131-645: And see text Phacops is a genus of trilobites in the order Phacopida , family Phacopidae , that lived in Europe, northwestern Africa, North and South America and China from the Late Ordovician until the very end of the Devonian , with a broader time range described from the Late Ordovician . It was a rounded animal, with a globose head and large eyes, and probably fed on detritus . Phacops
262-467: A balanced budget, some tax reforms , a labor-market reform, scheduled nuclear phase-out and instigated legislation allowing more stringent war crime and more lenient soft drug usage prosecution. Restrictions on euthanasia were reduced, and in 2003, Belgium became one of the first countries in the world to legalise same-sex marriage . The government promoted active diplomacy in Africa and opposed
393-557: A clade called Arachnomorpha , while others consider them to be more closely related to Mandibulata (which contains insects , crustaceans and myriapods ) as part of a clade called Antennulata . The earliest trilobites known from the fossil record are redlichiids and ptychopariid bigotinids dated to around 520 million years ago. Contenders for the earliest trilobites include Profallotaspis jakutensis (Siberia), Fritzaspis spp. (western USA), Hupetina antiqua (Morocco) and Serrania gordaensis (Spain). Trilobites appeared at
524-860: A decade after the other regions) is included in both the Flemish and French Communities, and the territory of the German-speaking Community lies wholly within the Walloon Region. Conflicts about jurisdiction between the bodies are resolved by the Constitutional Court of Belgium . The structure is intended as a compromise to allow different cultures to live together peacefully. The Federal State's authority includes justice, defense, federal police, social security, nuclear energy, monetary policy and public debt, and other aspects of public finances. State-owned companies include
655-686: A decrease in the number of lenses, which is a more widespread and recurring trend in many Phacopinae . Fossils of Phacops salteri have been found in the trilobite-rich Late Emsian to Early Givetian Floresta Formation of the Altiplano Cundiboyacense , Colombia. Phacops currently contains the following species: A number of species previously assigned to the genus Phacops have since been transferred to other genera: Trilobite Trilobites ( / ˈ t r aɪ l ə ˌ b aɪ t s , ˈ t r ɪ l ə -/ ; meaning "three lobes") are extinct marine arthropods that form
786-693: A large semi-independent enclave. The Eighty Years' War (1568–1648) was triggered by the Spanish government's policy towards Protestantism , which was becoming popular in the Low Countries. The rebellious northern United Provinces ( Belgica Foederata in Latin , the "Federated Netherlands") eventually separated from the Southern Netherlands ( Belgica Regia , the "Royal Netherlands"). The southern part continued to be ruled successively by
917-526: A lattice of chitin , and is curled round the lower edge to produce a small fringe called the "doublure". Their appendages and soft underbelly were non-mineralized. Three distinctive tagmata (sections) are present: cephalon (head); thorax (body) and pygidium (tail). As might be expected for a group of animals comprising c. 5,000 genera, the morphology and description of trilobites can be complex. Despite morphological complexity and an unclear position within higher classifications, there are
1048-519: A natant (unattached) hypostome . The most recently recognized of the nine trilobite orders, Harpetida, was erected in 2002. The progenitor of order Phacopida is unclear. When trilobites are found, only the exoskeleton is preserved (often in an incomplete state) in all but a handful of locations. A few locations ( Lagerstätten ) preserve identifiable soft body parts (legs, gills, musculature & digestive tract) and enigmatic traces of other structures (e.g. fine details of eye structure) as well as
1179-552: A new government . By 30 March 2011, this set a new world record for the elapsed time without an official government, previously held by war-torn Iraq . Finally, in December 2011 the Di Rupo Government led by Walloon socialist Prime Minister Elio Di Rupo was sworn in. The 2014 federal election (coinciding with the regional elections ) resulted in a further electoral gain for the Flemish nationalist N-VA, although
1310-427: A new order, Eodiscida. Over 20,000 species of trilobite have been described. Despite their rich fossil record with thousands of described genera found throughout the world, the taxonomy and phylogeny of trilobites have many uncertainties. Except possibly for the members of the orders Phacopida and Lichida (which first appear during the early Ordovician ), nine of the eleven trilobite orders appear prior to
1441-430: A number of characteristics which distinguish the trilobites from other arthropods: a generally sub-elliptical, dorsal , chitinous exoskeleton divided longitudinally into three distinct lobes (from which the group gets its name); having a distinct, relatively large head shield (cephalon) articulating axially with a thorax comprising articulated transverse segments, the hindmost of which are almost invariably fused to form
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#17327729982201572-986: A policy of mediation. The Belgians have been strong advocates of European integration . The headquarters of NATO and of several of the institutions of the European Union are located in Belgium. The Belgian Armed Forces had 23,200 active personnel in 2023, including 8,500 in the Land Component , 1,400 in the Naval Component , 4,900 in the Air Component , 1,450 in the Medical Component , and 6,950 in joint service, in addition to 5,900 reserve personnel. In 2019, Belgium's defense budget totaled €4.303 billion ($ 4.921 billion) representing .93% of its GDP. The operational commands of
1703-408: A result, Boeckops , Chotecops , Paciphacops , Prokops and Viaphacops have been erected as subgenera of Phacops , and are now widely regarded as genera in their own right. Most recent, it was considered that some North-American and North-African species on the one hand and European species on the other hand differ sufficiently from each other to be assigned to separate genera. As the type species
1834-640: A roughly equivalent time in Laurentia , Siberia and West Gondwana . All Olenellina lack facial sutures (see below ), and this is thought to represent the original state. The earliest sutured trilobite found so far ( Lemdadella ), occurs almost at the same time as the earliest Olenellina, suggesting the trilobites origin lies before the start of the Atdabanian, but without leaving fossils. Other groups show secondary lost facial sutures, such as all Agnostina and some Phacopina . Another common feature of
1965-471: A tail shield ( pygidium ). When describing differences between trilobite taxa , the presence, size, and shape of the cephalic features are often mentioned. During moulting , the exoskeleton generally splits between the head and thorax, which is why so many trilobite fossils are missing one or the other. In most groups facial sutures on the cephalon helped facilitate moulting. Similar to lobsters and crabs , trilobites would have physically "grown" between
