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Totternhoe Knolls

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83-841: Totternhoe Knolls is a 13.1-hectare (32-acre) Site of Special Scientific Interest (SSSI) in Totternhoe in Bedfordshire . It is also a local nature reserve , and part of the Chilterns Area of Outstanding Natural Beauty . The site is owned by Central Bedfordshire Council and leased to the National Trust . Most of the site is maintained jointly by the National Trust and the Wildlife Trust for Bedfordshire, Cambridgeshire and Northamptonshire (WTBCN), and

166-442: A taxonomic rank ; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain , kingdom , phylum ( division is sometimes used in botany in place of phylum ), class , order , family , genus , and species . The Swedish botanist Carl Linnaeus is regarded as the founder of the current system of taxonomy, as he developed

249-469: A basis of morphological and physiological facts as possible, and one in which "place is found for all observational and experimental data relating, even if indirectly, to the constitution, subdivision, origin, and behaviour of species and other taxonomic groups". Ideals can, it may be said, never be completely realized. They have, however, a great value of acting as permanent stimulants, and if we have some, even vague, ideal of an "omega" taxonomy we may progress

332-557: A chaotic and disorganized taxonomic literature. He not only introduced the standard of class, order, genus, and species, but also made it possible to identify plants and animals from his book, by using the smaller parts of the flower (known as the Linnaean system ). Plant and animal taxonomists regard Linnaeus' work as the "starting point" for valid names (at 1753 and 1758 respectively). Names published before these dates are referred to as "pre-Linnaean", and not considered valid (with

415-591: A continuation of the historical management of the land. Where an owner or occupier is unwilling or unable to carry out management, ultimately the conservation body can require it to be done. Public bodies which own or occupy an SSSI have a duty to manage it properly. Site management statements for SSSI in Scotland are available to download from the NatureScot website using the "Sitelink" facility. The law protecting SSSIs now covers everyone, not just public bodies and

498-443: A different sense, to mean the delimitation of species (not subspecies or taxa of other ranks), using whatever investigative techniques are available, and including sophisticated computational or laboratory techniques. Thus, Ernst Mayr in 1968 defined " beta taxonomy " as the classification of ranks higher than species. An understanding of the biological meaning of variation and of the evolutionary origin of groups of related species

581-455: A little way down the Greek alphabet. Some of us please ourselves by thinking we are now groping in a "beta" taxonomy. Turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy. Later authors have used the term in

664-478: A long way upstream of a wetland SSSI might require consultation. Some developments might be neutral or beneficial, even if they are within the SSSI itself – the critical point is whether they harm the interest features. The owners and occupiers of SSSIs are required (Scotland, England, Wales) to obtain consent from the relevant nature conservation body if they want to carry out, cause or permit to be carried out within

747-504: A notable renaissance, principally with respect to theoretical content. Part of the theoretical material has to do with evolutionary areas (topics e and f above), the rest relates especially to the problem of classification. Taxonomy is that part of Systematics concerned with topics (a) to (d) above. A whole set of terms including taxonomy, systematic biology, systematics , scientific classification, biological classification, and phylogenetics have at times had overlapping meanings – sometimes

830-407: A proposed activity would not affect the interest or is beneficial to it, then the conservation body will issue a "consent" allowing it to be carried out without further consultation. If it would be harmful, the conservation body may issue consent subject to conditions or refuse the application. If consent in writing is not given the operation must not proceed. Conditions may cover any relevant aspect of

913-516: A ranked system known as Linnaean taxonomy for categorizing organisms and binomial nomenclature for naming organisms. With advances in the theory, data and analytical technology of biological systematics, the Linnaean system has transformed into a system of modern biological classification intended to reflect the evolutionary relationships among organisms, both living and extinct. The exact definition of taxonomy varies from source to source, but

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996-477: A single continuum, as per the scala naturae (the Natural Ladder). This, as well, was taken into consideration in the great chain of being. Advances were made by scholars such as Procopius , Timotheus of Gaza , Demetrios Pepagomenos , and Thomas Aquinas . Medieval thinkers used abstract philosophical and logical categorizations more suited to abstract philosophy than to pragmatic taxonomy. During

1079-428: A site may contain strata containing vertebrate fossils, insect fossils and plant fossils and it may also be of importance for stratigraphy . Geological sites fall into two types, having different conservation priorities: exposure sites, and deposit sites. Exposure sites are where quarries , disused railway cuttings, cliffs or outcrops give access to extensive geological features, such as particular rock layers. If

