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100-534: The physical appearance of this domestic cat breed barely changed when compared to photos dating back to the late 1800s. However, since some breeders in the United States and other parts of the world introduced the brachycephalic mutation into the breed, the short nose and clear break became shorter and higher. This resulted in the modification of the original Persian breed standard so that modern show quality Persians must have pushed-in faces . This resulted in

200-666: A family that had a common ancestor about 10 to 15 million years ago . The evolutionary radiation of the Felidae began in Asia during the Miocene around 8.38 to 14.45 million years ago . Analysis of mitochondrial DNA of all Felidae species indicates a radiation at 6.46 to 16.76 million years ago . The genus Felis genetically diverged from other Felidae around 6 to 7 million years ago . Results of phylogenetic research shows that

300-464: A nictitating membrane , allowing them to blink without hindering their vision. The domestic cat's hearing is most acute in the range of 500 Hz to 32 kHz. It can detect an extremely broad range of frequencies ranging from 55 Hz to 79 kHz, whereas humans can only detect frequencies between 20 Hz and 20 kHz. It can hear a range of 10.5 octaves , while humans and dogs can hear ranges of about 9 octaves. Its hearing sensitivity

400-420: A "broad" definition: "a coevolutionary process that arises from a mutualism, in which one species (the domesticator) constructs an environment where it actively manages both the survival and reproduction of another species (the domesticate) in order to provide the former with resources and/or services." He comments that this adds niche construction to the activities of the domesticator. Domestication syndrome

500-520: A "diagonal" gait: The diagonally opposite hind and fore legs move simultaneously. Cats are generally fond of sitting in high places or perching . A higher place may serve as a concealed site from which to hunt; domestic cats strike prey by pouncing from a perch such as a tree branch. Another possible explanation is that height gives the cat a better observation point, allowing it to survey its territory. A cat falling from heights of up to 3 m (9.8 ft) can right itself and land on its paws. During

600-507: A cat's pupils expand to cover most of the exposed surface of its eyes. The domestic cat has rather poor color vision and only two types of cone cells , optimized for sensitivity to blue and yellowish green; its ability to distinguish between red and green is limited. A response to middle wavelengths from a system other than the rod cells might be due to a third type of cone. This appears to be an adaptation to low light levels rather than representing true trichromatic vision. Cats also have

700-592: A cosegregation between the ASIP allele and coat black coloration. Cats have excellent night vision and can see at one sixth the light level required for human vision. This is partly the result of cat eyes having a tapetum lucidum , which reflects any light that passes through the retina back into the eye, thereby increasing the eye's sensitivity to dim light. Large pupils are an adaptation to dim light. The domestic cat has slit pupils , which allow it to focus bright light without chromatic aberration . At low light,

800-430: A fall from a high place, a cat reflexively twists its body and rights itself to land on its feet using its acute sense of balance and flexibility. This reflex is known as the cat righting reflex . A cat always rights itself in the same way during a fall, if it has enough time to do so, which is the case in falls of 90 cm (3.0 ft) or more. How cats are able to right themselves when falling has been investigated as

900-860: A few have been domesticated, including squid , cuttlefish and octopus , all used in research on behaviour and neurology . Terrestrial snails in the genera Helix are raised for food. Several parasitic or parasitoidal insects, including the fly Eucelatoria , the beetle Chrysolina , and the wasp Aphytis are raised for biological control. Conscious or unconscious artificial selection has many effects on species under domestication; variability can readily be lost by inbreeding, selection against undesired traits, or genetic drift, while in Drosophila , variability in eclosion time (when adults emerge) has increased. Humans foraged for wild cereals, seeds, and nuts thousands of years before they were domesticated; wild wheat and barley, for example, were gathered in

1000-495: A fresh kill; some cats reject cold food (which would signal to the cat that the "prey" item is long dead and therefore possibly toxic or decomposing). To aid with navigation and sensation, cats have dozens of movable whiskers (vibrissae) over their body, especially their faces. These provide information on the width of gaps and on the location of objects in the dark, both by touching objects directly and by sensing air currents; they also trigger protective blink reflexes to protect

1100-710: A large surface of olfactory mucosa , about 5.8 cm (0.90 in ) in area, which is about twice that of humans. Cats and many other animals have a Jacobson's organ in their mouths that is used in the behavioral process of flehmening . It allows them to sense certain aromas in a way that humans cannot. Cats are sensitive to pheromones such as 3-mercapto-3-methylbutan-1-ol , which they use to communicate through urine spraying and marking with scent glands . Many cats also respond strongly to plants that contain nepetalactone , especially catnip , as they can detect that substance at less than one part per billion. About 70–80% of cats are affected by nepetalactone. This response

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1200-484: A means of communication between far-flung places through the exploitation of the pigeon's homing instinct; research suggests it was domesticated as early as 10,000 years ago. Chicken fossils in China have been dated to 7,400 years ago. The chicken's wild ancestor is Gallus gallus , the red junglefowl of Southeast Asia. The species appears to have been kept initially for cockfighting rather than for food. Two insects ,

