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105-636: The Toad and Grayling strata have yielded fossils of marine organisms, including ammonites , brachiopods , and bivalves . The Toad and Grayling Formations were originally described by E.D. Kindle in 1944, who named them for the Toad and Grayling Rivers, which are tributaries of the Liard River in northeasternmost British Columbia. They were combined as the Toad-Grayling Formation by A.D. Hunt and J.D. Ratcliffe in 1959. The Toad-Grayling

210-423: A "daughter" nuclide or decay product . In many cases, the daughter nuclide itself is radioactive, resulting in a decay chain , eventually ending with the formation of a stable (nonradioactive) daughter nuclide; each step in such a chain is characterized by a distinct half-life. In these cases, usually the half-life of interest in radiometric dating is the longest one in the chain, which is the rate-limiting factor in

315-510: A better time resolution than that available from long-lived isotopes, short-lived isotopes that are no longer present in the rock can be used. At the beginning of the solar system, there were several relatively short-lived radionuclides like Al, Fe, Mn, and I present within the solar nebula. These radionuclides—possibly produced by the explosion of a supernova—are extinct today, but their decay products can be detected in very old material, such as that which constitutes meteorites . By measuring

420-471: A certain temperature, the crystal structure has formed sufficiently to prevent diffusion of isotopes. Thus an igneous or metamorphic rock or melt, which is slowly cooling, does not begin to exhibit measurable radioactive decay until it cools below the closure temperature. The age that can be calculated by radiometric dating is thus the time at which the rock or mineral cooled to closure temperature. This temperature varies for every mineral and isotopic system, so

525-408: A consequence of background radiation on certain minerals. Over time, ionizing radiation is absorbed by mineral grains in sediments and archaeological materials such as quartz and potassium feldspar . The radiation causes charge to remain within the grains in structurally unstable "electron traps". Exposure to sunlight or heat releases these charges, effectively "bleaching" the sample and resetting

630-472: A constant rate. These " molecular clocks ", however, are fallible, and provide only approximate timing: for example, they are not sufficiently precise and reliable for estimating when the groups that feature in the Cambrian explosion first evolved, and estimates produced by different techniques may vary by a factor of two. Organisms are only rarely preserved as fossils in the best of circumstances, and only

735-426: A fraction of such fossils have been discovered. This is illustrated by the fact that the number of species known through the fossil record is less than 5% of the number of known living species, suggesting that the number of species known through fossils must be far less than 1% of all the species that have ever lived. Because of the specialized and rare circumstances required for a biological structure to fossilize, only

840-402: A half-life of 1.3 billion years, so this method is applicable to the oldest rocks. Radioactive potassium-40 is common in micas , feldspars , and hornblendes , though the closure temperature is fairly low in these materials, about 350 °C (mica) to 500 °C (hornblende). This is based on the beta decay of rubidium-87 to strontium-87 , with a half-life of 50 billion years. This scheme

945-499: A hierarchical classification system still in use today. Darwin and his contemporaries first linked the hierarchical structure of the tree of life with the then very sparse fossil record. Darwin eloquently described a process of descent with modification, or evolution, whereby organisms either adapt to natural and changing environmental pressures, or they perish. When Darwin wrote On the Origin of Species by Means of Natural Selection, or

1050-525: A high-temperature furnace. This field is known as thermochronology or thermochronometry. The mathematical expression that relates radioactive decay to geologic time is where The equation is most conveniently expressed in terms of the measured quantity N ( t ) rather than the constant initial value N o . To calculate the age, it is assumed that the system is closed (neither parent nor daughter isotopes have been lost from system), D 0 either must be negligible or can be accurately estimated, λ

1155-479: A higher time resolution at the expense of timescale. I beta-decays to Xe with a half-life of 16.14 ± 0.12 million years . The iodine-xenon chronometer is an isochron technique. Samples are exposed to neutrons in a nuclear reactor. This converts the only stable isotope of iodine ( I ) into Xe via neutron capture followed by beta decay (of I ). After irradiation, samples are heated in

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1260-399: A kiln. Other methods include: Absolute radiometric dating requires a measurable fraction of parent nucleus to remain in the sample rock. For rocks dating back to the beginning of the solar system, this requires extremely long-lived parent isotopes, making measurement of such rocks' exact ages imprecise. To be able to distinguish the relative ages of rocks from such old material, and to get

1365-418: A mid-Ordovician age. Such index fossils must be distinctive, be globally distributed and occupy a short time range to be useful. Misleading results are produced if the index fossils are incorrectly dated. Stratigraphy and biostratigraphy can in general provide only relative dating ( A was before B ), which is often sufficient for studying evolution. However, this is difficult for some time periods, because of