1162-425: A standard list for that country. The ORCs/OLDs are not "banned" activities – the list includes activities which would damage the interest, but also many which might be beneficial. For example, " grazing " (a standard item on the list) would require consent, even on a chalk grassland or heathland where grazing is an essential part of management. In England and Wales the list of OLDs is almost the same for each SSSI – and

1245-524: A truly scientific attempt to classify organisms did not occur until the 18th century, with the possible exception of Aristotle, whose works hint at a taxonomy. Earlier works were primarily descriptive and focused on plants that were useful in agriculture or medicine. There are a number of stages in this scientific thinking. Early taxonomy was based on arbitrary criteria, the so-called "artificial systems", including Linnaeus 's system of sexual classification for plants (Linnaeus's 1735 classification of animals

1328-770: A wide variety of invertebrates, including butterflies such as the common blue , the chalkhill blue , and the scarce small blue and Duke of Burgundy fritillary . Totternhoe Castle was probably built in the late eleventh century. Only the earthworks survive, with a mound five metres tall and 40 metres wide. It is unusual in having three baileys. There is access from the National Trust car park off Castle Hill Road. 51°53′07″N 0°34′08″W  /  51.8852°N 0.5688°W  / 51.8852; -0.5688 Site of Special Scientific Interest A site of special scientific interest ( SSSI ) in Great Britain , or an area of special scientific interest ( ASSI ) in

1411-497: Is a critical component of the taxonomic process. As a result, it informs the user as to what the relatives of the taxon are hypothesized to be. Biological classification uses taxonomic ranks, including among others (in order from most inclusive to least inclusive): Domain , Kingdom , Phylum , Class , Order , Family , Genus , Species , and Strain . The "definition" of a taxon is encapsulated by its description or its diagnosis or by both combined. There are no set rules governing

1494-400: Is a novel analysis of the variation patterns in a particular taxon . This analysis may be executed on the basis of any combination of the various available kinds of characters, such as morphological, anatomical , palynological , biochemical and genetic . A monograph or complete revision is a revision that is comprehensive for a taxon for the information given at a particular time, and for

1577-458: Is a resource for fossils. Biological taxonomy is a sub-discipline of biology , and is generally practiced by biologists known as "taxonomists", though enthusiastic naturalists are also frequently involved in the publication of new taxa. Because taxonomy aims to describe and organize life , the work conducted by taxonomists is essential for the study of biodiversity and the resulting field of conservation biology . Biological classification

1660-454: Is considered a part of taxonomy (definitions 1 and 2), or a part of systematics outside taxonomy. For example, definition 6 is paired with the following definition of systematics that places nomenclature outside taxonomy: In 1970, Michener et al. defined "systematic biology" and "taxonomy" (terms that are often confused and used interchangeably) in relation to one another as follows: Systematic biology (hereafter called simply systematics)

1743-419: Is even more important for the second stage of taxonomic activity, the sorting of species into groups of relatives ("taxa") and their arrangement in a hierarchy of higher categories. This activity is what the term classification denotes; it is also referred to as "beta taxonomy". How species should be defined in a particular group of organisms gives rise to practical and theoretical problems that are referred to as

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1826-516: Is governed by published SSSI Selection Guidelines. Within each area, a representative series of the best examples of each significant natural habitat may be notified, and for rarer habitats all examples may be included. Sites of particular significance for various taxonomic groups may be selected (for example birds, dragonflies , butterflies , reptiles, amphibians , etc.)—each of these groups has its own set of selection guidelines. Conservation of biological SSSI/ASSIs usually involves continuation of

1909-420: Is not necessarily absolute—generally it requires the SSSI interest to be considered properly against other factors. Local planning authorities are required to have policies in their development plans which protect SSSIs. They are then required to consult the appropriate conservation body over planning applications which might affect the interest of an SSSI (such a development might not be within or even close to

1992-461: Is now grassland with a rich variety of plant species, including some that are now rare; these are characteristic species of chalk downland and include kidney vetch , horseshoe vetch , large thyme , squinancywort , autumn gentian , clustered bellflower , sainfoin and dwarf thistle . Orchids that grow here include common spotted orchid , Herminium monorchis Musk orchid, Orchis anthropophora Man orchid, bee orchid and twayblade . There are