1300-468: A population bottleneck where genetic drift is enhanced by the small population size. Mutations can also be fixed in a population by a selective sweep . Mutational load can be increased by reduced selective pressure against moderately harmful traits when reproductive fitness is controlled by human management. However, there is evidence against a bottleneck in crops, such as barley, maize, and sorghum, where genetic diversity slowly declined rather than showing

1400-779: A predictable supply of a resource, resulting in mutual benefits . She noted further that it is not synonymous with agriculture since agriculture depends on domesticated organisms but does not automatically result from domestication. Michael D. Purugganan notes that domestication has been hard to define, despite the "instinctual consensus" that it means "the plants and animals found under the care of humans that provide us with benefits and which have evolved under our control." He comments that insects such as termites , ambrosia beetles , and leafcutter ants have domesticated some species of fungi , and notes further that other groups such as weeds and commensals have wrongly been called domesticated. Starting from Zeder's definition, Purugganan proposes

1500-537: A project to separate the breed from the modern Persian standard. This standard was only to be applicable to color-bred silvers, commonly known as "Chinchillas" in South Africa. The breed was accepted under the name "Chinchilla Longhair" with breed code "CHL" in the SA Cat Council (SACC) in 1996. A global standard for the "Traditional Longhair silver and golden, shaded and tipped varieties" was accepted by

1600-479: A project was started to obtain breed recognition of the Original Longhair in all WCC members. Domestic cat The cat ( Felis catus ), also referred to as domestic cat or house cat , is a small domesticated carnivorous mammal. It is the only domesticated species of the family Felidae . Advances in archaeology and genetics have shown that the domestication of the cat occurred in

1700-425: A protracted coevolutionary process with multiple stages along different pathways. There are three proposed major pathways that most mammal domesticates followed into domestication: Humans did not intend to domesticate mammals from either the commensal or prey pathways, or at least they did not envision a domesticated animal would result from it. In both of those cases, humans became entangled with these species as

1800-575: A rapid initial fall at the point of domestication. Further, the genetic diversity of these crops was regularly replenished from the natural population. Similar evidence exists for horses, pigs, cows, and goats. At least three groups of insects, namely ambrosia beetles, leafcutter ants, and fungus-growing termites, have domesticated species of fungi . Ambrosia beetles in the weevil subfamilies Scolytinae and Platypodinae excavate tunnels in dead or stressed trees into which they introduce fungal gardens, their sole source of nutrition. After landing on

1900-457: A steady supply of resources, such as meat, milk, or labor. The process is gradual and geographically diffuse, based on trial and error. Domestication affected genes for behavior in animals, making them less aggressive. In plants, domestication affected genes for morphology, such as increasing seed size and stopping the shattering of cereal seedheads. Such changes both make domesticated organisms easier to handle and reduce their ability to survive in

2000-402: A suitable tree, an ambrosia beetle excavates a tunnel in which it releases its fungal symbiont . The fungus penetrates the plant's xylem tissue, extracts nutrients from it, and concentrates the nutrients on and near the surface of the beetle gallery. Ambrosia fungi are typically poor wood degraders and instead utilize less demanding nutrients. Symbiotic fungi produce and detoxify ethanol, which

2100-437: A thicker protective layer of enamel , a less damaging saliva, less retention of food particles between teeth, and a diet mostly devoid of sugar. Nonetheless, they are subject to occasional tooth loss and infection. Cats have protractible and retractable claws. In their normal, relaxed position, the claws are sheathed with the skin and fur around the paw's toe pads. This keeps the claws sharp by preventing wear from contact with

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2200-514: A variable number of caudal vertebrae in the tail (humans have only three to five vestigial caudal vertebrae, fused into an internal coccyx ). The extra lumbar and thoracic vertebrae account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis . Unlike human arms, cat forelimbs are attached to the shoulder by free-floating clavicle bones which allow them to pass their body through any space into which they can fit their head. The cat skull

2300-474: A working animal. Among invertebrates , the silkworm and the western honey bee were domesticated over 5,000 years ago for silk and honey , respectively. The domestication of plants began around 13,000–11,000 years ago with cereals such as wheat and barley in the Middle East , alongside crops such as lentil , pea , chickpea , and flax . Beginning around 10,000 years ago, Indigenous peoples in

2400-511: Is digitigrade . It walks on the toes, with the bones of the feet making up the lower part of the visible leg. Unlike most mammals, it uses a "pacing" gait and moves both legs on one side of the body before the legs on the other side. It registers directly by placing each hind paw close to the track of the corresponding fore paw, minimizing noise and visible tracks. This also provides sure footing for hind paws when navigating rough terrain. As it speeds up from walking to trotting, its gait changes to

2500-496: Is proximal to the other claws. More proximally is a protrusion which appears to be a sixth "finger". This special feature of the front paws on the inside of the wrists has no function in normal walking but is thought to be an antiskidding device used while jumping. Some cat breeds are prone to having extra digits (" polydactyly "). Polydactylous cats occur along North America's northeast coast and in Great Britain. The cat