1470-536: A mutation first appeared. Phylogenetics and paleontology work together in the clarification of science's still dim view of the appearance of life and its evolution. Niles Eldredge 's study of the Phacops trilobite genus supported the hypothesis that modifications to the arrangement of the trilobite's eye lenses proceeded by fits and starts over millions of years during the Devonian . Eldredge's interpretation of

1575-458: A particular element is called a nuclide . Some nuclides are inherently unstable. That is, at some point in time, an atom of such a nuclide will undergo radioactive decay and spontaneously transform into a different nuclide. This transformation may be accomplished in a number of different ways, including alpha decay (emission of alpha particles ) and beta decay ( electron emission, positron emission, or electron capture ). Another possibility

1680-467: A polished slice of a material to determine the density of "track" markings left in it by the spontaneous fission of uranium-238 impurities. The uranium content of the sample has to be known, but that can be determined by placing a plastic film over the polished slice of the material, and bombarding it with slow neutrons . This causes induced fission of U, as opposed to the spontaneous fission of U. The fission tracks produced by this process are recorded in

1785-570: A portion of the deceased organism, usually that portion that was partially mineralized during life, such as the bones and teeth of vertebrates , or the chitinous or calcareous exoskeletons of invertebrates . Fossils may also consist of the marks left behind by the organism while it was alive, such as animal tracks or feces ( coprolites ). These types of fossil are called trace fossils or ichnofossils , as opposed to body fossils . Some fossils are biochemical and are called chemofossils or biosignatures . Gathering fossils dates at least to

1890-438: A range of several hundred thousand years. A related method is ionium–thorium dating , which measures the ratio of ionium (thorium-230) to thorium-232 in ocean sediment . Radiocarbon dating is also simply called carbon-14 dating. Carbon-14 is a radioactive isotope of carbon, with a half-life of 5,730 years (which is very short compared with the above isotopes), and decays into nitrogen. In other radiometric dating methods,

1995-400: A richly diverse assembly of early multicellular eukaryotes . The fossil record and faunal succession form the basis of the science of biostratigraphy or determining the age of rocks based on embedded fossils. For the first 150 years of geology , biostratigraphy and superposition were the only means for determining the relative age of rocks. The geologic time scale was developed based on

2100-406: A series of steps and the xenon isotopic signature of the gas evolved in each step is analysed. When a consistent Xe / Xe ratio is observed across several consecutive temperature steps, it can be interpreted as corresponding to a time at which the sample stopped losing xenon. Samples of a meteorite called Shallowater are usually included in the irradiation to monitor

2205-435: A single sample to accurately measure them. A faster method involves using particle counters to determine alpha, beta or gamma activity, and then dividing that by the number of radioactive nuclides. However, it is challenging and expensive to accurately determine the number of radioactive nuclides. Alternatively, decay constants can be determined by comparing isotope data for rocks of known age. This method requires at least one of

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2310-448: A small percentage of life-forms can be expected to be represented in discoveries, and each discovery represents only a snapshot of the process of evolution. The transition itself can only be illustrated and corroborated by transitional fossils, which will never demonstrate an exact half-way point. The fossil record is strongly biased toward organisms with hard-parts, leaving most groups of soft-bodied organisms with little to no role. It

2415-448: A system can be closed for one mineral but open for another. Dating of different minerals and/or isotope systems (with differing closure temperatures) within the same rock can therefore enable the tracking of the thermal history of the rock in question with time, and thus the history of metamorphic events may become known in detail. These temperatures are experimentally determined in the lab by artificially resetting sample minerals using

2520-460: A variable amount of uranium content. Because the fission tracks are healed by temperatures over about 200 °C the technique has limitations as well as benefits. The technique has potential applications for detailing the thermal history of a deposit. Large amounts of otherwise rare Cl (half-life ~300ky) were produced by irradiation of seawater during atmospheric detonations of nuclear weapons between 1952 and 1958. The residence time of Cl in

2625-437: Is spontaneous fission into two or more nuclides. While the moment in time at which a particular nucleus decays is unpredictable, a collection of atoms of a radioactive nuclide decays exponentially at a rate described by a parameter known as the half-life , usually given in units of years when discussing dating techniques. After one half-life has elapsed, one half of the atoms of the nuclide in question will have decayed into

2730-449: Is a notable example of how knowledge encoded by the fossil record continues to contribute otherwise unattainable information on the emergence and development of life on Earth. For example, the research suggests Markuelia has closest affinity to priapulid worms, and is adjacent to the evolutionary branching of Priapulida , Nematoda and Arthropoda . Despite significant advances in uncovering and identifying paleontological specimens, it

2835-445: Is a technique which is used to date materials such as rocks or carbon , in which trace radioactive impurities were selectively incorporated when they were formed. The method compares the abundance of a naturally occurring radioactive isotope within the material to the abundance of its decay products, which form at a known constant rate of decay. The use of radiometric dating was first published in 1907 by Bertram Boltwood and

