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Theridiidae

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Order ( Latin : ordo ) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy . It is classified between family and class . In biological classification , the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes . An immediately higher rank, superorder , is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families.

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22-420: Theridiidae , also known as the tangle-web spiders , cobweb spiders and comb-footed spiders , is a large family of araneomorph spiders first described by Carl Jakob Sundevall in 1833. This diverse, globally distributed family includes over 3,000 species in 124 genera , and is the most common arthropod found in human dwellings throughout the world. Theridiid spiders are both entelegyne , meaning that

44-550: Is Theridion with over 600 species, but it is not monophyletic . Parasteatoda , previously Achaearanea , is another large genus that includes the North American common house spider . As of April 2019, the World Spider Catalog accepts the following genera: About 35 extinct genera have also been placed in the family. The oldest known stem-group member of the family is Cretotheridion from

66-459: Is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. For some groups of organisms, their orders may follow consistent naming schemes . Orders of plants , fungi , and algae use

88-659: The Prodromus Systematis Naturalis Regni Vegetabilis of Augustin Pyramus de Candolle and the Genera Plantarum of Bentham & Hooker, it indicated taxa that are now given the rank of family (see ordo naturalis , ' natural order '). In French botanical publications, from Michel Adanson 's Familles naturelles des plantes (1763) and until the end of the 19th century, the word famille (plural: familles )

110-597: The Cenomanian aged Burmese amber of Myanmar. Araneomorphae The Araneomorphae (also called the Labidognatha or " true spiders " ) are an infraorder of spiders . They are distinguishable by chelicerae (fangs) that point diagonally forward and cross in a pinching action, in contrast to the Mygalomorphae (tarantulas and their close kin), where they point straight down. Araneomorphs comprise

132-586: The Hypochilae (containing only the family Hypochilidae ), and the Neocribellatae . The Neocribellatae were in turn divided into the Austrochiloidea, and the two series Haplogynae and Entelogynae, each containing several superfamilies . Molecular phylogenetic studies have shown that the haplogynes in particular are not a monophyletic group. A 2020 study suggested the relationships among

154-496: The contrary, include the weavers of spiral webs ; the cobweb spiders that live in the corners of rooms, and between windows and screens; the crab spiders that lurk on the surfaces of flowers in gardens; the jumping spiders that are visible hunting on surfaces; the wolf spiders that carpet hunting sites in sunny spots; and the large huntsman spiders . In older schemes, the Araneomorphae were divided into two lineages,

176-472: The ending -anae that was initiated by Armen Takhtajan 's publications from 1966 onwards. The order as a distinct rank of biological classification having its own distinctive name (and not just called a higher genus ( genus summum )) was first introduced by the German botanist Augustus Quirinus Rivinus in his classification of plants that appeared in a series of treatises in the 1690s. Carl Linnaeus

198-564: The females have a genital plate, and ecribellate , meaning that they spin sticky capture silk instead of woolly silk. They have a comb of serrated bristles ( setae ) on the tarsus of the fourth leg. The family includes some model organisms for research, including the medically important widow spiders . They are important to studies characterizing their venom and its clinical manifestation, but widow spiders are also used in research on spider silk and sexual biology, including sexual cannibalism . Anelosimus are also model organisms, used for

220-910: The field of zoology , the Linnaean orders were used more consistently. That is, the orders in the zoology part of the Systema Naturae refer to natural groups. Some of his ordinal names are still in use, e.g. Lepidoptera (moths and butterflies) and Diptera (flies, mosquitoes, midges, and gnats). In virology , the International Committee on Taxonomy of Viruses 's virus classification includes fifteen taxomomic ranks to be applied for viruses , viroids and satellite nucleic acids : realm , subrealm , kingdom , subkingdom, phylum , subphylum , class, subclass, order, suborder, family, subfamily , genus, subgenus , and species. There are currently fourteen viral orders, each ending in

242-445: The genetic basis of color polymorphism within species. T. grallator is known as the "happyface" spider, as certain morphs have a pattern resembling a smiley face or a grinning clown face on their yellow body. They often build tangle space webs , hence the common name, but Theridiidae has a large diversity of spider web forms. Many trap ants and other ground dwelling insects using elastic, sticky silk trap lines leading to