2600-489: Is a social species , but a solitary hunter and a crepuscular predator . Cat communication includes vocalizations—including meowing , purring , trilling, hissing , growling , and grunting —as well as body language . It can hear sounds too faint or too high in frequency for human ears , such as those made by small mammals . It secretes and perceives pheromones . Female domestic cats can have kittens from spring to late autumn in temperate zones and throughout

2700-571: Is also possible, producing hybrids such as the Kellas cat in Scotland . Development of cat breeds started in the mid 19th century. An analysis of the domestic cat genome revealed that the ancestral wildcat genome was significantly altered in the process of domestication, as specific mutations were selected to develop cat breeds. Most breeds are founded on random-bred domestic cats. Genetic diversity of these breeds varies between regions, and

2800-669: Is also produced by other plants, such as silver vine ( Actinidia polygama ) and the herb valerian ; it may be caused by the smell of these plants mimicking a pheromone and stimulating cats' social or sexual behaviors. Cats have relatively few taste buds compared to humans (470 or so, compared to more than 9,000 on the human tongue). Domestic and wild cats share a taste receptor gene mutation that keeps their sweet taste buds from binding to sugary molecules, leaving them with no ability to taste sweetness . They, however, possess taste bud receptors specialized for acids , amino acids like protein, and bitter tastes. Their taste buds possess

2900-405: Is an attractant for ambrosia beetles and likely prevents the growth of antagonistic pathogens and selects for other beneficial symbionts. Ambrosia beetles mainly colonise wood of recently dead trees. The leafcutter ants are any of some 47 species of leaf-chewing ants in the genera Acromyrmex and Atta . The ants carry the discs of leaves that they have cut back to their nest, where they feed

3000-459: Is attested from the 16th century and may have been introduced from Dutch poes or from Low German puuskatte , related to Swedish kattepus , or Norwegian pus , pusekatt . Similar forms exist in Lithuanian puižė and Irish puisín or puiscín . The etymology of this word is unknown, but it may have arisen from a sound used to attract a cat. A male cat

3100-400: Is called a tom or tomcat (or a gib , if neutered ). A female is called a queen (or sometimes a molly , if spayed ). A juvenile cat is referred to as a kitten . In Early Modern English , the word kitten was interchangeable with the now-obsolete word catling . A group of cats can be referred to as a clowder , a glaring , or a colony . The scientific name Felis catus

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3200-432: Is domesticated wheat, which relies on farmers for its reproduction and dissemination. Domesticated plants differ from their wild relatives in many ways, including Plant defenses against herbivory , such as thorns, spines, and prickles , poison, protective coverings, and sturdiness may have been reduced in domesticated plants. This would make them more likely to be eaten by herbivores unless protected by humans, but there

3300-463: Is enhanced by its large movable outer ears, the pinnae , which amplify sounds and help detect the location of a noise. It can detect ultrasound , which enables it to detect ultrasonic calls made by rodent prey. Recent research has shown that cats have socio-spatial cognitive abilities to create mental maps of owners' locations based on hearing owners' voices. Cats have an acute sense of smell, due in part to their well-developed olfactory bulb and

3400-719: Is lowest in purebred populations, which show more than 20 deleterious genetic disorders . The domestic cat has a smaller skull and shorter bones than the European wildcat . It averages about 46 cm (18 in) in head-to-body length and 23–25 cm (9.1–9.8 in) in height, with about 30 cm (12 in) long tails. Males are larger than females. Adult domestic cats typically weigh 4–5 kg (8.8–11.0 lb). Cats have seven cervical vertebrae (as do most mammals ); 13 thoracic vertebrae (humans have 12); seven lumbar vertebrae (humans have five); three sacral vertebrae (as do most mammals, but humans have five); and

3500-669: Is only weak support for most of this. Farmers did select for reduced bitterness and lower toxicity and for food quality, which likely increased crop palatability to herbivores as to humans. However, a survey of 29 plant domestications found that crops were as well-defended against two major insect pests ( beet armyworm and green peach aphid ) both chemically (e.g. with bitter substances) and morphologically (e.g. with toughness) as their wild ancestors. During domestication, crop species undergo intense artificial selection that alters their genomes, establishing core traits that define them as domesticated, such as increased grain size. Comparison of

3600-544: Is quite flexible and varied but being low-light predators, they are generally crepuscular , which means they tend to be more active near dawn and dusk. However, house cats' behavior is also influenced by human activity and they may adapt to their owners' sleeping patterns to some extent. Domestication Domestication is a multi-generational mutualistic relationship in which an animal species, such as humans or leafcutter ants , takes over control and care of another species, such as sheep or fungi, to obtain from them

3700-668: Is the suite of phenotypic traits that arose during the initial domestication process and which distinguish crops from their wild ancestors . It can also mean a set of differences now observed in domesticated mammals, not necessarily reflecting the initial domestication process. The changes include increased docility and tameness, coat coloration, reductions in tooth size, craniofacial morphology, ear and tail form (e.g., floppy ears), estrus cycles, levels of adrenocorticotropic hormone and neurotransmitters, prolongations in juvenile behavior, and reductions in brain size and of particular brain regions. The domestication of animals and plants