2940-412: Is any preserved remains, impression, or trace of any once-living thing from a past geological age . Examples include bones , shells , exoskeletons , stone imprints of animals or microbes , objects preserved in amber , hair , petrified wood and DNA remnants. The totality of fossils is known as the fossil record . Though the fossil record is incomplete, numerous studies have demonstrated that there

3045-538: Is enough information available to give a good understanding of the pattern of diversification of life on Earth. In addition, the record can predict and fill gaps such as the discovery of Tiktaalik in the arctic of Canada . Paleontology includes the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years to 4.1 billion years old. The observation in

3150-744: Is equivalent to the lower Montney Formation in the subsurface of the Peace River plains and to the Phroso Siltstone Member of the Sulphur Mountain Formation in west-central and southwestern Alberta . Its contact with the overlying Toad Formation is gradational. The Toad Formation is conformably overlain by the Liard Formation . North of the Peace River it is overlain, possibly unconformably, by

3255-479: Is generally accepted that the fossil record is vastly incomplete. Approaches for measuring the completeness of the fossil record have been developed for numerous subsets of species, including those grouped taxonomically, temporally, environmentally/geographically, or in sum. This encompasses the subfield of taphonomy and the study of biases in the paleontological record. Paleontology seeks to map out how life evolved across geologic time. A substantial hurdle

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3360-432: Is known to high precision, and one has accurate and precise measurements of D* and N ( t ). The above equation makes use of information on the composition of parent and daughter isotopes at the time the material being tested cooled below its closure temperature . This is well established for most isotopic systems. However, construction of an isochron does not require information on the original compositions, using merely

3465-570: Is more calcareous and less dolomitic than the Grayling. It consists of dark grey calcareous siltstone and silty limestone, with minor amounts of silty dolomite and calcareous sandstone and, in the lower part, minor thin, randomly dispersed lenses and nodules of phosphate . It reaches a maximum thickness of about 825 metres (2700 ft). The Grayling Formation unconformably overlies the Permian Fantasque Formation . It

3570-409: Is now the principal source of information about the absolute age of rocks and other geological features , including the age of fossilized life forms or the age of Earth itself, and can also be used to date a wide range of natural and man-made materials . Together with stratigraphic principles , radiometric dating methods are used in geochronology to establish the geologic time scale . Among

3675-533: Is often performed on the mineral zircon (ZrSiO 4 ), though it can be used on other materials, such as baddeleyite and monazite (see: monazite geochronology ). Zircon and baddeleyite incorporate uranium atoms into their crystalline structure as substitutes for zirconium , but strongly reject lead. Zircon has a very high closure temperature, is resistant to mechanical weathering and is very chemically inert. Zircon also forms multiple crystal layers during metamorphic events, which each may record an isotopic age of

3780-419: Is released, the intensity of which varies depending on the amount of radiation absorbed during burial and specific properties of the mineral. These methods can be used to date the age of a sediment layer, as layers deposited on top would prevent the grains from being "bleached" and reset by sunlight. Pottery shards can be dated to the last time they experienced significant heat, generally when they were fired in

3885-438: Is replete with the mollusks , the vertebrates , the echinoderms , the brachiopods and some groups of arthropods . Fossil sites with exceptional preservation—sometimes including preserved soft tissues—are known as Lagerstätten —German for "storage places". These formations may have resulted from carcass burial in an anoxic environment with minimal bacteria, thus slowing decomposition. Lagerstätten span geological time from

3990-438: Is the Al – Mg chronometer, which can be used to estimate the relative ages of chondrules . Al decays to Mg with a half-life of 720 000 years. The dating is simply a question of finding the deviation from the natural abundance of Mg (the product of Al decay) in comparison with the ratio of

4095-470: Is the difficulty of working out fossil ages. Beds that preserve fossils typically lack the radioactive elements needed for radiometric dating . This technique is our only means of giving rocks greater than about 50 million years old an absolute age, and can be accurate to within 0.5% or better. Although radiometric dating requires careful laboratory work, its basic principle is simple: the rates at which various radioactive elements decay are known, and so

4200-413: Is the science of deciphering the "layer-cake" that is the sedimentary record. Rocks normally form relatively horizontal layers, with each layer younger than the one underneath it. If a fossil is found between two layers whose ages are known, the fossil's age is claimed to lie between the two known ages. Because rock sequences are not continuous, but may be broken up by faults or periods of erosion , it

4305-535: Is the solid foundation of the common measurement of radioactivity. The accuracy and precision of the determination of an age (and a nuclide's half-life) depends on the accuracy and precision of the decay constant measurement. The in-growth method is one way of measuring the decay constant of a system, which involves accumulating daughter nuclides. Unfortunately for nuclides with high decay constants (which are useful for dating very old samples), long periods of time (decades) are required to accumulate enough decay products in