2096-475: Is Brussels ; other major cities are Antwerp , Ghent , Charleroi , Liège , Bruges , Namur , and Leuven . Belgium is a sovereign state and a federal constitutional monarchy with a parliamentary system . Its institutional organization is complex and is structured on both regional and linguistic grounds. It is divided into three highly autonomous regions : the Flemish Region (Flanders) in
2227-573: Is 72 cm (28 in) in length. It was found in 1998 by Canadian scientists in Ordovician rocks on the shores of Hudson Bay . However, a partial specimen of the Ordovician trilobite Hungioides bohemicus found in 2009 in Arouca , Portugal is estimated to have measured when complete 86.5 cm (34.1 in) in length. Only the upper (dorsal) part of their exoskeleton is mineralized, composed of calcite and calcium phosphate minerals in
2358-465: Is a strong indication that novel morphologies were developing very rapidly. Changes within the trilobite fauna during the Ordovician foreshadowed the mass extinction at the end of the Ordovician, allowing many families to continue into the Silurian with little disturbance. Ordovician trilobites were successful at exploiting new environments, notably reefs . The Ordovician mass extinction did not leave
2489-440: Is around 23% of the total land area, equivalent to 689,300 hectares (ha) of forest in 2020, up from 677,400 hectares (ha) in 1990. In 2020, naturally regenerating forest covered 251,200 hectares (ha) and planted forest covered 438,200 hectares (ha). For the year 2015, 47% of the forest area was reported to be under public ownership , 53% private ownership and 0% with ownership listed as other or unknown. The territory of Belgium
2620-483: Is based on civil law and originates from the Napoleonic code . The Court of Cassation is the court of last resort, with the courts of appeal one level below. Belgium's political institutions are complex; most political power rests on representation of the main cultural communities . Since about 1970, the significant national Belgian political parties have split into distinct components that mainly represent
2751-426: Is based on the use of trilobite marker fossils. Trilobites are the state fossils of Ohio ( Isotelus ), Wisconsin ( Calymene celebra ) and Pennsylvania ( Phacops rana ). The 10 most commonly recognized trilobite orders are Agnostida , Redlichiida , Corynexochida , Lichida , Odontopleurida , Phacopida , Proetida , Asaphida , Harpetida and Ptychopariida . In 2020, an 11th order, Trinucleida ,
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#17327729982202882-575: Is best known from these samples preserved similarly to bodies in Pompeii. The French palaeontologist Joachim Barrande (1799–1883) carried out his landmark study of trilobites in the Cambrian, Ordovician and Silurian of Bohemia , publishing the first volume of Système silurien du centre de la Bohême in 1852. The study of Paleozoic trilobites in the Welsh-English borders by Niles Eldredge
3013-505: Is divided into three Regions, two of which, the Flemish Region and Walloon Region , are in turn subdivided into provinces ; the third Region, the Brussels Capital Region , is neither a province nor a part of a province. Belgium is a constitutional , popular monarchy and a federal parliamentary democracy . The bicameral federal parliament is composed of a Senate and a Chamber of Representatives . The former
3144-429: Is foreshadowed. Some of the genera of Trilobites appearing in the Ordovician include: Most Early Silurian families constitute a subgroup of the Late Ordovician fauna. Few, if any, of the dominant Early Ordovician fauna survived to the end of the Ordovician, yet 74% of the dominant Late Ordovician trilobite fauna survived the Ordovician. Late Ordovician survivors account for all post-Ordovician trilobite groups except
3275-1830: Is found in the Silurian Wenlock Group . This trilobite is featured on the town's coat of arms and was named the Dudley Bug or Dudley Locust by quarrymen who once worked the now abandoned limestone quarries. Llandrindod Wells , Powys , Wales , is another famous trilobite location. The well-known Elrathia kingi trilobite is found in abundance in the Cambrian Wheeler Shale of Utah . Spectacularly preserved trilobite fossils, often showing soft body parts (legs, gills, antennae, etc.) have been found in British Columbia , Canada (the Cambrian Burgess Shale and similar localities); New York , U.S.A. (Ordovician Walcott–Rust quarry , near Russia , and Beecher's Trilobite Bed , near Rome ); China (Lower Cambrian Maotianshan Shales near Chengjiang ); Germany (the Devonian Hunsrück Slates near Bundenbach ) and, much more rarely, in trilobite-bearing strata in Utah (Wheeler Shale and other formations), Ontario , and Manuels River, Newfoundland and Labrador . Sites in Morocco also yield very well-preserved trilobites, many buried in mudslides alive and so perfectly preserved. An industry has developed around their recovery, leading to controversies about practices in restoral. The variety of eye and upper body forms and fragile protuberances
3406-417: Is in the frontal half of the cephalon a continuous furrow, delineated by ridges, and with notches laterally. This so-called vindicular furrow serves to lock the rim of the tailshield to the headshield when the trilobite is enrolled. The axial rings of the thorax do not have convex lateral axial nodes on its outer surface. The tailshield (or pygidium ) is well segmented. The pygidial axis has 9 to 11 rings, and
3537-667: Is lowest in January at 3 °C (37.4 °F) and highest in July at 18 °C (64.4 °F). The average precipitation per month varies between 54 mm (2.1 in) for February and April, to 78 mm (3.1 in) for July. Averages for the years 2000 to 2006 show daily temperature minimums of 7 °C (44.6 °F) and maximums of 14 °C (57.2 °F) and monthly rainfall of 74 mm (2.9 in); these are about 1 °C and nearly 10 millimeters above last century's normal values, respectively. Phytogeographically , Belgium
3668-497: Is made up of 50 senators appointed by the parliaments of the communities and regions and 10 co-opted senators . Prior to 2014, most of the Senate's members were directly elected. The Chamber 's 150 representatives are elected under a proportional voting system from 11 electoral districts . Belgium has compulsory voting and thus maintains one of the highest rates of voter turnout in the world. The King (currently Philippe )
3799-668: Is more than double that of Flanders, which boomed after the Second World War. Belgium is a developed country , with an advanced high-income economy . The country is one of the six founding members of the European Union , and its capital, Brussels, is the de facto capital of the European Union itself, hosting the official seats of the European Commission , the Council of the European Union , and
3930-517: Is no fold right behind the posterior vertical row of lenses, nor an isolated raised area just below the lenses. Not all of these characters may differentiate between Eldredgeops and other Phacops species however. During the Eifelian in the present-day Belgian Ardennes , several Phacops species developed from each other, the oldest being P. imitator , followed by P. fragosus , then P. latifrons and finally P. sartenaeri . These species show
4061-474: Is no surprise that trilobite evolutionary history is marked by a number of extinction events where some groups perished, and surviving groups diversified to fill ecological niches with comparable or unique adaptations. Generally, trilobites maintained high diversity levels throughout the Cambrian and Ordovician periods before entering a drawn-out decline in the Devonian , culminating in the final extinction of