3800-725: Is today Baluchistan in Pakistan. In East Asia 8,000 years ago, pigs were domesticated from wild boar genetically different from those found in the Fertile Crescent. The cat was domesticated in the Fertile Crescent, perhaps 10,000 years ago, from European wildcats , possibly to control rodents that were damaging stored food. The domestication of vertebrate animals is the relationship between non-human vertebrates and humans who have an influence on their care and reproduction. In his 1868 book The Variation of Animals and Plants Under Domestication , Charles Darwin recognized

3900-482: Is unusual among mammals in having very large eye sockets and a powerful specialized jaw. Within the jaw, cats have teeth adapted for killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal neck bite with its two long canine teeth , inserting them between two of the prey's vertebrae and severing its spinal cord , causing irreversible paralysis and death. Compared to other felines, domestic cats have narrowly spaced canine teeth relative to

4000-636: Is widely grown for food. The yeast Saccharomyces cerevisiae have been used for thousands of years to ferment beer and wine , and to leaven bread . Mould fungi including Penicillium are used to mature cheeses and other dairy products, as well as to make drugs such as antibiotics . Selection of animals for visible traits may have undesired consequences for the genetics of domestic animals. A side effect of domestication has been zoonotic diseases. For example, cattle have given humanity various viral poxes , measles , and tuberculosis ; pigs and ducks have contributed influenza ; and horses have brought

4100-718: The Indian Ocean and those in the Pacific Ocean . The coconut experienced a founder effect , where a small number of individuals with low diversity founded the modern population, permanently losing much of the genetic variation of the wild population. Population bottlenecks which reduced variation throughout the genome at some later date after domestication are evident in crops such as pearl millet , cotton , common bean and lima bean . In wheat, domestication involved repeated hybridization and polyploidy . These steps are large and essentially instantaneous changes to

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4200-634: The Levant at least 23,000 years ago. Neolithic societies in West Asia first began to cultivate and then domesticate some of these plants around 13,000 to 11,000 years ago. The founder crops of the West Asian Neolithic included cereals ( emmer , einkorn wheat , barley ), pulses ( lentil , pea , chickpea , bitter vetch ), and flax . Other plants were independently domesticated in 13 centers of origin (subdivided into 24 areas) of

4300-493: The Near East around 7500 BC . It is commonly kept as a pet and farm cat , but also ranges freely as a feral cat avoiding human contact. Valued by humans for companionship and its ability to kill vermin , the cat's retractable claws are adapted to killing small prey like mice and rats . It has a strong, flexible body, quick reflexes , and sharp teeth, and its night vision and sense of smell are well developed. It

4400-619: The coding DNA of chromosome 8 in rice between fragrant and non-fragrant varieties showed that aromatic and fragrant rice, including basmati and jasmine , is derived from an ancestral rice domesticate that suffered a deletion in exon 7 which altered the coding for betaine aldehyde dehydrogenase (BADH2). Comparison of the potato genome with that of other plants located genes for resistance to potato blight caused by Phytophthora infestans . In coconut , genomic analysis of 10 microsatellite loci (of noncoding DNA ) found two episodes of domestication based on differences between individuals in

4500-470: The domestication of wheat . Wild wheat shatters and falls to the ground to reseed itself when ripe, but domesticated wheat stays on the stem for easier harvesting. This change was possible because of a random mutation in the wild populations at the beginning of wheat's cultivation . Wheat with this mutation was harvested more frequently and became the seed for the next crop. Therefore, without realizing it, early farmers selected for this mutation. The result

4600-417: The domestication syndrome , traits presumed essential in the early stages of domestication, while others represent later improvement traits. Domesticated mammals in particular tend to be smaller and less aggressive than their wild counterparts; other common traits are floppy ears, a smaller brain, and a shorter muzzle. Domestication traits are generally fixed within all domesticates, and were selected during

4700-437: The rhinoviruses . Many parasites , too, have their origins in domestic animals. Alongside these, the advent of domestication resulted in denser human populations, which provided ripe conditions for pathogens to reproduce, mutate, spread, and eventually find a new host in humans. Scholars have expressed widely differing viewpoints on domestication's effects on society. Anarcho-primitivism critiques domestication as destroying

4800-436: The silkworm and the western honey bee , have been domesticated for over 5,000 years, often for commercial use. The silkworm is raised for the silk threads wound around its pupal cocoon; the western honey bee, for honey , and, from the 20th century, for pollination of crops. Several other invertebrates have been domesticated, both terrestrial and aquatic, including some such as Drosophila melanogaster fruit flies and

4900-481: The " falling cat problem ". The cat family (Felidae) can pass down many colors and patterns to their offspring. The domestic cat genes MC1R and ASIP allow color variety in their coats. The feline ASIP gene consists of three coding exons. Three novel microsatellite markers linked to ASIP were isolated from a domestic cat BAC clone containing this gene to perform linkage analysis on 89 domestic cats segregated for melanism . The domestic cat family demonstrated