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4410-418: Is used to date old igneous and metamorphic rocks , and has also been used to date lunar samples . Closure temperatures are so high that they are not a concern. Rubidium-strontium dating is not as precise as the uranium–lead method, with errors of 30 to 50 million years for a 3-billion-year-old sample. Application of in situ analysis (Laser-Ablation ICP-MS) within single mineral grains in faults have shown that

4515-467: Is very difficult to match up rock beds that are not directly adjacent. However, fossils of species that survived for a relatively short time can be used to match isolated rocks: this technique is called biostratigraphy . For instance, the conodont Eoplacognathus pseudoplanus has a short range in the Middle Ordovician period. If rocks of unknown age have traces of E. pseudoplanus , they have

4620-801: The Cambrian period to the present . Worldwide, some of the best examples of near-perfect fossilization are the Cambrian Maotianshan Shales and Burgess Shale , the Devonian Hunsrück Slates , the Jurassic Solnhofen Limestone , and the Carboniferous Mazon Creek localities. A fossil is said to be recrystallized when the original skeletal compounds are still present but in a different crystal form, such as from aragonite to calcite . Replacement occurs when

4725-807: The Ludington Formation , and in the Liard River area it is unconformably overlain by the Fort St. John Group . It is laterally equivalent to the Doig Formation and the upper two-thirds of the Montney Formation in the subsurface of the Peace River plains, and to part of the Llama Member of the Sulphur Mountain Formation in west-central and southwestern Alberta. Fossil A fossil (from Classical Latin fossilis , lit.   ' obtained by digging ' )

4830-586: The Phacops fossil record was that the aftermaths of the lens changes, but not the rapidly occurring evolutionary process, were fossilized. This and other data led Stephen Jay Gould and Niles Eldredge to publish their seminal paper on punctuated equilibrium in 1971. Synchrotron X-ray tomographic analysis of early Cambrian bilaterian embryonic microfossils yielded new insights of metazoan evolution at its earliest stages. The tomography technique provides previously unattainable three-dimensional resolution at

4935-569: The Renaissance . Leonardo da Vinci concurred with Aristotle's view that fossils were the remains of ancient life. For example, Leonardo noticed discrepancies with the biblical flood narrative as an explanation for fossil origins: If the Deluge had carried the shells for distances of three and four hundred miles from the sea it would have carried them mixed with various other natural objects all heaped up together; but even at such distances from

5040-602: The Song dynasty during the 11th century, who kept a specific seashell fossil with his own poem engraved on it. In his Dream Pool Essays published in 1088, Song dynasty Chinese scholar-official Shen Kuo hypothesized that marine fossils found in a geological stratum of mountains located hundreds of miles from the Pacific Ocean was evidence that a prehistoric seashore had once existed there and shifted over centuries of time . His observation of petrified bamboos in

5145-493: The biosphere as a consequence of industrialization have also depressed the proportion of carbon-14 by a few percent; in contrast, the amount of carbon-14 was increased by above-ground nuclear bomb tests that were conducted into the early 1960s. Also, an increase in the solar wind or the Earth's magnetic field above the current value would depress the amount of carbon-14 created in the atmosphere. This involves inspection of

5250-402: The law of superposition ) preserved different assemblages of fossils, and that these assemblages succeeded one another in a regular and determinable order. He observed that rocks from distant locations could be correlated based on the fossils they contained. He termed this the principle of faunal succession . This principle became one of Darwin's chief pieces of evidence that biological evolution

5355-403: The thunderbird . There is no such direct mythological connection known from prehistoric Africa, but there is considerable evidence of tribes there excavating and moving fossils to ceremonial sites, apparently treating them with some reverence. In Japan, fossil shark teeth were associated with the mythical tengu , thought to be the razor-sharp claws of the creature, documented some time after

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5460-453: The 16th century. Roman naturalist Pliny the Elder wrote of " tongue stones ", which he called glossopetra . These were fossil shark teeth, thought by some classical cultures to look like the tongues of people or snakes. He also wrote about the horns of Ammon , which are fossil ammonites , whence the group of shelled octopus-cousins ultimately draws its modern name. Pliny also makes one of

5565-739: The 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolute ages of rocks and the fossils they host. There are many processes that lead to fossilization , including permineralization , casts and molds, authigenic mineralization , replacement and recrystallization, adpression, carbonization , and bioimmuration. Fossils vary in size from one- micrometre (1 μm) bacteria to dinosaurs and trees, many meters long and weighing many tons. A fossil normally preserves only

5670-413: The 8th century AD. In medieval China, the fossil bones of ancient mammals including Homo erectus were often mistaken for " dragon bones" and used as medicine and aphrodisiacs . In addition, some of these fossil bones are collected as "art" by scholars, who left scripts on various artifacts, indicating the time they were added to a collection. One good example is the famous scholar Huang Tingjian of