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4192-632: Is now the Benelux, the so-called Burgundian Netherlands . "Burgundy" and "Flanders" were the first two common names used for the Burgundian Netherlands which was the predecessor of the Austrian Netherlands, the predecessor of modern Belgium. The union, technically stretching between two kingdoms, gave the area economic and political stability which led to an even greater prosperity and artistic creation. Born in Belgium,
4323-540: Is often found rolled up (" volvation "), a biological defense mechanism that is widespread among smaller trilobites but further perfected in this genus. Like in all sighted Phacopina , the eyes of Phacops are compounded of very large, separately set lenses without a common cornea (so called schizochroal eyes), and like almost all other Phacopina, the articulate mid-length part of the body (or thorax ) in Phacops has 11 segments. The central raised area (or glabella ) of
4454-428: Is recorded at the same time as the extinctions, suggesting major environmental upheaval. Notable trilobite genera appearing in the Cambrian include: The Early Ordovician is marked by vigorous radiations of articulate brachiopods, bryozoans, bivalves, echinoderms, and graptolites, with many groups appearing in the fossil record for the first time. Although intra-species trilobite diversity seems to have peaked during
4585-729: Is shared between the Atlantic European and Central European provinces of the Circumboreal Region within the Boreal Kingdom . According to the World Wide Fund for Nature , the territory of Belgium belongs to the terrestrial ecoregions of Atlantic mixed forests and Western European broadleaf forests . Belgium had a 2018 Forest Landscape Integrity Index mean score of 1.36/10, ranking it 163rd globally out of 172 countries. In Belgium forest cover
4716-631: Is the head of state , though with limited prerogatives . He appoints ministers, including a Prime Minister, that have the confidence of the Chamber of Representatives to form the federal government . The Council of Ministers is composed of no more than fifteen members. With the possible exception of the Prime Minister, the Council of Ministers is composed of an equal number of Dutch-speaking members and French-speaking members. The judicial system
4847-583: Is the European P. latifrons , the North-American species are now called Eldredgeops . However, the previous assigned species, like Phacops rana , are still widely used among fossil collectors. Eldredgeops has a raised ridge along the ventral margin of the cephalon, the glabella is more inflated, the lateral parts of the preoccipital ring are not round but rectangular, the palpebral area and palpebral lobe are larger than in P. latifrons , and there
4978-404: Is weakly divided into a strongly convex central lobe and weakly convex lateral lobes. The large to medium size eyes have a crescent shaped outline, and are elevated high above the cheeks. The steep visual surface is kidney shaped. The back corners of the cephalon (or genal angles) are acutely to bluntly rounded, but a genal spine is lacking in adults. In the ventral surface of the seam (or doublure)
5109-517: The Artiopoda , a group of extinct arthropods morphologically similar to trilobites, though only the trilobites had mineralised exoskeletons. Thus, other artiopodans are typically only found in exceptionally preserved deposits, mostly during the Cambrian period. The exact relationships of artiopods to other arthropods is uncertain. They have been considered closely related to chelicerates (which include horseshoe crabs and arachnids ) as part of
5240-500: The Belgian Post Group and Belgian Railways . The Federal Government is responsible for the obligations of Belgium and its federalized institutions towards the European Union and NATO. It controls substantial parts of public health, home affairs and foreign affairs. The budget—without the debt—controlled by the federal government amounts to about 50% of the national fiscal income. The federal government employs around 12% of
5371-715: The European Council , as well as one of two seats of the European Parliament (the other being Strasbourg ). Belgium is also a founding member of the Eurozone , NATO , OECD , and WTO , and a part of the trilateral Benelux Union and the Schengen Area . Brussels also hosts the headquarters of many major international organizations, such as NATO . According to Julius Caesar , the Belgae were
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5502-758: The Habsburg Emperor Charles ;V was heir of the Burgundians, but also of the royal families of Austria , Castile and Aragon . With the Pragmatic Sanction of 1549 he gave the Seventeen Provinces more legitimacy as a stable entity, rather than just a temporary personal union . He also increased the influence of these Netherlands over the Prince-Bishopric of Liège , which continued to exist as
5633-548: The Harpetida . Silurian and Devonian trilobite assemblages are superficially similar to Ordovician assemblages, dominated by Lichida and Phacopida (including the well-known Calymenina ). A number of characteristic forms do not extend far into the Devonian and almost all the remainder were wiped out by a series of dramatic Middle and Late Devonian extinctions . Three orders and all but five families were exterminated by
5764-537: The Holy Roman Empire . The lordships and bishoprics along the "March" (frontier) between the two great kingdoms maintained important connections between each other. For example, the county of Flanders expanded over the Scheldt into the empire, and during several periods was ruled by the same lords as the county of Hainaut . In the 13th and 14th centuries, the cloth industry and commerce boomed especially in
5895-724: The Industrial Revolution , and during the course of the 20th century, possessed a number of colonies , notably the Belgian Congo and Ruanda-Urundi . These colonies gained independence between 1960 and 1962. The second half of the 20th century was marked by rising tensions between the Dutch-speakers and French-speakers, fueled by differences in political culture and the unequal economic development of Flanders and Wallonia. This continuing antagonism has led to several far-reaching state reforms , resulting in
6026-822: The Michel Government . It was the first time N-VA was part of the federal cabinet, while the French-speaking side was represented only by the MR, which achieved a minority of the public votes in Wallonia. In May 2019 federal elections in the Flemish-speaking northern region of Flanders far-right Vlaams Belang party made major gains. In the French-speaking southern area of Wallonia the Socialists were strong. The moderate Flemish nationalist party
6157-471: The Netherlands ( 450 km ). Its total surface, including water area, is 30,689 km (11,849 sq mi). Before 2018, its total area was believed to be 30,528 km (11,787 sq mi). However, when the country's statistics were measured in 2018, a new calculation method was used. Unlike previous calculations, this one included the area from the coast to the low-water line, revealing