2075-441: Is often pronounced "triple-S I". Sites notified for their biological interest are known as Biological SSSIs (or ASSIs), and those notified for geological or physiographic interest are Geological SSSIs (or ASSIs). Sites may be divided into management units, with some areas including units that are noted for both biological and geological interest. Biological SSSI/ASSIs may be selected for various reasons, which for Great Britain

2158-536: Is part of the WTBCN Totternhoe nature reserve , which also includes Totternhoe Chalk Quarry and Totternhoe Stone Pit . The SSSI also includes Totternhoe Castle , the earthworks of a Norman motte-and-bailey castle which is a Scheduled monument . Part of the site was formerly a quarry where Totternhoe Stone , a strong type of chalk that was used in Westminster Abbey , was mined. This part

2241-400: Is the field that (a) provides scientific names for organisms, (b) describes them, (c) preserves collections of them, (d) provides classifications for the organisms, keys for their identification, and data on their distributions, (e) investigates their evolutionary histories, and (f) considers their environmental adaptations. This is a field with a long history that in recent years has experienced

2324-529: The Aristotelian system , with additions concerning the philosophical and existential order of creatures. This included concepts such as the great chain of being in the Western scholastic tradition, again deriving ultimately from Aristotle. The Aristotelian system did not classify plants or fungi , due to the lack of microscopes at the time, as his ideas were based on arranging the complete world in

2407-658: The Isle of Man and Northern Ireland , is a conservation designation denoting a protected area in the United Kingdom and Isle of Man . SSSI/ASSIs are the basic building block of site-based nature conservation legislation and most other legal nature/geological conservation designations in the United Kingdom are based upon them, including national nature reserves , Ramsar sites , Special Protection Areas , and Special Areas of Conservation . The acronym "SSSI"

2490-453: The Marine and Coastal Access Act 2010 ). Access to SSSIs is the same as for the rest of the countryside of the relevant country. Most SSSIs/ASSIs are in private ownership and form parts of working farms, forests and estates. In Scotland, people may use their rights of responsible access to visit SSSIs. When designating an SSSI/ASSI, the relevant nature conservation body must formally notify

2573-983: The National Parks and Access to the Countryside Act 1949 , but the current legal framework for SSSIs is provided in England and Wales by the Wildlife and Countryside Act 1981 , amended in 1985 and further substantially amended in 2000 (by the Countryside and Rights of Way Act 2000 ), in Scotland by the Nature Conservation (Scotland) Act 2004 and in Northern Ireland by the Nature Conservation and Amenity Lands (Northern Ireland) Order 1985 . SSSIs are also covered under

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2656-585: The Neomura , the clade that groups together the Archaea and Eucarya , would have evolved from Bacteria, more precisely from Actinomycetota . His 2004 classification treated the archaeobacteria as part of a subkingdom of the kingdom Bacteria, i.e., he rejected the three-domain system entirely. Stefan Luketa in 2012 proposed a five "dominion" system, adding Prionobiota ( acellular and without nucleic acid ) and Virusobiota (acellular but with nucleic acid) to

2739-512: The Renaissance and the Age of Enlightenment , categorizing organisms became more prevalent, and taxonomic works became ambitious enough to replace the ancient texts. This is sometimes credited to the development of sophisticated optical lenses, which allowed the morphology of organisms to be studied in much greater detail. One of the earliest authors to take advantage of this leap in technology

2822-570: The Water Resources Act 1991 and related legislation. An SSSI may be made on any area of land which is considered to be of special interest by virtue of its fauna , flora , geological or physiographical / geomorphological features. SSSI notification can cover any "land" within the area of the relevant nature conservation body, including dry land, land covered by fresh water . The extent to which an SSSI/ASSI may extend seawards differs between countries. In Scotland an SSSI may include

2905-439: The species problem . The scientific work of deciding how to define species has been called microtaxonomy. By extension, macrotaxonomy is the study of groups at the higher taxonomic ranks subgenus and above, or simply in clades that include more than one taxon considered a species, expressed in terms of phylogenetic nomenclature . While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations,

2988-467: The vertebrates ), as well as groups like the sharks and cetaceans , are commonly used. His student Theophrastus (Greece, 370–285 BC) carried on this tradition, mentioning some 500 plants and their uses in his Historia Plantarum . Several plant genera can be traced back to Theophrastus, such as Cornus , Crocus , and Narcissus . Taxonomy in the Middle Ages was largely based on