5775-615: The Preservation of Favoured Races in the Struggle for Life , the oldest animal fossils were those from the Cambrian Period, now known to be about 540 million years old. He worried about the absence of older fossils because of the implications on the validity of his theories, but he expressed hope that such fossils would be found, noting that: "only a small portion of the world is known with accuracy." Darwin also pondered

5880-566: The Proterozoic and deeper still in the Archean is only "recounted by microscopic fossils and subtle chemical signals." Molecular biologists, using phylogenetics , can compare protein amino acid or nucleotide sequence homology (i.e., similarity) to evaluate taxonomy and evolutionary distances among organisms, with limited statistical confidence. The study of fossils, on the other hand, can more specifically pinpoint when and in what organism

5985-600: The Rb-Sr method can be used to decipher episodes of fault movement. A relatively short-range dating technique is based on the decay of uranium-234 into thorium-230, a substance with a half-life of about 80,000 years. It is accompanied by a sister process, in which uranium-235 decays into protactinium-231, which has a half-life of 32,760 years. While uranium is water-soluble, thorium and protactinium are not, and so they are selectively precipitated into ocean-floor sediments , from which their ratios are measured. The scheme has

6090-520: The age of the sample even if some of the lead has been lost. This can be seen in the concordia diagram, where the samples plot along an errorchron (straight line) which intersects the concordia curve at the age of the sample. This involves the alpha decay of Sm to Nd with a half-life of 1.06 x 10 years. Accuracy levels of within twenty million years in ages of two-and-a-half billion years are achievable. This involves electron capture or positron decay of potassium-40 to argon-40. Potassium-40 has

6195-444: The ages of the same materials are consistent from one method to another. It is not affected by external factors such as temperature , pressure , chemical environment, or presence of a magnetic or electric field . The only exceptions are nuclides that decay by the process of electron capture, such as beryllium-7 , strontium-85 , and zirconium-89 , whose decay rate may be affected by local electron density. For all other nuclides,

6300-422: The atmosphere is about 1 week. Thus, as an event marker of 1950s water in soil and ground water, Cl is also useful for dating waters less than 50 years before the present. Cl has seen use in other areas of the geological sciences, including dating ice and sediments. Luminescence dating methods are not radiometric dating methods in that they do not rely on abundances of isotopes to calculate age. Instead, they are

6405-413: The beach, indicating the fossils were once living animals. He had previously explained them in terms of vaporous exhalations , which Persian polymath Avicenna modified into the theory of petrifying fluids ( succus lapidificatus ). Recognition of fossil seashells as originating in the sea was built upon in the 14th century by Albert of Saxony , and accepted in some form by most naturalists by

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6510-714: The beginning of recorded history. The fossils themselves are referred to as the fossil record. The fossil record was one of the early sources of data underlying the study of evolution and continues to be relevant to the history of life on Earth . Paleontologists examine the fossil record to understand the process of evolution and the way particular species have evolved. Fossils have been visible and common throughout most of natural history, and so documented human interaction with them goes back as far as recorded history, or earlier. There are many examples of paleolithic stone knives in Europe, with fossil echinoderms set precisely at

6615-430: The best-known techniques are radiocarbon dating , potassium–argon dating and uranium–lead dating . By allowing the establishment of geological timescales, it provides a significant source of information about the ages of fossils and the deduced rates of evolutionary change. Radiometric dating is also used to date archaeological materials, including ancient artifacts. Different methods of radiometric dating vary in

6720-416: The clock to zero. The trapped charge accumulates over time at a rate determined by the amount of background radiation at the location where the sample was buried. Stimulating these mineral grains using either light ( optically stimulated luminescence or infrared stimulated luminescence dating) or heat ( thermoluminescence dating ) causes a luminescence signal to be emitted as the stored unstable electron energy

6825-453: The conversion efficiency from I to Xe . The difference between the measured Xe / Xe ratios of the sample and Shallowater then corresponds to the different ratios of I / I when they each stopped losing xenon. This in turn corresponds to a difference in age of closure in the early solar system. Another example of short-lived extinct radionuclide dating

6930-535: The cups, the ions set up a very weak current that can be measured to determine the rate of impacts and the relative concentrations of different atoms in the beams. Uranium–lead radiometric dating involves using uranium-235 or uranium-238 to date a substance's absolute age. This scheme has been refined to the point that the error margin in dates of rocks can be as low as less than two million years in two-and-a-half billion years. An error margin of 2–5% has been achieved on younger Mesozoic rocks. Uranium–lead dating

7035-472: The decay products of extinct radionuclides with a mass spectrometer and using isochronplots, it is possible to determine relative ages of different events in the early history of the solar system. Dating methods based on extinct radionuclides can also be calibrated with the U–Pb method to give absolute ages. Thus both the approximate age and a high time resolution can be obtained. Generally a shorter half-life leads to