6288-621: The Permian (when the vast majority of species on Earth were wiped out ). It is unknown why the order Proetida alone survived the Devonian. The Proetida maintained relatively diverse faunas in both deep and shallow water shelf environments throughout the Carboniferous. For many millions of years the Proetida existed untroubled in their ecological niche . An analogy would be today's crinoids , which mostly exist as deep-water species; in
6419-543: The Precambrian this is no longer supported, and it is thought that trilobites originated shortly before they appeared in the fossil record. Very shortly after trilobite fossils appeared in the lower Cambrian, they rapidly diversified into the major orders that typified the Cambrian— Redlichiida , Ptychopariida , Agnostida , and Corynexochida . The first major crisis in the trilobite fossil record occurred in
6550-796: The Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized mineralised exoskeleton , they left an extensive fossil record. The study of their fossils has facilitated important contributions to biostratigraphy , paleontology , evolutionary biology , and plate tectonics . Trilobites are placed within
6681-759: The Spanish ( Spanish Netherlands ) and the Austrian House of Habsburgs ( Austrian Netherlands ) and comprised most of modern Belgium. This was the theatre of several more protracted conflicts during much of the 17th and 18th centuries involving France, including the Franco-Dutch War (1672–1678), the Nine Years' War (1688–1697), the War of the Spanish Succession (1701–1714), and part of
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#17327729982206812-429: The United Kingdom of the Netherlands occurred at the dissolution of the First French Empire in 1814, after the abdication of Napoleon. In 1830, the Belgian Revolution led to the re-separation of the Southern Provinces from the Netherlands and to the establishment of a Catholic and bourgeois, officially French-speaking and neutral, independent Belgium under a provisional government and a national congress . Since
6943-399: The War of the Austrian Succession (1740–1748). Following the campaigns of 1794 in the French Revolutionary Wars , the Low Countries – including territories that were never nominally under Habsburg rule, such as the Prince-Bishopric of Liège – were annexed by the French First Republic , ending Austrian rule in the region. A reunification of the Low Countries as
7074-425: The Western Roman Empire , the Roman provinces of Belgica and Germania were inhabited by a mix of a Romanized population and Germanic -speaking Franks who came to dominate the military and political class. During the 5th century, the area came under the rule of the Frankish Merovingian kings, who initially established a kingdom ruling over the Romanized population in what is now northern France, and then conquered
7205-402: The class Trilobita . Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period ( 521 million years ago ) and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian , all trilobite orders except
7336-406: The labrum in well-preserved trilobite specimens from Cambrian Stage 4 of Morocco, providing new anatomical information regarding the external and internal morphology of trilobites, and the cause of such extraordinary preservation is probably due to their rapid death after an underwater pyroclastic flow. Trilobites saw great diversification over time. For such a long-lasting group of animals, it
7467-419: The taxonomy and phylogeny of trilobites. The dorsal surface of the trilobite cephalon (the frontmost tagma , or the 'head') can be divided into two regions—the cranidium and the librigena ("free cheeks"). The cranidium can be further divided into the glabella (the central lobe in the cephalon) and the fixigena ("fixed cheeks"). The facial sutures lie along the anterior edge, at the division between
7598-436: The 1970s by Dan Cooper. As a well-known rock collector, he incited scientific and public interest in the location. The fossils are dated to the Givetian (387.2 - 382.7 million years ago) when the Western New York Region was 30 degrees south of the equator and completely covered in water. The site was purchased from Vincent C. Bonerb by the Town of Hamburg with the cooperation of the Hamburg Natural History Society to protect
7729-422: The 19th century were the Catholic Party and the Liberal Party , with the Belgian Labour Party emerging towards the end of the 19th century. French was originally the official language used by the nobility and the bourgeoisie , especially after the rejection of the Dutch monarchy. French progressively lost its dominance as Dutch began to recover its status. This recognition became official in 1898, and in 1967,
7860-474: The Ardennes are more rugged and rocky with caves and small gorges . Extending westward into France, this area is eastwardly connected to the Eifel in Germany by the High Fens plateau, on which the Signal de Botrange forms the country's highest point at 694 m (2,277 ft). The climate is maritime temperate with significant precipitation in all seasons ( Köppen climate classification : Cfb ), like most of northwest Europe. The average temperature
7991-428: The Belgian state to assume responsibility for the government of the colony, henceforth called the Belgian Congo . A Belgian commission in 1919 estimated that Congo's population was half what it was in 1879. Germany invaded Belgium in August 1914 as part of the Schlieffen Plan to attack France , and much of the Western Front fighting of World War I occurred in western parts of the country. The opening months of
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#17327729982208122-402: The Cambrian, trilobites were still active participants in the Ordovician radiation event, with a new fauna taking over from the old Cambrian one. Phacopida and Trinucleioidea are characteristic forms, highly differentiated and diverse, most with uncertain ancestors. The Phacopida and other "new" clades almost certainly had Cambrian forebears, but the fact that they have avoided detection
8253-428: The Communities and Regions were created in 1980, Flemish politicians decided to merge both. Thus the Flemings just have one single institutional body of parliament and government is empowered for all except federal and specific municipal matters. The overlapping boundaries of the Regions and Communities have created two notable peculiarities: the territory of the Brussels-Capital Region (which came into existence nearly
8384-490: The County of Flanders and it became one of the richest areas in Europe. This prosperity played a role in conflicts between Flanders and the king of France . Famously, Flemish militias scored a surprise victory at the Battle of the Golden Spurs against a strong force of mounted knights in 1302, but France soon regained control of the rebellious province. In the 15th century, the Duke of Burgundy in France took control of Flanders, and from there proceeded to unite much of what
8515-548: The European Union, for which Belgium hosts major administrations and institutions, including the European Commission , the Council of the European Union and the extraordinary and committee sessions of the European Parliament . In the early 1990s, Belgium saw several large corruption scandals notably surrounding Marc Dutroux , Andre Cools , the Dioxin Affair , Agusta Scandal and the murder of Karel van Noppen . Belgium shares borders with France ( 620 km ), Germany ( 162/167 km ), Luxembourg ( 148 km ), and