3071-488: The 1960s. In 1958, Julian Huxley used the term clade . Later, in 1960, Cain and Harrison introduced the term cladistic . The salient feature is arranging taxa in a hierarchical evolutionary tree , with the desideratum that all named taxa are monophyletic. A taxon is called monophyletic if it includes all the descendants of an ancestral form. Groups that have descendant groups removed from them are termed paraphyletic , while groups representing more than one branch from

3154-563: The ORC lists for each SSSI and removed those activities that were unlikely to happen and if they were to would be unlikely to damage the protected natural features, and other activities adequately regulated by other statutory regimes. The intention of this was to remove the need for owners and occupiers to obtain SSSI consent as well as licences/ permits from other authorities (who must consult NatureScot prior to determining such applications). Purely geological SSSIs often have much shorter OLD lists. If

3237-491: The Origin of Species (1859) led to a new explanation for classifications, based on evolutionary relationships. This was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler (1883) and Engler (1886–1892). The advent of cladistic methodology in the 1970s led to classifications based on the sole criterion of monophyly , supported by the presence of synapomorphies . Since then,

3320-607: The SSSI Register, hosted by The Registers of Scotland . Further information about SSSIs in Scotland is available on the NatureScot website. The decision to notify an SSSI is made by the relevant nature conservation body (the appropriate conservation body ) for that part of the United Kingdom: Northern Ireland Environment Agency , Natural England , NatureScot or Natural Resources Wales . SSSIs were originally set up by

3403-414: The SSSI any of the activities listed in the notification. Formerly these activities were called 'potentially damaging operations' or PDOs. Under the current legal arrangements they are called 'operations requiring consent' or ORCs (Scotland), or 'operations likely to damage the SSSI interest' or OLDs (England & Wales). The list of ORCs/OLDs for each SSSI is unique to that site – though all are derived from

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3486-402: The SSSI itself). The effect of this is to prevent development which harms the interest – except where the value of that interest is over-ridden by some more important factor, for example a requirement for a major road or port or oil pipe. The requirement for consultation covers any development which might affect the interest, not just developments within the SSSI itself – for example, a development

3569-522: The animal and plant kingdoms toward the end of the 18th century, well before Charles Darwin's On the Origin of Species was published. The pattern of the "Natural System" did not entail a generating process, such as evolution, but may have implied it, inspiring early transmutationist thinkers. Among early works exploring the idea of a transmutation of species were Zoonomia in 1796 by Erasmus Darwin (Charles Darwin's grandfather), and Jean-Baptiste Lamarck 's Philosophie zoologique of 1809. The idea

3652-426: The core of the discipline remains: the conception, naming, and classification of groups of organisms. As points of reference, recent definitions of taxonomy are presented below: The varied definitions either place taxonomy as a sub-area of systematics (definition 2), invert that relationship (definition 6), or appear to consider the two terms synonymous. There is some disagreement as to whether biological nomenclature

3735-550: The definition of taxa, but the naming and publication of new taxa is governed by sets of rules. In zoology , the nomenclature for the more commonly used ranks ( superfamily to subspecies ), is regulated by the International Code of Zoological Nomenclature ( ICZN Code ). In the fields of phycology , mycology , and botany , the naming of taxa is governed by the International Code of Nomenclature for algae, fungi, and plants ( ICN ). The initial description of

3818-867: The designating authority is NatureScot ; the role in Wales is performed by Natural Resources Wales (formerly the Countryside Council for Wales ). In the Isle of Man the role is performed by the Department of Environment, Food and Agriculture . Geological SSSI/ASSIs are selected by a different mechanism to biological ones, with a minimalistic system selecting one site for each geological feature in Great Britain. Academic geological specialists have reviewed geological literature, selecting sites within Great Britain of at least national importance for each of

3901-399: The entire world. Other (partial) revisions may be restricted in the sense that they may only use some of the available character sets or have a limited spatial scope. A revision results in a conformation of or new insights in the relationships between the subtaxa within the taxon under study, which may lead to a change in the classification of these subtaxa, the identification of new subtaxa, or

3984-494: The evidentiary basis has been expanded with data from molecular genetics that for the most part complements traditional morphology . Naming and classifying human surroundings likely began with the onset of language. Distinguishing poisonous plants from edible plants is integral to the survival of human communities. Medicinal plant illustrations show up in Egyptian wall paintings from c.  1500 BC , indicating that