7140-670: The deity Sopdu , the Morning Star, equivalent of Venus in Roman mythology. Fossils appear to have directly contributed to the mythology of many civilizations, including the ancient Greeks. Classical Greek historian Herodotos wrote of an area near Hyperborea where gryphons protected golden treasure. There was indeed gold mining in that approximate region , where beaked Protoceratops skulls were common as fossils. A later Greek scholar, Aristotle , eventually realized that fossil seashells from rocks were similar to those found on

7245-466: The dry northern climate zone of what is now Yan'an , Shaanxi province, China, led him to advance early ideas of gradual climate change due to bamboo naturally growing in wetter climate areas. In medieval Christendom , fossilized sea creatures on mountainsides were seen as proof of the biblical deluge of Noah's Ark . After observing the existence of seashells in mountains, the ancient Greek philosopher Xenophanes (c. 570 – 478 BC) speculated that

7350-415: The earlier known references to toadstones , thought until the 18th century to be a magical cure for poison originating in the heads of toads, but which are fossil teeth from Lepidotes , a Cretaceous ray-finned fish. The Plains tribes of North America are thought to have similarly associated fossils, such as the many intact pterosaur fossils naturally exposed in the region, with their own mythology of

7455-550: The earliest known stromatolites are over 3.4 billion years old. The fossil record is life's evolutionary epic that unfolded over four billion years as environmental conditions and genetic potential interacted in accordance with natural selection. The Virtual Fossil Museum Paleontology has joined with evolutionary biology to share the interdisciplinary task of outlining the tree of life, which inevitably leads backwards in time to Precambrian microscopic life when cell structure and functions evolved. Earth's deep time in

7560-464: The earth during earthquake and subsidences, and petrifies whatever comes into contact with it. As a matter of fact, the petrifaction of the bodies of plants and animals is not more extraordinary than the transformation of waters. From the 13th century to the present day, scholars pointed out that the fossil skulls of Deinotherium giganteum , found in Crete and Greece, might have been interpreted as being

7665-422: The event. In situ micro-beam analysis can be achieved via laser ICP-MS or SIMS techniques. One of its great advantages is that any sample provides two clocks, one based on uranium-235's decay to lead-207 with a half-life of about 700 million years, and one based on uranium-238's decay to lead-206 with a half-life of about 4.5 billion years, providing a built-in crosscheck that allows accurate determination of

7770-478: The existence of a world previous to ours, destroyed by some kind of catastrophe. Interest in fossils, and geology more generally, expanded during the early nineteenth century. In Britain, Mary Anning 's discoveries of fossils, including the first complete ichthyosaur and a complete plesiosaurus skeleton, sparked both public and scholarly interest. Early naturalists well understood the similarities and differences of living species leading Linnaeus to develop

7875-750: The existing isotope decays with a characteristic half-life (5730 years). The proportion of carbon-14 left when the remains of the organism are examined provides an indication of the time elapsed since its death. This makes carbon-14 an ideal dating method to date the age of bones or the remains of an organism. The carbon-14 dating limit lies around 58,000 to 62,000 years. The rate of creation of carbon-14 appears to be roughly constant, as cross-checks of carbon-14 dating with other dating methods show it gives consistent results. However, local eruptions of volcanoes or other events that give off large amounts of carbon dioxide can reduce local concentrations of carbon-14 and give inaccurate dates. The releases of carbon dioxide into

7980-423: The half-life of the parent is accurately known, and enough of the daughter product is produced to be accurately measured and distinguished from the initial amount of the daughter present in the material. The procedures used to isolate and analyze the parent and daughter nuclides must be precise and accurate. This normally involves isotope-ratio mass spectrometry . The precision of a dating method depends in part on

8085-455: The half-life of the radioactive isotope involved. For instance, carbon-14 has a half-life of 5,730 years. After an organism has been dead for 60,000 years, so little carbon-14 is left that accurate dating cannot be established. On the other hand, the concentration of carbon-14 falls off so steeply that the age of relatively young remains can be determined precisely to within a few decades. The closure temperature or blocking temperature represents

8190-519: The hand grip, dating back to Homo heidelbergensis and Neanderthals . These ancient peoples also drilled holes through the center of those round fossil shells, apparently using them as beads for necklaces. The ancient Egyptians gathered fossils of species that resembled the bones of modern species they worshipped. The god Set was associated with the hippopotamus , therefore fossilized bones of hippo-like species were kept in that deity's temples. Five-rayed fossil sea urchin shells were associated with