8646-491: The Flemings refused to do so and succeeded progressively in making Dutch an equal language in the education system. Following World War II, Belgian politics became increasingly dominated by the autonomy of its two main linguistic communities. Intercommunal tensions rose and the constitution was amended to minimize the potential for conflict. Based on the four language areas defined in 1962–63 (the Dutch, bilingual, French and German language areas), consecutive revisions of
8777-400: The Latin name of a large Roman province covering most of Northern Gaul, including the Belgae and Treveri. However, areas closer to the lower Rhine frontier, including the eastern part of modern Belgium, subsequently became part of the frontier province of Germania Inferior , which continued to interact with their neighbours outside the empire. At the time when central government collapsed in
8908-421: The Middle Cambrian ; surviving orders developed isopygius or macropygius bodies and developed thicker cuticles, allowing better defense against predators (see Thorax below). The end- Cambrian mass extinction event marked a major change in trilobite fauna; almost all Redlichiida (including the Olenelloidea) and most Late Cambrian stocks became extinct. A continuing decrease in Laurentian continental shelf area
9039-458: The N-VA remained the largest party in parliament. In July 2019 prime minister Charles Michel was selected to hold the post of President of the European Council . His successor Sophie Wilmès was Belgium's first female prime minister. She led the caretaker government since October 2019. The Flemish Liberal party politician Alexander De Croo became new prime minister in October 2020. The parties had agreed on federal government 16 months after
9170-515: The Olenellina also suggests this suborder to be the ancestral trilobite stock: early protaspid stages have not been found, supposedly because these were not calcified, and this also is supposed to represent the original state. Earlier trilobites may be found and could shed more light on their origins. Three specimens of a trilobite from Morocco, Megistaspis hammondi , dated 478 million years old contain fossilized soft parts. In 2024, researchers discovered soft tissues and other structures including
9301-411: The Paleozoic era, vast 'forests' of crinoids lived in shallow near-shore environments. Some of the genera of trilobites during the Carboniferous and Permian periods include: Exactly why the trilobites became extinct is not clear; with repeated extinction events (often followed by apparent recovery) throughout the trilobite fossil record, a combination of causes is likely. After the extinction event at
9432-570: The Redlichiida or Corynexochida in the Middle Cambrian. Order Ptychopariida is the most problematic order for trilobite classification. In the 1959 Treatise on Invertebrate Paleontology , what are now members of orders Ptychopariida, Asaphida , Proetida and Harpetida were grouped together as order Ptychopariida; subclass Librostoma was erected in 1990 to encompass all of these orders, based on their shared ancestral character of
9563-600: The Regions' and Communities' Governments is the broadest of all the Federating units of all the Federations all over the world. Because of its location at the crossroads of Western Europe , Belgium has historically been the route of invading armies from its larger neighbors. With virtually defenseless borders, Belgium has traditionally sought to avoid domination by the more powerful nations which surround it through
9694-521: The age of the rocks in which they are found. They were among the first fossils to attract widespread attention, and new species are being discovered every year. In the United States, the best open-to-the-public collection of trilobites is located in Hamburg, New York . The shale quarry, informally known as Penn Dixie, stopped mining in the 1960s. The large amounts of trilobites were discovered in
9825-399: The cabinet to the king, as no progress in constitutional reforms had been made. In December 2008, Leterme once more offered his resignation after a crisis surrounding the sale of Fortis to BNP Paribas . At this juncture, his resignation was accepted and Christian Democratic and Flemish Herman Van Rompuy was sworn in as Prime Minister on 30 December 2008. After Herman Van Rompuy
9956-731: The civil servants. Communities exercise their authority only within linguistically determined geographical boundaries, originally oriented towards the individuals of a Community's language: culture (including audiovisual media), education and the use of the relevant language. Extensions to personal matters less directly connected with language comprise health policy (curative and preventive medicine) and assistance to individuals (protection of youth, social welfare, aid to families, immigrant assistance services, and so on.). Regions have authority in fields that can be broadly associated with their territory. These include economy, employment, agriculture, water policy, housing, public works, energy, transport,
10087-473: The clade Artiopoda , which includes many organisms that are morphologically similar to trilobites, but are largely unmineralised. The relationship of Artiopoda to other arthropods is uncertain. Trilobites evolved into many ecological niches; some moved over the seabed as predators , scavengers , or filter feeders , and some swam, feeding on plankton . Some even crawled onto land. Most lifestyles expected of modern marine arthropods are seen in trilobites, with
10218-518: The combination of sea level changes and a break in the redox equilibrium (a meteorite impact has also been suggested as a cause). Only a single order, the Proetida , survived into the Carboniferous. Genera of trilobites during the Silurian and Devonian periods include: The Proetida survived for millions of years, continued through the Carboniferous period and lasted until the end of
10349-574: The country in May 1940 , and 40,690 Belgians, over half of them Jews, were killed during the subsequent occupation and the Holocaust . From September 1944 to February 1945 the Allies liberated Belgium. After World War II , a general strike forced King Leopold III to abdicate in 1951 in favour of his son, Prince Baudouin , since many Belgians thought he had collaborated with Germany during
10480-535: The country to be 160 km (62 sq mi) larger in surface area than previously thought. Its land area alone is 30,494 square kilometers. It lies between latitudes 49°30' and 51°30' N, and longitudes 2°33' and 6°24' E. Belgium has three main geographical regions; the coastal plain in the northwest and the central plateau both belong to the Anglo-Belgian Basin, and the Ardennes uplands in
10611-407: The country's constitution in 1970, 1980, 1988 and 1993 established a unique form of a federal state with segregated political power into three levels: The constitutional language areas determine the official languages in their municipalities, as well as the geographical limits of the empowered institutions for specific matters. Although this would allow for seven parliaments and governments when
10742-644: The cranidium and the librigena. Belgium – in Europe (light green & dark grey) – in the European Union (light green) Belgium , officially the Kingdom of Belgium , is a country in Northwestern Europe . The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to the south, and the North Sea to