4067-524: The exception of spiders published in Svenska Spindlar ). Even taxonomic names published by Linnaeus himself before these dates are considered pre-Linnaean. Modern taxonomy is heavily influenced by technology such as DNA sequencing , bioinformatics , databases , and imaging . A pattern of groups nested within groups was specified by Linnaeus' classifications of plants and animals, and these patterns began to be represented as dendrograms of

4150-461: The exposure becomes obscured, the feature could in principle be re-exposed elsewhere. Conservation of these sites usually concentrates on maintenance of access for future study. Deposit sites are features which are limited in extent or physically delicate—for example, they include small lenses of sediment , mine tailings , caves and other landforms . If such features become damaged they cannot be recreated, and conservation usually involves protecting

4233-507: The feature from erosion or other damage. Following devolution, legal arrangements for SSSIs (Scotland, England, Wales) and ASSIs (Northern Ireland) differ between the countries of the UK. The Isle of Man ASSI system is a separate entity. NatureScot publishes a summary of the SSSI arrangements for SSSI owners and occupiers (other than public bodies) which can be downloaded from its website. Legal documents for all SSSIs in Scotland are available on

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4316-486: The first modern groups tied to fossil ancestors was birds. Using the then newly discovered fossils of Archaeopteryx and Hesperornis , Thomas Henry Huxley pronounced that they had evolved from dinosaurs, a group formally named by Richard Owen in 1842. The resulting description, that of dinosaurs "giving rise to" or being "the ancestors of" birds, is the essential hallmark of evolutionary taxonomic thinking. As more and more fossil groups were found and recognized in

4399-687: The formal naming of clades. Linnaean ranks are optional and have no formal standing under the PhyloCode , which is intended to coexist with the current, rank-based codes. While popularity of phylogenetic nomenclature has grown steadily in the last few decades, it remains to be seen whether a majority of systematists will eventually adopt the PhyloCode or continue using the current systems of nomenclature that have been employed (and modified, but arguably not as much as some systematists wish) for over 250 years. Well before Linnaeus, plants and animals were considered separate Kingdoms. Linnaeus used this as

4482-405: The interest), but not illegal trail biking. This loophole was closed by the Countryside and Rights of Way Act 2000 and section 19 of the Nature Conservation (Scotland) Act 2004. Funding for the monitoring of SSSIs in England has been cut from £1.58 million in 2010 to £700,000 in 2018, causing concern that many have not been inspected over the last six years, as required by guidelines. Since

4565-473: The interested parties and allow a period for them to make representations before confirming the notification. When creating a new SSSI/ASSI the designation has legal effect from the date of notification. The interested parties include central government, local planning authorities , national park authorities, all the owners and occupiers of the land, relevant public bodies such as the utility providers e.g., water companies . In Scotland, NatureScot must also notify

4648-469: The intertidal land down to mean low water spring or to the extent of the local planning authority area, thus only limited areas of estuaries and coastal waters beyond MLWS may be included. In England, Natural England may notify an SSSI over estuarial waters and further adjacent waters in certain circumstances (section 28(1A & 1B) of the Wildlife and Countryside Act 1981 as amended by Part 2 of Annex 13 of

4731-466: The late 19th and early 20th centuries, palaeontologists worked to understand the history of animals through the ages by linking together known groups. With the modern evolutionary synthesis of the early 1940s, an essentially modern understanding of the evolution of the major groups was in place. As evolutionary taxonomy is based on Linnaean taxonomic ranks, the two terms are largely interchangeable in modern use. The cladistic method has emerged since

4814-402: The list for an SSSI will only omit activities impossible on the particular SSSI (such as fishing where there is no water), and things requiring planning permission (which are covered by the local planning authority consultation process). In Scotland, and following the implementation of the Nature Conservation (Scotland) Act 2004, Scottish Natural Heritage (the former name for NatureScot) reviewed

4897-401: The merger of previous subtaxa. Taxonomic characters are the taxonomic attributes that can be used to provide the evidence from which relationships (the phylogeny ) between taxa are inferred. Kinds of taxonomic characters include: The term " alpha taxonomy " is primarily used to refer to the discipline of finding, describing, and naming taxa , particularly species. In earlier literature,

4980-531: The most important features within each geological topic (or block ). Each of these sites is described, with most published in the Geological Conservation Review series, and so becomes a GCR site . Almost all GCR sites (but no other sites) are subsequently notified as geological SSSIs, except some that coincide with designated biological SSSI management units. A GCR site may contain features from several different topic blocks, for example

