ), and the Carnivora (including cats , dogs , and seals ). Mammals are the only living members of Synapsida ; this clade , together with Sauropsida (reptiles and birds), constitutes

2040-439: The sea cows are mere internal vestiges . Mammals range in size from the 30–40 millimetres (1.2–1.6 in) bumblebee bat to the 30 metres (98 ft) blue whale —possibly the largest animal to have ever lived. Maximum lifespan varies from two years for the shrew to 211 years for the bowhead whale . All modern mammals give birth to live young, except the five species of monotremes , which lay eggs. The most species-rich group

2100-473: The French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or the scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of

2160-479: The Iberian Peninsula, feed on wallflower flowering and fruiting stalks. Erysimum crepidifolium (pale wallflower) is toxic to some generalist vertebrate herbivores. Most wallflowers are pollinator-generalists, their flowers being visited by many different species of bees, bee flies, hoverflies, butterflies, beetles, and ants. However, there are some specialist species. For example, Erysimum scoparium

2220-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

2280-492: The case of humans, complex language . Mammals can organize themselves into fission–fusion societies , harems , and hierarchies —but can also be solitary and territorial . Most mammals are polygynous , but some can be monogamous or polyandrous . Domestication of many types of mammals by humans played a major role in the Neolithic Revolution , and resulted in farming replacing hunting and gathering as

2340-812: The clade originating with the last common ancestor of Sinoconodon and living mammals. The earliest-known synapsid satisfying Kemp's definitions is Tikitherium , dated 225 Ma , so the appearance of mammals in this broader sense can be given this Late Triassic date. However, this animal may have actually evolved during the Neogene. As of the early 21st century, molecular studies based on DNA analysis have suggested new relationships among mammal families. Most of these findings have been independently validated by retrotransposon presence/absence data . Classification systems based on molecular studies reveal three major groups or lineages of placental mammals— Afrotheria , Xenarthra and Boreoeutheria —which diverged in

2400-444: The closest thing to an official classification of mammals, despite its known issues. Most mammals, including the six most species-rich orders , belong to the placental group. The three largest orders in numbers of species are Rodentia : mice , rats , porcupines , beavers , capybaras , and other gnawing mammals; Chiroptera : bats; and Eulipotyphla : shrews , moles , and solenodons . The next three biggest orders, depending on

2460-585: The earlier Triassic , despite the fact that Triassic fossils in the Haramiyida have been referred to the Mammalia since the mid-19th century. If Mammalia is considered as the crown group, its origin can be roughly dated as the first known appearance of animals more closely related to some extant mammals than to others. Ambondro is more closely related to monotremes than to therian mammals while Amphilestes and Amphitherium are more closely related to

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2520-451: The end of the 20th century. However, since 1945, a large amount of new and more detailed information has gradually been found: The paleontological record has been recalibrated, and the intervening years have seen much debate and progress concerning the theoretical underpinnings of systematization itself, partly through the new concept of cladistics . Though fieldwork and lab work progressively outdated Simpson's classification, it remains

2580-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

2640-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

2700-490: The hypothesis of enhanced speciation after escape from herbivory, phylogenetic studies involving 128 Erysimum species indicate diversification in Eurasia between 0.5 and 2 million years ago, and in North America between 0.7 and 1.65 million years ago.) This evolutionarily rapid expansion of the Erysimum genus has resulted in several hundred known species distributed throughout the northern hemisphere. Erysimum species have

2760-432: The idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of the same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but

2820-668: The larger Amniota clade. Early synapsids are referred to as " pelycosaurs ." The more advanced therapsids became dominant during the Guadalupian . Mammals originated from cynodonts , an advanced group of therapsids, during the Late Triassic to Early Jurassic . Mammals achieved their modern diversity in the Paleogene and Neogene periods of the Cenozoic era, after the extinction of non-avian dinosaurs , and have been

2880-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

2940-447: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Mammal A mammal (from Latin mamma  'breast') is a vertebrate animal of the class Mammalia ( / m ə ˈ m eɪ l i . ə / ). Mammals are characterized by the presence of milk -producing mammary glands for feeding their young,

3000-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

3060-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

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3120-716: The primary source of food for humans. This led to a major restructuring of human societies from nomadic to sedentary, with more co-operation among larger and larger groups, and ultimately the development of the first civilizations . Domesticated mammals provided, and continue to provide, power for transport and agriculture, as well as food ( meat and dairy products ), fur , and leather . Mammals are also hunted and raced for sport, kept as pets and working animals of various types, and are used as model organisms in science. Mammals have been depicted in art since Paleolithic times, and appear in literature, film, mythology, and religion. Decline in numbers and extinction of many mammals

3180-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

3240-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

3300-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

3360-1098: The stem ranges from 4 to 7mm. There are four pouch-shaped sepals , light green, 5–7mm × 1.5–2mm. The genus name Erysimum is derived from the Ancient Greek erysimon ( ἐρύσιμον , Sisymbrium officinale or Sisymbrium polyceratium    [ d ] , the hedgenettle), itself from the word eryo ( ἐρύω ) meaning to drag or eryso , a form of rhyomai ( ῥύομαι ), meaning "to ward off" or "to heal" in reference to its medicinal properties. Wallflowers are native to temperate Eurasia, North Africa and Macaronesia , and North America south to Costa Rica. Many wallflowers are endemic to small areas, such as: Most wallflower garden cultivars (e.g. Erysimum 'Chelsea Jacket' ) are derived from E. cheiri (often placed in Cheiranthus ), from southern Europe. They are often attacked by fungal and bacterial disease, so they are best grown as biennials and discarded after flowering. They are also susceptible to clubroot ,

3420-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

3480-548: The therians; as fossils of all three genera are dated about 167 million years ago in the Middle Jurassic , this is a reasonable estimate for the appearance of the crown group. T. S. Kemp has provided a more traditional definition: " Synapsids that possess a dentary – squamosal jaw articulation and occlusion between upper and lower molars with a transverse component to the movement" or, equivalently in Kemp's view,

3540-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

3600-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

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