10873-647: The elections. Following a usage which can be traced back to the Burgundian and Habsburg courts, in the 19th century it was necessary to speak French to belong to the governing upper class, and those who could only speak Dutch were effectively second-class citizens. Late that century, and continuing into the 20th century, Flemish movements evolved to counter this situation. While the people in Southern Belgium spoke French or dialects of French, and most Brusselers adopted French as their first language,
11004-418: The end of nearly 300 million successful years for the trilobites would not have been unexpected at the time. Trilobites appear to have been primarily marine organisms, since the fossilized remains of trilobites are always found in rocks containing fossils of other salt-water animals such as brachiopods, crinoids, and corals. Some trackways suggest trilobites made at least temporary excursions onto land. Within
11135-610: The end of the Cambrian . Most scientists believe that order Redlichiida , more specifically its suborder Redlichiina , contains a common ancestor of all other orders, with the possible exception of the Agnostina. While many potential phylogenies are found in the literature, most have suborder Redlichiina giving rise to orders Corynexochida and Ptychopariida during the Lower Cambrian, and the Lichida descending from either
11266-467: The end of the Devonian period, what trilobite diversity remained was bottlenecked into the order Proetida. Decreasing diversity of genera limited to shallow-water shelf habitats coupled with a drastic lowering of sea level ( regression ) meant that the final decline of trilobites happened shortly before the end Permian mass extinction event . With so many marine species involved in the Permian extinction,
11397-681: The environment, town and country planning, nature conservation, credit and foreign trade. They supervise the provinces, municipalities and intercommunal utility companies. In several fields, the different levels each have their own say on specifics. With education, for instance, the autonomy of the Communities neither includes decisions about the compulsory aspect nor allows for setting minimum requirements for awarding qualifications, which remain federal matters. Each level of government can be involved in scientific research and international relations associated with its powers. The treaty-making power of
11528-477: The exoskeleton. Of the 20,000 known species only 38 have fossils with preserved appendages. Trilobites range in length from minute (less than 1 millimetre (0.039 in)) to very large (over 70 centimetres (28 in)), with an average size range of 3–10 cm (1.2–3.9 in). Supposedly the smallest species is Acanthopleurella stipulae with a maximum of 1.5 millimetres (0.059 in). The world's largest-known trilobite specimen, assigned to Isotelus rex
11659-682: The feeding trace, are furrows through the sediment, which are believed to represent the movement of trilobites while deposit feeding. Many of the Diplichnites fossils are believed to be traces made by trilobites walking on the sediment surface. Care must be taken as similar trace fossils are recorded in freshwater and post-Paleozoic deposits, representing non-trilobite origins. Trilobite fossils are found worldwide, with thousands of known species. Because they appeared quickly in geological time, and moulted like other arthropods, trilobites serve as excellent index fossils , enabling geologists to date
11790-795: The four components are subordinate to the Staff Department for Operations and Training of the Ministry of Defense , which is headed by the Assistant Chief of Staff Operations and Training , and to the Chief of Defense . The Belgian military consists of volunteers (conscription was abolished in 1995), and citizens of other EU states, Iceland, Norway, Switzerland, or Lichtenstein are also able to join. Belgium has troops deployed in several African countries as part of UN or EU missions, in Iraq for
11921-404: The glabella (impendent). Many variations in shape and placement of the hypostome have been described. The size of the glabella and the lateral fringe of the cephalon, together with hypostome variation, have been linked to different lifestyles, diets and specific ecological niches . The anterior and lateral fringe of the cephalon is greatly enlarged in the Harpetida , in other species a bulge in
12052-548: The government, and on 26 April 2010 King Albert officially accepted the resignation. The Parliamentary elections in Belgium on 13 June 2010 saw the Flemish nationalist N-VA become the largest party in Flanders, and the Socialist Party PS the largest party in Wallonia. Until December 2011, Belgium was governed by Leterme's caretaker government awaiting the end of the deadlocked negotiations for formation of
12183-402: The headshield (or cephalon ) is moderately to strongly inflated near to its front, more or less flattened on the top, falling vertically to or slightly overhanging the border furrow. Up to three lateral furrows may be discernable on the glabella behind the utterly dominating frontal lobe. From the back there is a very distinct occipital ring, and in front of that a distinct preoccipital ring which
12314-568: The incumbent coalition (composed of Flemish and French-speaking Social Democrats, Liberals, and Christian Democrats) maintains a solid majority in Parliament and in all electoral constituencies. On 22 July 2014, King Philippe nominated Charles Michel (MR) and Kris Peeters (CD&V) to lead the formation of a new federal cabinet composed of the Flemish parties N-VA, CD&V, Open Vld and the French-speaking MR, which resulted in
12445-586: The inhabitants of the northernmost part of Gaul . They lived in a region stretching from Paris to the Rhine , which is much bigger than modern Belgium. However, he also specifically used the Latin word " Belgium " to refer to a politically dominant part of that region, which is now in northernmost France. In contrast, modern Belgium, together with neighbouring parts of the Netherlands and Germany, corresponds to
12576-463: The installation of Leopold I as king on 21 July 1831, now celebrated as Belgium's National Day , Belgium has been a constitutional monarchy and parliamentary democracy , with a laicist constitution based on the Napoleonic code . Although the franchise was initially restricted, universal suffrage for men was introduced after the general strike of 1893 (with plural voting until 1919) and for women in 1949. The main political parties of
12707-471: The invasion of Iraq . It is the only country that does not have age restrictions on euthanasia. Verhofstadt's coalition fared badly in the June 2007 elections . For more than a year, the country experienced a political crisis . This crisis was such that many observers speculated on a possible partition of Belgium . From 21 December 2007 until 20 March 2008 the temporary Verhofstadt III Government
12838-546: The land from development. In 1994, the quarry became Penn Dixie Fossil Park & Nature Reserve when they received 501(c)3 status and was opened for visitation and collection of trilobite samples. The two most common found samples are Eldredgeops rana and Greenops . A famous location for trilobite fossils in the United Kingdom is Wren's Nest , Dudley , in the West Midlands , where Calymene blumenbachii