5000-707: The "Sterling" accepted by the International Cat Exhibitors (ICE) for championship status in 1998. The breed had its own registry under the International Sterling Society. However, not many breeders of chinchilla Persian elected to switch over from the existing Persian standard to the new Sterling standard. Mrs Johnson died in 2006 before she could get wider buy-in for the breed. Due to lack of support, this Sterling breed does not exist anymore. In South Africa, all breeds cat judge and Chinchilla breeder Stella Slabber, headed up

5100-690: The 2010 WCF breed recognition show in Cape Town was based on the SACC standard with a request to include the golden varieties. After consultation with the Chairperson of the WCF Standard- and Judges Commission, Cornelia Hungerecker, the breed name was changed to "Tradition Longhair". One other minor differentiation was that the eye shape should read "walnut shaped" instead of "round". The 2010 standard for Traditional Longhair silver and golden varieties

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5200-465: The American type and coat as well as the attractiveness of the kittens. She proposed a new breed, called the "Sterling" because of their silver color. These new cats were bred to look like Persians but with a short, dense plush coat. The name was changed to "Exotic Shorthair" when all colors were accepted in 1979. In September 1992 the "English Sterling" as a New Breed was presented by Jeanne Johnson at

5300-555: The Americas began to cultivate peanuts , squash , maize , potatoes , cotton , and cassava . Rice was first domesticated in China some about 9,000 years ago. In Africa, crops such as sorghum were domesticated. Agriculture developed in some 13 centres around the world, domesticating different crops and animals. Three groups of insects, namely ambrosia beetles , leafcutter ants , and fungus-growing termites have independently domesticated species of fungi, on which they feed. In

5400-592: The Americas, Africa, and Asia (the Middle East, South Asia, the Far East, and New Guinea and Wallacea); in some thirteen of these regions people began to cultivate grasses and grains. Rice was first cultivated in East Asia. Sorghum was widely cultivated in sub-Saharan Africa, while peanuts, squash, cotton, maize , potatoes , and cassava were domesticated in the Americas. Continued domestication

5500-561: The Dragon . Persian cats originate from Persia ( Iran ). The cats were introduced in Europe in the 1500s as highly valued items of trade. The Europeans were impressed by the Persian's long, silky coat and purposefully bred the cats to perpetuate the trait. Two mutations of note influenced the Persian cats: the color inhibitor gene (1882) that gave rise to the so-called "chinchilla" coat and

5600-595: The Late Pleistocene. Domestication reduces genetic diversity of the domesticated population, especially of alleles of genes targeted by selection. One reason is a population bottleneck created by artificially selecting the most desirable individuals to breed from. Most of the domesticated strain is then born from just a few ancestors, creating a situation similar to the founder effect . Domesticated populations such as of dogs, rice, sunflowers, maize, and horses have an increased mutation load , as expected in

5700-675: The TICA 13th Annual Meeting. Follow-up work resulted in the breed with the name "Sterling" (without 'English' in front) was accepted as Category 1 in TICA in Feb 1994. (Category 1 in TICA means "Established Breeds".) The Sterling had Category 1 status only for 18 months, before it was scrapped in total by TICA in September 1995. The assumed reason is that there was a lack of buy-in from enough fellow breeders as required by TICA breed recognition rules. Despite this set-back, Jeannie Johnson continued by getting

5800-447: The US advocated pure color breeding. A key person in this regard was Jeanne Ramsdale, author of the book "Persian Cats and Other Longhairs". Despite the non-conformance to the modern Persian breed standards, some breeders, and in particular those that practiced controlled color-breeding, continued to breed and register their original longhair cats. The following attempts were made to restore

5900-505: The WCF in August 2010. The breed proposal was compiled in Cape Town by Alida Delport. The facilitator of the breed recognition process was Dr. Johan Lamprecht, who also did the breed proposal presentation at the WCF. Nestle Purina sponsored the breed recognition show due to these cats being the brand animal used in their cat food advertisements world-wide. The initial standard that was proposed at

6000-458: The beginning of the Holocene 11,700 years ago, a warmer climate and increasing human populations led to small-scale animal and plant domestication and an increased supply of food. The appearance of the domestic dog in the archaeological record, at least 15,000 years ago, was followed by domestication of livestock and of crops such as wheat and barley , the invention of agriculture , and

6100-648: The beginning of the 1st century AD. By the end of the Western Roman Empire in the 5th century, the Egyptian domestic cat lineage had arrived in a Baltic Sea port in northern Germany . The leopard cat ( Prionailurus bengalensis ) was tamed independently in China around 5500 BC. This line of partially domesticated cats leaves no trace in the domestic cat populations of today. During domestication, cats have undergone only minor changes in anatomy and behavior, and they are still capable of surviving in

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6200-470: The brachycephalic mutation (circa 1942) that gave rise to the so-called "peke-faced" or "snub-nosed" Persian cats. The brachycephalic mutation in the Persian breed occurred during World War II in the US. This led to a concerted effort to breed Persians with rounder heads and smaller ears than their ancient ancestors after World War II. The resultant brachycephalic heads lead to the much desired snub noses favored by many modern Persian breeders and which dictates