5063-400: The natural and artificial processes which resulted in their development and survival, for example the continued traditional grazing of heathland or chalk grassland . In England, the designating body for SSSIs, Natural England , selects biological SSSIs from within natural areas which are areas with particular landscape and ecological characteristics, or on a county basis. In Scotland,

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5146-1003: The new Act, often with boundary changes. This complex process took some ten years to complete for the several thousand SSSIs. For the purposes of selecting the original tranche of SSSIs, Natural England's predecessors (the Nature Conservancy, the Nature Conservancy Council and English Nature ) used a system termed "areas of search" (AOSs). In England these were largely based on the 1974–1996 administrative counties (with larger counties divided into two or more areas), whereas in Scotland and Wales they are based around districts. The individual AOSs are between 400 km (150 sq mi) and 4,000 km (1,500 sq mi) in size. There were 59 AOSs in England, 12 in Wales, and 44 in Scotland. Watsonian vice-counties were formerly used for selection over

5229-399: The notification is then confirmed or withdrawn (in whole or part). At the time of the passing of the Wildlife and Countryside Act 1981 , many SSSIs were already in existence, having been notified over the previous decades under the National Parks and Access to the Countryside Act 1949 . Each of these was considered in turn, and either denotified, or renotified —brought under the provisions of

5312-447: The owners and occupiers of SSSIs. Previously, activities by "third parties" were not illegal under the SSSI legislation. This meant that damaging activities such as fly-tipping , intensive bait-digging or trail biking on an SSSI were only prevented if done (or permitted) by the owner or occupier – not if done by trespassers or under public rights. The effect was, for example, to allow control of legal trail biking on SSSIs (where damaging to

5395-434: The possibilities of closer co-operation with their cytological, ecological and genetics colleagues and to acknowledge that some revision or expansion, perhaps of a drastic nature, of their aims and methods, may be desirable ... Turrill (1935) has suggested that while accepting the older invaluable taxonomy, based on structure, and conveniently designated "alpha", it is possible to glimpse a far-distant taxonomy built upon as wide

5478-517: The proposed operation and may, for example, limit its timing, location or intensity. The process is slightly different where the owner or occupier is a public body, but the effect is broadly similar. The relevant nature conservation body sends all SSSI owners and occupiers a site-specific 'site management statement' describing the ideal management (there may be grants available to help fund management). Owners and occupiers are encouraged to carry out this management, which in many (but not all) cases will be

5561-818: The rank of Order, although both exclude fossil representatives. A separate compilation (Ruggiero, 2014) covers extant taxa to the rank of Family. Other, database-driven treatments include the Encyclopedia of Life , the Global Biodiversity Information Facility , the NCBI taxonomy database , the Interim Register of Marine and Nonmarine Genera , the Open Tree of Life , and the Catalogue of Life . The Paleobiology Database

5644-572: The referendum to leave the EU in 2016, more than 450 staff have been transferred to the Department for Environment, Food and Rural Affairs (Defra). Neglected areas include Exmoor , the Lake District , the Pennines , and The Wash . The process of designating a site as of Special Scientific Interest is called notification; this is followed by consultation with the site's owners and occupiers, and

5727-565: The relevant community councils and community group having registered an interest in the land. The notification includes a description of the land and the natural features for which it is notified ("the citation"), a boundary map, and a list of the acts or omissions (activities) that the nature conservation body regulates through the issue of consents. The various laws protect the interest features of SSSIs from development, from other damage, and (since 2000 in England) also from neglect. Protection

5810-407: The same, sometimes slightly different, but always related and intersecting. The broadest meaning of "taxonomy" is used here. The term itself was introduced in 1813 by de Candolle , in his Théorie élémentaire de la botanique . John Lindley provided an early definition of systematics in 1830, although he wrote of "systematic botany" rather than using the term "systematics". Europeans tend to use

5893-472: The term had a different meaning, referring to morphological taxonomy, and the products of research through the end of the 19th century. William Bertram Turrill introduced the term "alpha taxonomy" in a series of papers published in 1935 and 1937 in which he discussed the philosophy and possible future directions of the discipline of taxonomy. ... there is an increasing desire amongst taxonomists to consider their problems from wider viewpoints, to investigate