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264-673: The host web, and sometimes attack and eat the host itself. Theridiid gumfoot-webs consist of frame lines that anchor them to surroundings and of support threads, which possess viscid silk. These can either have a central retreat ( Achaearanea -type) or a peripheral retreat ( Latrodectus -type). Building gum-foot lines is a unique, stereotyped behaviour, and is likely homologous for Theridiidae and its sister family Nesticidae . Among webs without gumfooted lines, some contain viscid silk ( Theridion -type) and some that are sheet-like, which do not contain viscid silk ( Coleosoma -type). However, there are many undescribed web forms. The largest genus

286-472: The long axis of the spider's body, thus they have only one orientation they can employ during prey capture. They have two pairs of book lungs, and the females often live many years. Almost all of the familiar spiders are included in the Araneomorphae group, one major exception being the tarantulas . There are a few other Mygalomorphae species that live around homes or gardens, but they typically are relatively small and not easily noticed. The Araneomorphae, to

308-446: The major groups were as shown in the following cladogram. Hypochilidae Filistatidae Synspermiata Leptonetidae Austrochiloidea Palpimanoidea Entelegynae The blue bar to the right shows the former Haplogynae in the sense of Coddington (2005). Infraorder What does and does not belong to each order is determined by a taxonomist , as is whether a particular order should be recognized at all. Often there

330-420: The same position. Michael Benton (2005) inserted them between superorder and magnorder instead. This position was adopted by Systema Naturae 2000 and others. In botany , the ranks of subclass and suborder are secondary ranks pre-defined as respectively above and below the rank of order. Any number of further ranks can be used as long as they are clearly defined. The superorder rank is commonly used, with

352-410: The soil surface. Webs remain in place for extended periods and are expanded and repaired, but no regular pattern of web replacement has been observed. The well studied kleptoparasitic members of Argyrodinae ( Argyrodes , Faiditus , and Neospintharus ) live in the webs of larger spiders and pilfer small prey caught by their host's web. They eat prey killed by the host spider, consume silk from

374-423: The study of sociality, because it has evolved frequently within the genus, allowing comparative studies across species, and because it contains species varying from solitary to permanently social. These spiders are also a promising model for the study of inbreeding because all permanently social species are highly inbred. The Hawaiian Theridion grallator is used as a model to understand the selective forces and

396-777: The suffix -ales (e.g. Dictyotales ). Orders of birds and fishes use the Latin suffix -iformes meaning 'having the form of' (e.g. Passeriformes ), but orders of mammals and invertebrates are not so consistent (e.g. Artiodactyla , Actiniaria , Primates ). For some clades covered by the International Code of Zoological Nomenclature , several additional classifications are sometimes used, although not all of these are officially recognized. In their 1997 classification of mammals , McKenna and Bell used two extra levels between superorder and order: grandorder and mirorder . Michael Novacek (1986) inserted them at

418-406: The vast majority (about 93% ) of living spiders. Most spider species are Araneomorphae, which have fangs that face towards each other, increasing the orientations that they can employ during prey-capture. They have fewer book lungs (when present) - usually one pair - and the females typically live one year. The Mygalomorphae have fangs that face towards the ground, and which are parallel to

440-578: The word family ( familia ) was assigned to the rank indicated by the French famille , while order ( ordo ) was reserved for a higher rank, for what in the 19th century had often been named a cohors (plural cohortes ). Some of the plant families still retain the names of Linnaean "natural orders" or even the names of pre-Linnaean natural groups recognized by Linnaeus as orders in his natural classification (e.g. Palmae or Labiatae ). Such names are known as descriptive family names. In

462-551: Was the first to apply it consistently to the division of all three kingdoms of nature (then minerals , plants , and animals ) in his Systema Naturae (1735, 1st. Ed.). For plants, Linnaeus' orders in the Systema Naturae and the Species Plantarum were strictly artificial, introduced to subdivide the artificial classes into more comprehensible smaller groups. When the word ordo was first consistently used for natural units of plants, in 19th-century works such as

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484-561: Was used as a French equivalent for this Latin ordo . This equivalence was explicitly stated in the Alphonse Pyramus de Candolle 's Lois de la nomenclature botanique (1868), the precursor of the currently used International Code of Nomenclature for algae, fungi, and plants . In the first international Rules of botanical nomenclature from the International Botanical Congress of 1905,

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