6300-533: The breed disappearing from cat shows. However, this older form of Persian cat was made iconic to the general public through its use in popular films, even as it was being phased out in the cat fancy. For example, the Traditional Persian has been featured in eight of the James Bond movies, beginning with " From Russia with Love " in 1963. It was also featured in the kung fu cult classic Enter

6400-506: The case of the termites, the relationship is a fully obligate symbiosis on both sides. Domestication (not to be confused with the taming of an individual animal ), is from the Latin domesticus , 'belonging to the house'. The term remained loosely defined until the 21st century, when the American archaeologist Melinda A. Zeder defined it as a long-term relationship in which humans take over control and care of another organism to gain

6500-624: The cat and other wild mammals to the island from the Middle Eastern mainland. Scientists therefore assume that African wildcats were attracted to early human settlements in the Fertile Crescent by rodents, in particular the house mouse ( Mus musculus ), and were tamed by Neolithic farmers. This mutual relationship between early farmers and tamed cats lasted thousands of years. As agricultural practices spread, so did tame and domesticated cats. Wildcats of Egypt contributed to

6600-509: The domestication of animals involved violence against animals and damage to the environment. This, in turn, he argues, corrupted human ethics and paved the way for "conquest, extermination, displacement, repression, coerced and enslaved servitude, gender subordination and sexual exploitation, and hunger." Domesticated ecosystems provide food, reduce predator and natural dangers, and promote commerce, but their creation has resulted in habitat alteration or loss, and multiple extinctions commencing in

6700-515: The domestication syndrome; it is predicted that domestic animals are less resistant to parasites than their wild relatives. Domesticated birds principally mean poultry , raised for meat and eggs: some Galliformes ( chicken , turkey , guineafowl ) and Anseriformes (waterfowl: ducks , geese , and swans ). Also widely domesticated are cagebirds such as songbirds and parrots ; these are kept both for pleasure and for use in research. The domestic pigeon has been used both for food and as

6800-574: The extinction of bird , mammal , and reptile species. As of 2017, the domestic cat was the second most popular pet in the United States, with 95.6 million cats owned and around 42 million households owning at least one cat. In the United Kingdom, 26% of adults have a cat, with an estimated population of 10.9 million pet cats as of 2020. As of 2021, there were an estimated 220 million owned and 480 million stray cats in

6900-407: The eyes from damage. Outdoor cats are active both day and night, although they tend to be slightly more active at night. Domestic cats spend the majority of their time in the vicinity of their homes but can range many hundreds of meters from this central point. They establish territories that vary considerably in size, in one study ranging 7–28 ha (17–69 acres). The timing of cats' activity

7000-447: The freshwater cnidarian Hydra for research into genetics and physiology. Few have a long history of domestication. Most are used for food or other products such as shellac and cochineal . The phyla involved are Cnidaria , Platyhelminthes (for biological pest control ), Annelida , Mollusca , Arthropoda (marine crustaceans as well as insects and spiders), and Echinodermata . While many marine mollusks are used for food, only

7100-810: The genome and the epigenome , enabling a rapid evolutionary response to artificial selection. Polyploidy increases the number of chromosomes, bringing new combinations of genes and alleles, which in turn enable further changes such as by chromosomal crossover . The microbiome , the collection of microorganisms inhabiting the surface and internal tissue of plants, is affected by domestication. This includes changes in microbial species composition and diversity. Plant lineage, including speciation , domestication, and breeding , have shaped plant endophytes ( phylosymbiosis ) in similar patterns as plant genes. Several species of fungi have been domesticated for use directly as food, or in fermentation to produce foods and drugs. The cultivated mushroom Agaricus bisporus

7200-483: The ground and allows for the silent stalking of prey. The claws on the forefeet are typically sharper than those on the hindfeet. Cats can voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing, kneading , or for extra traction on soft surfaces. Cats shed the outside layer of their claw sheaths when scratching rough surfaces. Most cats have five claws on their front paws and four on their rear paws. The dewclaw

7300-414: The hair length is darker pigmented). The sub-set of chinchilla varieties experienced problems during the brachycephalic modernization phase with regards to the original chinchilla coloring features. These included loss or incomplete lip- and nose liners; patched-colored paw pads and eye color faults. Once lost, it is hard, if not impossible, to recover. By the mid-1950s, concerned breeders of chinchilla in

7400-411: The high IMP and free l-Histidine contents of tuna, which produces a strong umami taste synergy that is highly preferred by cats". One of the researchers in this research has stated, "I think umami is as important for cats as sweet is for humans". Cats also have a distinct temperature preference for their food, preferring food with a temperature around 38 °C (100 °F) which is similar to that of

7500-532: The homogenizing effect of gene flow from wild boars into pigs, and created domestication islands in the genome. The same process may apply to other domesticated animals. The 2023 parasite-mediated domestication hypothesis suggests that endoparasites such as helminths and protozoa could have mediated the domestication of mammals. Domestication involves taming, which has an endocrine component; and parasites can modify endocrine activity and microRNAs . Genes for resistance to parasites might be linked to those for