5976-482: The terms "systematics" and "biosystematics" for the study of biodiversity as a whole, whereas North Americans tend to use "taxonomy" more frequently. However, taxonomy, and in particular alpha taxonomy , is more specifically the identification, description, and naming (i.e., nomenclature) of organisms, while "classification" focuses on placing organisms within hierarchical groups that show their relationships to other organisms. A taxonomic revision or taxonomic review

6059-510: The three-domain method is the separation of Archaea and Bacteria , previously grouped into the single kingdom Bacteria (a kingdom also sometimes called Monera ), with the Eukaryota for all organisms whose cells contain a nucleus . A small number of scientists include a sixth kingdom, Archaea, but do not accept the domain method. Thomas Cavalier-Smith , who published extensively on the classification of protists , in 2002 proposed that

6142-427: The top rank, dividing the physical world into the vegetable, animal and mineral kingdoms. As advances in microscopy made the classification of microorganisms possible, the number of kingdoms increased, five- and six-kingdom systems being the most common. Domains are a relatively new grouping. First proposed in 1977, Carl Woese 's three-domain system was not generally accepted until later. One main characteristic of

6225-436: The traditional three domains. Partial classifications exist for many individual groups of organisms and are revised and replaced as new information becomes available; however, comprehensive, published treatments of most or all life are rarer; recent examples are that of Adl et al., 2012 and 2019, which covers eukaryotes only with an emphasis on protists, and Ruggiero et al., 2015, covering both eukaryotes and prokaryotes to

6308-518: The tree of life are called polyphyletic . Monophyletic groups are recognized and diagnosed on the basis of synapomorphies , shared derived character states. Cladistic classifications are compatible with traditional Linnean taxonomy and the Codes of Zoological and Botanical nomenclature , to a certain extent. An alternative system of nomenclature, the International Code of Phylogenetic Nomenclature or PhyloCode has been proposed, which regulates

6391-642: The uses of different species were understood and that a basic taxonomy was in place. Organisms were first classified by Aristotle ( Greece , 384–322 BC) during his stay on the Island of Lesbos . He classified beings by their parts, or in modern terms attributes , such as having live birth, having four legs, laying eggs, having blood, or being warm-bodied. He divided all living things into two groups: plants and animals . Some of his groups of animals, such as Anhaima (animals without blood, translated as invertebrates ) and Enhaima (animals with blood, roughly

6474-437: The whole of Great Britain. Taxonomy (biology) In biology , taxonomy (from Ancient Greek τάξις ( taxis )  'arrangement' and -νομία ( -nomia )  ' method ') is the scientific study of naming, defining ( circumscribing ) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given

6557-486: Was Methodus Plantarum Nova (1682), in which he published details of over 18,000 plant species. At the time, his classifications were perhaps the most complex yet produced by any taxonomist, as he based his taxa on many combined characters. The next major taxonomic works were produced by Joseph Pitton de Tournefort (France, 1656–1708). His work from 1700, Institutiones Rei Herbariae , included more than 9000 species in 698 genera, which directly influenced Linnaeus, as it

6640-555: Was entitled " Systema Naturae " ("the System of Nature"), implying that he, at least, believed that it was more than an "artificial system"). Later came systems based on a more complete consideration of the characteristics of taxa, referred to as "natural systems", such as those of de Jussieu (1789), de Candolle (1813) and Bentham and Hooker (1862–1863). These classifications described empirical patterns and were pre- evolutionary in thinking. The publication of Charles Darwin 's On

6723-615: Was popularized in the Anglophone world by the speculative but widely read Vestiges of the Natural History of Creation , published anonymously by Robert Chambers in 1844. With Darwin's theory, a general acceptance quickly appeared that a classification should reflect the Darwinian principle of common descent . Tree of life representations became popular in scientific works, with known fossil groups incorporated. One of

6806-543: Was the Italian physician Andrea Cesalpino (1519–1603), who has been called "the first taxonomist". His magnum opus De Plantis came out in 1583, and described more than 1500 plant species. Two large plant families that he first recognized are in use: the Asteraceae and Brassicaceae . In the 17th century John Ray ( England , 1627–1705) wrote many important taxonomic works. Arguably his greatest accomplishment

6889-429: Was the text he used as a young student. The Swedish botanist Carl Linnaeus (1707–1778) ushered in a new era of taxonomy. With his major works Systema Naturae 1st Edition in 1735, Species Plantarum in 1753, and Systema Naturae 10th Edition , he revolutionized modern taxonomy. His works implemented a standardized binomial naming system for animal and plant species, which proved to be an elegant solution to

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