8295-676: The heavy parent isotopes were produced by nucleosynthesis in supernovas, meaning that any parent isotope with a short half-life should be extinct by now. Carbon-14, though, is continuously created through collisions of neutrons generated by cosmic rays with nitrogen in the upper atmosphere and thus remains at a near-constant level on Earth. The carbon-14 ends up as a trace component in atmospheric carbon dioxide (CO 2 ). A carbon-based life form acquires carbon during its lifetime. Plants acquire it through photosynthesis , and animals acquire it from consumption of plants and other animals. When an organism dies, it ceases to take in new carbon-14, and

8400-456: The isotope systems to be very precisely calibrated, such as the Pb–Pb system . The basic equation of radiometric dating requires that neither the parent nuclide nor the daughter product can enter or leave the material after its formation. The possible confounding effects of contamination of parent and daughter isotopes have to be considered, as do the effects of any loss or gain of such isotopes since

8505-535: The limits of fossilization. Fossils of two enigmatic bilaterians, the worm-like Markuelia and a putative, primitive protostome , Pseudooides , provide a peek at germ layer embryonic development. These 543-million-year-old embryos support the emergence of some aspects of arthropod development earlier than previously thought in the late Proterozoic. The preserved embryos from China and Siberia underwent rapid diagenetic phosphatization resulting in exquisite preservation, including cell structures. This research

8610-425: The plastic film. The uranium content of the material can then be calculated from the number of tracks and the neutron flux . This scheme has application over a wide range of geologic dates. For dates up to a few million years micas , tektites (glass fragments from volcanic eruptions), and meteorites are best used. Older materials can be dated using zircon , apatite , titanite , epidote and garnet which have

8715-399: The present ratios of the parent and daughter isotopes to a standard isotope. An isochron plot is used to solve the age equation graphically and calculate the age of the sample and the original composition. Radiometric dating has been carried out since 1905 when it was invented by Ernest Rutherford as a method by which one might determine the age of the Earth . In the century since then

8820-545: The problems involved in matching rocks of the same age across continents . Family-tree relationships also help to narrow down the date when lineages first appeared. For instance, if fossils of B or C date to X million years ago and the calculated "family tree" says A was an ancestor of B and C, then A must have evolved earlier. It is also possible to estimate how long ago two living clades diverged, in other words approximately how long ago their last common ancestor must have lived, by assuming that DNA mutations accumulate at

8925-411: The proportion of the original nuclide to its decay products changes in a predictable way as the original nuclide decays over time. This predictability allows the relative abundances of related nuclides to be used as a clock to measure the time from the incorporation of the original nuclides into a material to the present. The radioactive decay constant, the probability that an atom will decay per year,

9030-443: The ratio of the radioactive element to its decay products shows how long ago the radioactive element was incorporated into the rock. Radioactive elements are common only in rocks with a volcanic origin, and so the only fossil-bearing rocks that can be dated radiometrically are volcanic ash layers, which may provide termini for the intervening sediments. Consequently, palaeontologists rely on stratigraphy to date fossils. Stratigraphy

9135-462: The relative ages of rock strata as determined by the early paleontologists and stratigraphers . Since the early years of the twentieth century, absolute dating methods, such as radiometric dating (including potassium/argon , argon/argon , uranium series , and, for very recent fossils, radiocarbon dating ) have been used to verify the relative ages obtained by fossils and to provide absolute ages for many fossils. Radiometric dating has shown that

9240-453: The reservoir when they formed, they should form an isochron . This can reduce the problem of contamination . In uranium–lead dating , the concordia diagram is used which also decreases the problem of nuclide loss. Finally, correlation between different isotopic dating methods may be required to confirm the age of a sample. For example, the age of the Amitsoq gneisses from western Greenland

9345-436: The sample was created. It is therefore essential to have as much information as possible about the material being dated and to check for possible signs of alteration . Precision is enhanced if measurements are taken on multiple samples from different locations of the rock body. Alternatively, if several different minerals can be dated from the same sample and are assumed to be formed by the same event and were in equilibrium with

9450-429: The sea and that they were still living when the strait of Gibraltar was cut through. In the mountains of Parma and Piacenza multitudes of shells and corals with holes may be seen still sticking to the rocks.... In 1666, Nicholas Steno examined a shark, and made the association of its teeth with the "tongue stones" of ancient Greco-Roman mythology, concluding that those were not in fact the tongues of venomous snakes, but

9555-423: The sea we see the oysters all together and also the shellfish and the cuttlefish and all the other shells which congregate together, found all together dead; and the solitary shells are found apart from one another as we see them every day on the sea-shores. And we find oysters together in very large families, among which some may be seen with their shells still joined together, indicating that they were left there by

9660-533: The shell, bone, or other tissue is replaced with another mineral. In some cases mineral replacement of the original shell occurs so gradually and at such fine scales that microstructural features are preserved despite the total loss of original material. Scientists can use such fossils when researching the anatomical structure of ancient species. Several species of saurids have been identified from mineralized dinosaur fossils. Radiometric dating Radiometric dating , radioactive dating or radioisotope dating