7600-426: The initial episode of domestication of that animal or plant, whereas improvement traits are present only in a proportion of domesticates, though they may be fixed in individual breeds or regional populations . Certain animal species, and certain individuals within those species, make better candidates for domestication because of their behavioral characteristics: The beginnings of mammal domestication involved

7700-580: The leaf material to the fungi that they tend. Some of these fungi are not fully domesticated: the fungi farmed by Mycocepurus smithii constantly produce spores that are not useful to the ants, which eat fungal hyphae instead. The process of domestication by Atta ants, on the other hand, is complete; it took 30 million years. Some 330 fungus-growing termite species of the subfamily Macrotermitinae cultivate Termitomyces fungi to eat; domestication occurred exactly once, 25–40 mya . The fungi, described by Roger Heim in 1942, grow on 'combs' formed from

7800-634: The maternal gene pool of the domestic cat at a later time. The earliest known evidence for the occurrence of the domestic cat in Greece dates to around 1200 BC. Greek, Phoenician , Carthaginian and Etruscan traders introduced domestic cats to southern Europe. By the 5th century BC, they were familiar animals around settlements in Magna Graecia and Etruria . During the Roman Empire , they were introduced to Corsica and Sardinia before

7900-602: The modern domesticated subspecies F. silvestris catus sampled worldwide was considered to have likely descended from the African wildcat ( F. lybica ), following results of phylogenetic research. In 2017, the IUCN Cat Classification Taskforce followed the recommendation of the ICZN in regarding the domestic cat as a distinct species, Felis catus . The domestic cat is a member of the Felidae,

8000-488: The modern standard for Persians in cat fancies throughout the world. The color inhibitor gene mutated in a Persian cross in 1882 in the UK in a cat called "Chinnie". The offspring of this mutation that express the gene are commonly known as "chinchillas" presumably due to the resemblance of their coat to that of the rodent by that name . They were classified as a Persian variety for breeding purposes. The chinchilla comes in both

8100-489: The original longhairs as breed in their own right, namely: There is some confusion around the name "Sterling". In the International Cat Association (TICA), there were some American Shorthair breeders who bred to Persians to obtain their silver color and green eyes. The kittens were attractive but did not meet the true American Shorthair type. Jane Martinke noticed the effect the outcrosses had on

8200-487: The receptors needed to detect umami . However, these receptors contain molecular changes that make cat taste umami different from that of humans. In humans, they detect the amino acids glutamic acid and aspartic acid , but in cats, they instead detect inosine monophosphate and l-Histidine . These molecules are particularly enriched in tuna . This, it has been argued, is why cats find tuna so palatable : as put by researchers into cat taste, "the specific combination of

8300-514: The relationship between them intensified, and humans' role in their survival and reproduction gradually led to formalized animal husbandry . Although the directed pathway for draft and riding animals proceeded from capture to taming, the other two pathways are not as goal-oriented, and archaeological records suggest that they took place over much longer time frames. Unlike other domestic species selected primarily for production-related traits, dogs were initially selected for their behaviors. The dog

8400-516: The silver and golden series, with black silver chinchilla as the most common. Chinchilla cats are almost pure white Persian cats with a sprinkling of darker colored tipping (shell) that gives them a sparkling metallic appearance. The chinchilla should have only a tiny amount of darker colored tipping on their face and legs. Only the hair tips are darker pigmented (approx. 1/8 of the hair length), and chinchillas are significantly lighter in overall color compared to Persians with shaded coats (approx. 1/3 of

8500-518: The size of their jaw, which is an adaptation to their preferred prey of small rodents, which have small vertebrae. The premolar and first molar together compose the carnassial pair on each side of the mouth, which efficiently shears meat into small pieces, like a pair of scissors. These are vital in feeding, since cats' small molars cannot chew food effectively, and cats are largely incapable of mastication. Cats tend to have better teeth than most humans, with decay generally less likely because of

8600-459: The small number of traits that made domestic species different from their wild ancestors. He was also the first to recognize the difference between conscious selective breeding in which humans directly select for desirable traits and unconscious selection, in which traits evolve as a by-product of natural selection or from selection on other traits. There is a difference between domestic and wild populations; some of these differences constitute

8700-451: The supposed primitive state of harmony with nature in hunter-gatherer societies, and replacing it, possibly violently or by enslavement, with a social hierarchy as property and power emerged. The dialectal naturalist Murray Bookchin has argued that domestication of animals, in turn, meant the domestication of humanity, both parties being unavoidably altered by their relationship with each other. The sociologist David Nibert asserts that

8800-571: The transition of humans from foraging to farming in different places and times across the planet. For instance, small-scale trial cultivation of cereals began some 28,000 years ago at the Ohalo II site in Israel. In the Fertile Crescent 11,000–10,000 years ago, zooarchaeology indicates that goats, pigs, sheep, and taurine cattle were the first livestock to be domesticated. Two thousand years later, humped zebu cattle were domesticated in what