12969-620: The lands of the most northerly Belgae – the Morini , Menapii , Nervii , Germani Cisrhenani , and Aduatuci . Caesar found these peoples particularly warlike and economically undeveloped, and described them as kinsmen of the Germanic tribes east of the Rhine. The area around Arlon in southern Belgium was a part of the country of the powerful Treveri , to whom some of them paid tribute. After Caesar's conquests, Gallia Belgica first came to be
13100-400: The last few survivors at the end of the Permian period. Principal evolutionary trends from primitive morphologies, such as exemplified by Eoredlichia , include the origin of new types of eyes, improvement of enrollment and articulation mechanisms, increased size of pygidium (micropygy to isopygy), and development of extreme spinosity in certain groups. Changes also included narrowing of
13231-439: The marine paleoenvironment, trilobites were found in a broad range from extremely shallow water to very deep water. Trilobites, like brachiopods, crinoids, and corals, are found on all modern continents, and occupied every ancient ocean from which Paleozoic fossils have been collected. The remnants of trilobites can range from the preserved body to pieces of the exoskeleton, which it shed in the process known as ecdysis. In addition,
13362-426: The moult stage and the hardening of the new exoskeleton. A trilobite's cephalon, or head section, is highly variable with a lot of morphological complexity. The glabella forms a dome underneath which sat the "crop" or "stomach". Generally, the exoskeleton has few distinguishing ventral features, but the cephalon often preserves muscle attachment scars and occasionally the hypostome , a small rigid plate comparable to
13493-739: The north, the Walloon Region (Wallonia) in the south, and the Brussels-Capital Region in the middle. Brussels is the smallest and most densely populated region, as well as the richest region in terms of GDP per capita . Belgium is also home to two main linguistic communities: the Flemish Community ( Dutch -speaking), which constitutes about 60 percent of the population, and the French Community ( French -speaking), which constitutes about 40 percent of
13624-657: The other Frankish kingdoms. During the 8th century, the empire of the Franks came to be ruled by the Carolingian dynasty , whose centre of power included the area which is now eastern Belgium. Over the centuries, it was divided up in many ways, but the Treaty of Verdun in 843 divided the Carolingian Empire into three kingdoms whose borders had a lasting impact on medieval political boundaries. Most of modern Belgium
13755-852: The parliament accepted a Dutch version of the Constitution . The Berlin Conference of 1885 ceded control of the Congo Free State to King Leopold II as his private possession. From around 1900 there was growing international concern for the extreme and savage treatment of the Congolese population under Leopold II , for whom the Congo was primarily a source of revenue from ivory and rubber production. Many Congolese were killed by Leopold's agents for failing to meet production quotas for ivory and rubber. In 1908, this outcry led
13886-413: The pleural areas to the sides have 5 to 8 pairs of recognizable ribs. Furrows between the ribs are deep, those that divide each rib in frontal and rear bands are very shallow, and the frontal bands are widest. The surface of the exoskeleton is covered in tubercles . There are specimens known of Phacops rana with many irregular black spots. Because similar spots in a specimen of Greenops boothi from
14017-504: The political and linguistic interests of these communities. The major parties in each community, though close to the political center, belong to three main groups: Christian Democrats , Liberals , and Social Democrats . Further notable parties came into being well after the middle of last century, mainly to represent linguistic , nationalist, or environmental interests, and recently smaller ones of some specific liberal nature. A string of Christian Democrat coalition governments from 1958
14148-551: The population. A small German-speaking Community , making up around one percent of the population, exists in the East Cantons . The Brussels-Capital Region is officially bilingual in French and Dutch, although French is the majority language and lingua franca . Belgium's linguistic diversity and related political conflicts are reflected in its complex system of governance, made up of six different governments . In antiquity
14279-404: The possible exception of parasitism (where scientific debate continues). Some trilobites (particularly the family Olenidae ) are even thought to have evolved a symbiotic relationship with sulfur-eating bacteria from which they derived food. The largest trilobites were more than 70 centimetres (28 in) long and may have weighed as much as 4.5 kilograms (9.9 lb). Trilobites belong to
14410-417: The pre-glabellar area is preserved that suggests a brood pouch. Highly complex compound eyes are another obvious feature of the cephalon. Facial or cephalic sutures are the natural fracture lines in the cephalon of trilobites. Their function was to assist the trilobite in shedding its old exoskeleton during ecdysis (or molting). All species assigned to the suborder Olenellina , that became extinct at
14541-405: The rate of speciation during the period known as the Cambrian explosion because they are the most diverse group of metazoans known from the fossil record of the early Cambrian. Trilobites are excellent stratigraphic markers of the Cambrian period: researchers who find trilobites with alimentary prosopon, and a micropygium, have found Early Cambrian strata. Most of the Cambrian stratigraphy
14672-776: The region around Belgium was inhabited by the Belgae , and became the Roman province of Gallia Belgica in 22 BC. In the Middle Ages , the region was part of the Carolingian Empire , and much of it was later part of the Holy Roman Empire , and subsequently the Burgundian Netherlands . Belgium's central location has meant that the area has been relatively prosperous, connected commercially and politically to its bigger neighbours. The country as it exists today
14803-529: The same site are arranged in rows, it may be assumed that they are original and not caused by the fossilisation process. The spots are irregular and have spurs branching outwardly, similar to the melanophores in many extant animals. In one specimen, the black spots are much larger than in another one. It is quite conceivable that changing the size of the melanophores enabled Phacops rana to camouflage itself in different environments. The concept of many fossil taxa has been tightened over time, including Phacops . As
14934-533: The southeast to the Hercynian orogenic belt . The Paris Basin reaches a small fourth area at Belgium's southernmost tip, Belgian Lorraine . The coastal plain consists mainly of sand dunes and polders . Further inland lies a smooth, slowly rising landscape irrigated by numerous waterways, with fertile valleys and the northeastern sandy plain of the Campine ( Kempen ). The thickly forested hills and plateaus of