9765-586: The skulls of the Cyclopes of Greek mythology , and are possibly the origin of that Greek myth. Their skulls appear to have a single eye-hole in the front, just like their modern elephant cousins, though in fact it's actually the opening for their trunk. In Norse mythology , echinoderm shells (the round five-part button left over from a sea urchin) were associated with the god Thor , not only being incorporated in thunderstones , representations of Thor's hammer and subsequent hammer-shaped crosses as Christianity

9870-505: The stable isotopes Al / Mg . The excess of Mg (often designated Mg *) is found by comparing the Mg / Mg ratio to that of other Solar System materials. The Al – Mg chronometer gives an estimate of the time period for formation of primitive meteorites of only a few million years (1.4 million years for Chondrule formation). In

9975-467: The sudden appearance of many groups (i.e. phyla ) in the oldest known Cambrian fossiliferous strata. Since Darwin's time, the fossil record has been extended to between 2.3 and 3.5 billion years. Most of these Precambrian fossils are microscopic bacteria or microfossils . However, macroscopic fossils are now known from the late Proterozoic. The Ediacara biota (also called Vendian biota) dating from 575 million years ago collectively constitutes

10080-520: The techniques have been greatly improved and expanded. Dating can now be performed on samples as small as a nanogram using a mass spectrometer . The mass spectrometer was invented in the 1940s and began to be used in radiometric dating in the 1950s. It operates by generating a beam of ionized atoms from the sample under test. The ions then travel through a magnetic field, which diverts them into different sampling sensors, known as " Faraday cups ," depending on their mass and level of ionization. On impact in

10185-451: The teeth of some long-extinct species of shark. Robert Hooke (1635–1703) included micrographs of fossils in his Micrographia and was among the first to observe fossil forams . His observations on fossils, which he stated to be the petrified remains of creatures some of which no longer existed, were published posthumously in 1705. William Smith (1769–1839) , an English canal engineer, observed that rocks of different ages (based on

10290-413: The temperature below which the mineral is a closed system for the studied isotopes. If a material that selectively rejects the daughter nuclide is heated above this temperature, any daughter nuclides that have been accumulated over time will be lost through diffusion , resetting the isotopic "clock" to zero. As the mineral cools, the crystal structure begins to form and diffusion of isotopes is less easy. At

10395-428: The timescale over which they are accurate and the materials to which they can be applied. All ordinary matter is made up of combinations of chemical elements , each with its own atomic number , indicating the number of protons in the atomic nucleus . Additionally, elements may exist in different isotopes , with each isotope of an element differing in the number of neutrons in the nucleus. A particular isotope of

10500-500: The ultimate transformation of the radioactive nuclide into its stable daughter. Isotopic systems that have been exploited for radiometric dating have half-lives ranging from only about 10 years (e.g., tritium ) to over 100 billion years (e.g., samarium-147 ). For most radioactive nuclides, the half-life depends solely on nuclear properties and is essentially constant. This is known because decay constants measured by different techniques give consistent values within analytical errors and

10605-524: The world was once inundated in a great flood that buried living creatures in drying mud. In 1027, the Persian Avicenna explained fossils' stoniness in The Book of Healing : If what is said concerning the petrifaction of animals and plants is true, the cause of this (phenomenon) is a powerful mineralizing and petrifying virtue which arises in certain stony spots, or emanates suddenly from

10710-458: Was adopted, but also kept in houses to garner Thor's protection. These grew into the shepherd's crowns of English folklore, used for decoration and as good luck charms, placed by the doorway of homes and churches. In Suffolk , a different species was used as a good-luck charm by bakers, who referred to them as fairy loaves , associating them with the similarly shaped loaves of bread they baked. More scientific views of fossils emerged during

10815-443: Was determined to be 3.60 ± 0.05 Ga (billion years ago) using uranium–lead dating and 3.56 ± 0.10 Ga (billion years ago) using lead–lead dating, results that are consistent with each other. Accurate radiometric dating generally requires that the parent has a long enough half-life that it will be present in significant amounts at the time of measurement (except as described below under "Dating with short-lived extinct radionuclides"),

10920-408: Was real. Georges Cuvier came to believe that most if not all the animal fossils he examined were remains of extinct species. This led Cuvier to become an active proponent of the geological school of thought called catastrophism . Near the end of his 1796 paper on living and fossil elephants he said: All of these facts, consistent among themselves, and not opposed by any report, seem to me to prove

11025-454: Was replaced by the Doig and Montney Formations by J.H. Armitage in 1962, and the names are now considered obsolete, although their usage persists. The Grayling Formation consists of dolomitic siltstone and silty shale , with minor silty limestone , dolomite , and very fine-grained sandstone . It reaches a maximum thickness of about 460 metres (1500 ft). The overlying Toad Formation

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