8900-862: The wild members of this genus evolved through sympatric or parapatric speciation , whereas the domestic cat evolved through artificial selection . The domestic cat and its closest wild ancestor are diploid and both possess 38 chromosomes and roughly 20,000 genes. Pantherinae other Felinae lineages Jungle cat ( F. chaus ) [REDACTED] Black-footed cat ( F. nigripes ) Sand cat ( F. margarita ) Chinese mountain cat ( F. bieti ) African wildcat ( F. lybica ) European wildcat ( F. silvestris ) [REDACTED] Domestic cat [REDACTED] Sand cat ( F. margarita ) Chinese mountain cat ( F. bieti ) European wildcat ( F. silvestris ) [REDACTED] Southern African wildcat (F. l. cafra) Asiatic wildcat (F. l. ornata) Near Eastern wildcat Domestic cat [REDACTED] It

9000-408: The wild. The first animal to be domesticated by humans was the dog , as a commensal , at least 15,000 years ago. Other animals, including goats , sheep , and cows , were domesticated around 11,000 years ago. Among birds, the chicken was first domesticated in East Asia, seemingly for cockfighting, some 7,000 years ago. The horse came under domestication around 5,500 years ago in central Asia as

9100-625: The wild. Several natural behaviors and characteristics of wildcats may have pre-adapted them for domestication as pets. These traits include their small size, social nature, obvious body language, love of play, and high intelligence. Since they practice rigorous grooming habits and have an instinctual drive to bury and hide their urine and feces, they are generally much less messy than other domesticated animals. Captive Leopardus cats may also display affectionate behavior toward humans but were not domesticated. House cats often mate with feral cats. Hybridization between domestic and other Felinae species

9200-548: The world. The origin of the English word cat , Old English catt , is thought to be the Late Latin word cattus , which was first used at the beginning of the 6th century. The Late Latin word may be derived from an unidentified African language . The Nubian word kaddîska 'wildcat' and Nobiin kadīs are possible sources or cognates. The forms might also have derived from an ancient Germanic word that

9300-403: The year in equatorial regions , with litter sizes often ranging from two to five kittens. Domestic cats are bred and shown at events as registered pedigreed cats , a hobby known as cat fancy . Animal population control of cats may be achieved by spaying and neutering , but their proliferation and the abandonment of pets has resulted in large numbers of feral cats worldwide, contributing to

9400-412: Was absorbed into Latin and then into Greek, Syriac, and Arabic. The word may be derived from Germanic and Northern European languages, and ultimately be borrowed from Uralic , cf. Northern Sámi gáđfi , 'female stoat ', and Hungarian hölgy , 'lady, female stoat'; from Proto-Uralic * käďwä , 'female (of a furred animal)'. The English puss , extended as pussy and pussycat ,

9500-500: Was domesticated long before other animals, becoming established across Eurasia before the end of the Late Pleistocene era, well before agriculture . The archaeological and genetic data suggest that long-term bidirectional gene flow between wild and domestic stocks – such as in donkeys , horses , New and Old World camelids, goats, sheep, and pigs – was common. Human selection for domestic traits likely counteracted

9600-474: Was extended by the World Cat Federation (WCF) in 2012 to include all Persian color varieties. In 2014 the World Cat Federation (WCF) changed the name "Traditional Longhair" to "Original Longhair". The World Cat Congress promotes better understanding and co-operation among the world's major cat associations in matters of mutual interest and concern. It has nine members. In 2014, after a webinar,

9700-532: Was gradual and geographically diffuse – happening in many small steps and spread over a wide area – on the evidence of both archaeology and genetics. It was a process of intermittent trial and error and often resulted in diverging traits and characteristics. Whereas domestication of animals impacted most on the genes that controlled behavior, that of plants impacted most on the genes that controlled morphology (seed size, plant architecture, dispersal mechanisms) and physiology (timing of germination or ripening), as in

9800-519: Was long thought that the domestication of the cat began in ancient Egypt , where cats were venerated from around 3100 BC. However, the earliest known indication for the taming of an African wildcat was excavated close by a human Neolithic grave in Shillourokambos , southern Cyprus , dating to about 7500–7200 BC. Since there is no evidence of native mammalian fauna on Cyprus, the inhabitants of this Neolithic village most likely brought

9900-631: Was proposed by Carl Linnaeus in 1758 for a domestic cat. Felis catus domesticus was proposed by Johann Christian Polycarp Erxleben in 1777. Felis daemon proposed by Konstantin Satunin in 1904 was a black cat from the Transcaucasus , later identified as a domestic cat. In 2003, the International Commission on Zoological Nomenclature ruled that the domestic cat is a distinct species, namely Felis catus . In 2007,

10000-557: Was triggered by the climatic and environmental changes that occurred after the peak of the Last Glacial Maximum and which continue to this present day. These changes made obtaining food by hunting and gathering difficult. The first animal to be domesticated was the dog at least 15,000 years ago. The Younger Dryas 12,900 years ago was a period of intense cold and aridity that put pressure on humans to intensify their foraging strategies but did not favour agriculture. By

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