15065-594: The thoracic furrows, is also a common evolutionary trend. Notable examples of this were the orders Agnostida and Asaphida , and the suborder Illaenina of the Corynexochida . Effacement is believed to be an indication of either a burrowing lifestyle or a pelagic one. Effacement poses a problem for taxonomists since the loss of details (particularly of the glabella ) can make the determination of phylogenetic relationships difficult. Although it has historically been suggested that trilobites originated during
15196-406: The thorax and increasing or decreasing numbers of thoracic segments. Specific changes to the cephalon are also noted; variable glabella size and shape, position of eyes and facial sutures, and hypostome specialization. Several morphologies appeared independently within different major taxa (e.g. eye reduction or miniaturization). Effacement, the loss of surface detail in the cephalon, pygidium, or
15327-505: The tracks left behind by trilobites living on the sea floor are often preserved as trace fossils . There are three main forms of trace fossils associated with trilobites: Rusophycus , Cruziana and Diplichnites —such trace fossils represent the preserved life activity of trilobites active upon the sea floor. Rusophycus , the resting trace, are trilobite excavations involving little or no forward movement and ethological interpretations suggest resting, protection and hunting. Cruziana ,
15458-448: The transition from a unitary to a federal arrangement between 1970 and 1993. Despite the reforms, tensions have persisted: there is particularly significant separatist sentiment among the Flemish ; language laws such as the municipalities with language facilities have been the source of much controversy; and the government formation period following the 2010 federal election set a world record at 589 days. Unemployment in Wallonia
15589-596: The trilobites unscathed; some distinctive and previously successful forms such as the Telephinidae and Agnostida became extinct. The Ordovician marks the last great diversification period amongst the trilobites: very few entirely new patterns of organisation arose post-Ordovician. Later evolution in trilobites was largely a matter of variations upon the Ordovician themes. By the Ordovician mass extinction , vigorous trilobite radiation has stopped, and gradual decline
15720-448: The ventral plate in other arthropods. A toothless mouth and stomach sat upon the hypostome with the mouth facing backward at the rear edge of the hypostome. Hypostome morphology is highly variable; sometimes supported by an un-mineralised membrane (natant), sometimes fused onto the anterior doublure with an outline very similar to the glabella above (conterminant) or fused to the anterior doublure with an outline significantly different from
15851-554: The very end of the Early Cambrian (like Fallotaspis , Nevadia , Judomia , and Olenellus ) lacked facial sutures. They are believed to have never developed facial sutures, having pre-dated their evolution. Because of this (along with other primitive characteristics), they are thought to be the earliest ancestors of later trilobites. Some other later trilobites also lost facial sutures secondarily. The type of sutures found in different species are used extensively in
15982-728: The war were known as the Rape of Belgium due to German excesses. Belgium assumed control of the German colonies of Ruanda-Urundi (modern-day Rwanda and Burundi ) during the war, and in 1924 the League of Nations mandated them to Belgium. In the aftermath of the First World War, Belgium annexed the Prussian districts of Eupen and Malmedy in 1925, thereby causing the presence of a German-speaking minority. German forces again invaded
16113-702: The war. The Belgian Congo gained independence in 1960 during the Congo Crisis ; Ruanda-Urundi followed with its independence two years later. Belgium joined NATO as a founding member and formed the Benelux group of nations with the Netherlands and Luxembourg. Belgium became one of the six founding members of the European Coal and Steel Community in 1951 and of the European Atomic Energy Community and European Economic Community , established in 1957. The latter has now become
16244-629: The west. It covers an area of 30,689 km (11,849 sq mi) and has a population of more than 11.7 million, making it the 22nd most densely populated country in the world and the 6th most densely populated country in Europe, with a density of 383/km (990/sq mi). Belgium is part of an area known as the Low Countries , historically a somewhat larger region than the Benelux group of states, as it also included parts of northern France. The capital and largest metropolitan region
16375-568: Was broken in 1999 after the first dioxin crisis , a major food contamination scandal. A "rainbow coalition" emerged from six parties: the Flemish and the French-speaking Liberals, Social Democrats and Greens. Later, a " purple coalition " of Liberals and Social Democrats formed after the Greens lost most of their seats in the 2003 election . The government led by Prime Minister Guy Verhofstadt from 1999 to 2007 achieved
16506-547: Was designated the first permanent President of the European Council on 19 November 2009, he offered the resignation of his government to King Albert II on 25 November 2009. A few hours later, the new government under Prime Minister Yves Leterme was sworn in. On 22 April 2010, Leterme again offered the resignation of his cabinet to the king after one of the coalition partners, the OpenVLD , withdrew from
16637-526: Was established following the 1830 Belgian Revolution , when it seceded from the United Kingdom of the Netherlands , which had incorporated the Southern Netherlands (which comprised most of modern-day Belgium) after the Congress of Vienna in 1815. Belgium has been called "the Battlefield of Europe", a reputation reinforced in the 20th century by both world wars . Belgium was an early participant in
16768-583: Was fundamental in formulating and testing punctuated equilibrium as a mechanism of evolution. Identification of the 'Atlantic' and 'Pacific' trilobite faunas in North America and Europe implied the closure of the Iapetus Ocean (producing the Iapetus suture), thus providing important supporting evidence for the theory of continental drift . Trilobites have been important in estimating
16899-541: Was in office. This was a coalition of the Flemish and Francophone Christian Democrats , the Flemish and Francophone Liberals together with the Francophone Social Democrats . On that day a new government , led by Flemish Christian Democrat Yves Leterme , the actual winner of the federal elections of June 2007 , was sworn in by the king. On 15 July 2008 Leterme offered the resignation of
17030-543: Was in the Middle Kingdom , later known as Lotharingia , but the coastal county of Flanders , west of the Scheldt , became the northernmost part of West Francia , the predecessor of France . In 870 in the Treaty of Meerssen , modern Belgium lands all became part of the western kingdom for a period, but in 880 in the Treaty of Ribemont , Lotharingia came under the lasting control of the eastern kingdom, which became
17161-469: Was proposed to be elevated out of the asaphid superfamily Trinucleioidea . Sometimes the Nektaspida are considered trilobites, but these lack a calcified exoskeleton and eyes. Some scholars have proposed that the order Agnostida is polyphyletic, with the suborder Agnostina representing non-trilobite arthropods unrelated to the suborder Eodiscina . Under this hypothesis, Eodiscina would be elevated to
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